ZOOTAXA ISSN 1175-5326 (Print Edition) & ISSN 1175-5334 (Online Edition)
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Tardigrada  

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Editor, Dr Clark W. Beasley (beasleyc@mcmurryadm.mcm.edu)
Clark W. Beasley, Department of Biology, McMurry University, Abilene, Texas 79697, U.S.A.

2009

 Article 
Zootaxa 2124: 1-20 (3 Jun. 2009) 7 plates; 10 references                            Accepted: 29 Apr. 2009
Tardigrades of the Seychelles Islands, with the description of three new species
GIOVANNI PILATO & OSCAR LISI (Italy)

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 Article 
Zootaxa 2095: 21-36 (8 May 2009) 5 plates; 44 references                          Accepted: 18 Apr. 2009
Tardigrades from northwestern Patagonia (Neuquén Province, Argentina) with
the description of three new species
GUSTAVO ROSSI, MARIA CLAPS & DIEGO ARDOHAIN (Argentina)

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 Article 
Zootaxa 2076: 63-68 (17 Apr. 2009) 2 plates; 6 references                           Accepted: 12 Mar. 2009
Revision of Biserovus xiae Li, Su & Yu, 2004 (Tardigrada: Macrobiotidae)
XIAOCHEN LI (China)

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 Article 
Zootaxa 2058: 62-68 (30 Mar. 2009) 2 plates; 12 references                           Accepted: 17 Feb. 2009
Bindius triquetrus gen. nov. sp. nov. (Eutardigrada, Hypsibiidae)
from Sicily (Italy)
GIOVANNI PILATO (Italy)

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 Article 
Zootaxa 2030: 21-38 (9 Mar. 2009) 4 plates; 76 references                           Accepted: 17 Dec. 2008
Tardigrades from Portugal: four new records and description of two new
species
PAULO FONTOURA (Portugal), GIOVANNI PILATO (Italy), OSCAR LISI (Italy) & PAULO MORAIS
(Portugal)
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 Article 
Zootaxa 2022: 45-57 (27 Feb. 2009) 7 plates; 30 references                           Accepted: 19 Jan. 2009
Macrobiotus (Eutardigrada, Macrobiotidae) from the Great Smoky Mountains
National Park, Tennessee/North Carolina, USA (North America): two new species and six new records
PAUL J. BARTELS (USA), GIOVANNI PILATO (Italy), OSCAR LISI (Italy) & DIANE R. NELSON
(USA)
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 Article 
Zootaxa 2005: 24-34 (9 Feb. 2009) 4 plates; 24 references                           Accepted: 17 Dec. 2008
Description of three new species of Tardigrada from the Seychelles
GIOVANNI PILATO & OSCAR LISI (Italy)

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2008

 Article 
Zootaxa 1940: 41-47 (24 Nov. 2008) 5 plates; 29 references                          Accepted: 22 Oct. 2008
Designation of Pseudobiotus kathmanae Nelson, Marley & Bertolani, 1999 as the type species for the genus Pseudobiotus Nelson, 1980 (Tardigrada)
NIGEL J. MARLEY (UK), ROBERTO BERTOLANI (Italy) & DIANE R. NELSON (USA)

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 Article 
Zootaxa 1856: 41-54 (25 Aug. 2008) 8 plates; 17 references                              Accepted: 4 Aug. 2008
New description of Echiniscus scabrospinosus Fontoura, 1982, and description of a new species of Echiniscus (Heterotardigrada) from China
Giovanni Pilato (Italy) , Paulo Fontoura (Portugal), OSCAR LISI (ITALY) & Clark Beasley (USA)

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 Article 
Zootaxa 1778: 37-47 (28 May 2008) 4 plates; 31 references                              Accepted: 11 Apr. 2008
New records of eutardigrades (Tardigrada) from Faial and Pico Islands, the Azores, with the description of two new species
PAULO FONTOURA (Portugal), GIOVANNI PILATO (Italy) & OSCAR LISI (Italy)

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 Article 
Zootaxa 1757: 1-30 (28 Apr. 2008) 7 plates; 54 references                              Accepted: 27 Mar. 2008
New records and within-species variability of Iberian tardigrades (Tardigrada), with comments on the species from the Echiniscus blumi-canadensis series
NOEMI GUIL (USA)

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 Article 
Zootaxa 1731: 1-23 (25 Mar. 2008) 17 plates; 18 references                              Accepted: 10 Jan. 2008
Macrobiotus derkai, a new species of Tardigrada (Eutardigrada, Macrobiotidae, huziori group) from the Colombian Andes (South America)
PETER DEGMA (Slovakia), LUKASZ MICHALCZYK (UK) & LUKASZ KACZMAREK (Poland)

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 Article 
Zootaxa 1693: 49-61 (31 Jan. 2008) 3 plates; 33 references                              Accepted: 7 Dec. 2007
Echiniscidae (Tardigrada, Heterotardigrada) from Faial and Pico Islands, the Azores, with the description of two new species
PAULO FONTOURA (Portugal), GIOVANNI PILATO (Italy) & OSCAR LISI (Italy)

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2007

 Article 
Zootaxa 1646: 59-65 (26 Nov. 2007)  3 plates; 10 references                            Accepted: 2 Oct. 2007
Doryphoribius smokiensis, a new species of Eutardigrada (Hypsibiidae) from the Great Smoky Mountains National Park, TN, USA (North America)
PAUL J. BARTELS (USA), DIANE R. NELSON (USA), LUKASZ KACZMAREK (Poland) & LUKASZ MICHALCZYK (UK)
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 Article 
Zootaxa 1626: 59-67 (31 Oct. 2007)  2 plates; 21 references                            Accepted: 27 Sep. 2007
Tardigrades from Guizhou Province (China) with description of a new species of genus Doryphoribius (Tardigrada: Hypsibiidae)
HANPING WANG, LIZHI WANG & XIAOCHEN LI (P.R. China)

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 Article 
Zootaxa 1589: 47-55 (19 Sep. 2007)  4 plates; 6 references                            Accepted: 11 Aug. 2007
Diphascon (Diphascon) faialense sp. nov. a new species of Tardigrada (Eutardigrada, Hypsibiidae) from the Azores and a key to the species of the D. pingue group
PAULO FontourA (Portugal) & GIOVANNI Pilato (Italy)

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 Article 
Zootaxa 1579: 41-53 (10 Sep. 2007)  0 plates; 46 references                            Accepted: 28 Jul. 2007
Notes to the current checklist of Tardigrada
PETER DEGMA (Slovakia) & ROBERTO GUIDETTI (Italy)

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 Article 
Zootaxa 1479: 1-7 (21 May 2007) 2 plates; 8 references                            Accepted: 18 Jan. 2007
Macrobiotus alvaroi
, a new species of eutardigrade (Tardigrada, Macrobiotidae) of the polyopus group from Costa Rica (Central America)
Giovanni Pilato (Italy) & Lukasz Kaczmarek (Poland)

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 Article 
Zootaxa 1471: 15-25 (10 May 2007) 6 plates; 9 references                            Accepted: 28 Feb. 2007
Echiniscus ganczareki, a new species of Tardigrada (Heterotardigrada: Echiniscidae, bigranulatus group) from Costa Rica
L
UKASZ MICHALCZYK (UK) & LUKASZ KACZMAREK (Poland)

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 Article 
Zootaxa 1471: 61-68 (10 May 2007) 2 plates; 28 references                              Accepted: 20 Mar. 2007
New taxonomic position of several Macrobiotus species (Eutardigrada: Macrobiotidae)
ROBERTO GUIDETTI (Italy), ROBERTO BERTOLANI (Italy) & PETER DEGMA (Slovakia)

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 Article 
Zootaxa 1389: 55-60 (11 Jan. 2007)  3 plates; 2 references                           Accepted: 29 Nov. 2006
Echiniscus quitensis, a new species of tardigrade from Ecuador (Heterotardigrada: Echiniscidae)
G. PILATO (Italy)
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2006

Zootaxa 1373: 65-68 (7 Dec. 2006) 1 plate; 8 references                             Accepted: 30 Oct. 2006
Thulinius itoi comb. nov. a new systematic position for Isohypsibius itoi (Tsurusaki, 1980) (Tardigrada: Eutardigrada: Hypsibiidae)
L. Kaczmarek (Poland) & L. Michalczyk (UK)

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Zootaxa 1371: 45-56 (4 Dec. 2006) 5 plates; 21 references                             Accepted: 26 Oct. 2006
Macrobiotus sklodowskae sp. nov. (Tardigrada: Eutardigrada: Macrobiotidae, richtersi group) from Cyprus
L. MICHALCZYK (UK), L. KACZMAREK (Poland) & B. WEGLARSKA (Poland)

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Zootaxa 1369: 57-68 (30 Nov. 2006) 32 references                                        Accepted: 16 Nov. 2006
Tardigrada nova: a bibliographic analysis and catalogue of new species described in Zootaxa before 2006
Z.-Q. ZHANG (New Zealand)
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Zootaxa 1298: 37-47 (17 Aug. 2006) 3 plates; 9 references                              Accepted:  28 Jul. 2006
Remarks on the Macrobiotus polyopus group, with the description
of two new species (Eutardigrada, Macrobiotidae)
G. PILATO (Italy)

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Zootaxa 1298: 49-60 (17 Aug. 2006) 5 plates; 19 references                              Accepted:  14 Jul. 2006
Redescription of Caprella dilatata Krøyer, 1843
(Caprellidae: Amphipoda: Crustacea) from Brazil, with note on its biogeographical distribution in South America
S. MASUNARI (Brazil) & I. TAKEUCHI (Japan)

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Zootaxa 1298: 37-47 (17 Aug. 2006) 3 plates; 9 references                              Accepted:  28 Jul. 2006
Remarks on the Macrobiotus polyopus group, with the description
of two new species (Eutardigrada, Macrobiotidae)
G. PILATO (Italy)

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Zootaxa 1236: 53-68 (19 Jun. 2006) 4 plates; 28 references                             Accepted: 19 Apr. 2006
Notes on some tardigrades from southern Mexico with description of three new species
G. PILATO & O. LISI (Italy) 
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Zootaxa 1229: 49-57 (12 Jun. 2006) 6 plates; 6 references                               Accepted: 15 May 2006
Ramazzottius bunikowskae, a new species of Tardigrada (Eutardigrada, Hypsibiidae) from Russia
L. KACZMAREK (Poland), L. MICHALCZYK (Poland) & D. DIDUSZKO (UK) 
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Zootaxa 1202: 53-59 (15 May 2006) 2 plates; 7 references                               Accepted: 18 Apr. 2006
Macrobiotus siamensis sp. n. (Eutardigrada, Macrobiotidae) from Thailand (Asia)
D.V. TUMANOV (Russia) 
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Zootaxa 1172: 68 (13 Apr. 2006
Corrigenda: Marley, N.J. (2006) Zootaxa, 1166, 35–48
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Zootaxa 1169: 47-59 (10 Apr. 2006) 12 plates; 7 references                               Accepted: 25 Jan. 2006
Macrobiotus huziori, a new species of Tardigrada (Eutardigrada: Macrobiotidae) from Costa Rica (Central America)
L. MICHALCZYK & L. KACZMAREK (Poland)
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Zootaxa 1166: 35-48 (3 Apr. 2006) 6 plates; 22 references                              Accepted: 24 Feb. 2006
A new species of Tardigrada, Platicrista ramsayi sp. nov. from the páramo of Volcán Chiles, Ecuador
N.J. MARLEY (UK)
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Zootaxa 1154: 1-26 (20 Mar. 2006) 17 plates; 5 references                              Accepted: 25 Jan. 2006
Revision of the Echiniscus bigranulatus group with a description of a new species Echiniscus madonnae (Tardigrada: Heterotardigrada: Echiniscidae) from South America
L. MICHALCZYK & L. KACZMAREK (Poland)
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Zootaxa 1122: 1-23 (2 Feb. 2006) 8 plates; 10 references                                Accepted: 16 Dec. 2006
Five new species of the genus Milnesium (Tardigrada, Eutardigrada, Milnesiidae)
D.V. TUMANOV (Russia)
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Zootaxa 1109: 49-55 (17 Jan. 2006) 2 plates; 9 references                                  Accepted: 7 Nov. 2005
Macrobiotus rigidus sp. nov., new species of eutardigrade from New Zealand
GIOVANNI PILATO & OSCAR LISI (Italy)
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Zootaxa 1105: 1-16 (10 Jan. 2006) 1 plate; 66 references                                Accepted: 26 Oct. 2005
The Bryofauna of Remote Coastal Labrador: including a review of current Canadian records
M. BOECKNER, M. COLLINS, J. FINNEY-CRAWLEY & L. BATEMAN (Canada)
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2005

Zootaxa 1093: 25-33 (8 Dec. 2005) 3 plates; 16 references                                 Accepted: 8 Oct. 2005
A new species and five new records of the family Echiniscidae (Tardigrada) from China
X.C. LI, L.Z. WANG, Y. LIU & L.N. SU (P.R. China)
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Zootaxa 1092: 39-45 (8 Dec. 2005) 3 plates; 3 references                                 Accepted: 19 Oct. 2005
Echiniscus taibaiensis sp. nov. and a new record of Echiniscus bisetosus Heinis (Tardigrada, Echiniscidae) from China
L.Z. WANG & X.C. LI  (P.R. China)
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Zootaxa 1080: 33-38 (15 Nov. 2005) 3 plates;  5 references                              Accepted: 30 Sept. 2005
A new species of Tardigrada Bryodelphax brevidentatus sp. nov. (Heterotardigrada: Echiniscidae) from China (Asia)
L. KACZMAREK (Poland) , L. MICHALCZYK (UK) & P. DEGMA (Slovakia)
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Zootaxa 1075: 55-68 (4 Nov. 2005) 5 plates; 29 references                             Accepted: 9 Sept. 2005
Remarks on Macrobiotus furciger Murray, 1906 and description of three new species of the furciger group (Eutardigrada, Macrobiotidae)
M. G. BINDA, G. PILATO & O. LISI (Italy)
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Zootaxa 1058: 43-49 (4 Oct. 2005) 1 plate; 26 references                             Accepted: 6 Sept. 2005
Tardigrade fauna of the South Sandwich Islands, maritime Antarctic
S. J. MCINNES & P. CONVEY  (UK)
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Zootaxa 1053: 35-45 (23 Sept. 2005) 3 plates; 20 references                           Accepted: 16 Sept. 2005
Some tardigrades from Siberia (Russia, Baikal region) with a description of Macrobiotus garynahi sp. nov. (Eutardigrada: Macrobiotidae: richtersi group)
L. KACZMAREK, L. MICHALCZYK & D. DIDUSZKO  (Poland)
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Zootaxa 1043: 33-46 (30 Aug. 2005) 5 plates; 17 references                               Accepted: 5 Aug. 2005
Two new species of Macrobiotus (Eutardigrada, Macrobiotidae) from Tien Shan (Kirghizia), with notes on Macrobiotus tenuis group
D.V. TUMANOV (Russia)
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Zootaxa 1036: 55-60 (15 Aug. 2005) 1 plate; 19 references                              Accepted: 27 May 2005
Isohypsibius taibaiensis sp. nov. (Tardigrada, Hypsibiidae) from China
X.C. LI & L.Z. WANG (P.R. China)
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Zootaxa 1027: 27-45 (29 Jul. 2005) 11 plates; 15 references                             Accepted: 29 May 2005
Remarks on some Echiniscidae (Heterotardigrada) from New Zealand with the description of two new species
G. PILATO, M.G. BINDA & O. LISI (Italy)
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Zootaxa 1022: 57-64 (25 Jul. 2005) 3 plates; 23 references                               Accepted: 28 May 2005
Tardigrades of the Australian Antarctic: Hypsibius heardensis (Eutardigrada: Hypsibiidae: dujardini group) a new species from sub-Antarctic Heard Island
W.R. MILLER (USA) , S.J. MCINNES (UK), & D.M. BERGSTROM  (Australia)
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Zootaxa 1008: 47-56 (14 Jun. 2005) 8 plates; 6 references                               Accepted: 20 May 2005
Minibiotus weglarskae
, a new species of Tardigrada (Eutardigrada: Macrobiotidae) from Mongolia
Ł. MICHALCZYK (Poland), Ł. KACZMAREK  (Poland) & S.K. CLAXTON (Australia)
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Zootaxa 999: 1-16 (27 May 2005) 8 plates; 25 references                                  Accepted: 11 May 2005
Notes on the Tardigrada of Thailand, with a description of Macrobiotus alekseevi sp. nov. (Eutardigrada, Macrobiotidae)
D.V. TUMANOV (Russia)
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Zootaxa 980: 1-4 (16 May 2005) 1 plate; 4 references                                  Accepted: 9 May 2005
Isohypsibius qinlingensis sp. nov. (Tardigrada, Hypsibiidae) from China
X. LI, L. WANG & D. YU (P.R. China)
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Zootaxa 914: 1-5 (24 Mar. 2005) 1 plate; 7 references                                    Accepted:4 Mar. 2005
Diphascon (Diphascon) dolomiticum, a new species of Hypsibiidae (Eutardigrada) from Italy
G. PILATO & R. BERTOLANI (Italy
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Zootaxa 889: 1-11 (7 Mar. 2005) 2 plates; 22 references                                   Accepted: 23 Feb. 2005
A new species of Tardigrada (Eutardigrada: Macrobiotidae) from Iberian Peninsula and Canary Islands (Spain)
N. GUIL (Spain) & R. GUIDETTI (Italy) 
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Zootaxa 879: 1-7 (4 Mar. 2005) 4 plates; 11 references                                   Accepted: 16 Feb. 2005
Haplomacrobiotus utahensis new species of Calohypsibiidae (Eutardigrada) from North America
GIOVANNI PILATO (Italy) & CLARK BEASLEY (USA)
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Zootaxa 845: 1-46 (1 Feb. 2005)  147 references                                           Accepted: 12 Jan. 2005
Tardigrade taxonomy: an updated check list of the taxa and a list of characters for their identification
R. GUIDETTI & R. BERTOLANI (Italy)
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Zootaxa 812: 1-4 (14 Jan. 2005) 2 plates, 5 references                                    Accepted: 29 Dec. 2004
Isohypsibius panovi, a new species of Tardigrada from Ireland (Eutardigrada, Hypsibiidae)
D.V. TUMANOV (Russia)
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2004

Zootaxa 743: 1-5 (25 November 2004) 2 plates; 10 references                      Accepted: 22 Nov. 2004
Milnesium katarzynae sp. nov., a new species of eutardigrade (Milnesiidae) from China
Ł. KACZMAREK (Poland), Ł. MICHALCZYK (Poland) & C.W. BEASLEY
(USA)
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Zootaxa 660: 1-7 (29 September 2004)  1 plate; 9 references                         Accepted: 6 Sept. 2004
Biserovus xiae sp. n. and Doryphoribius qinlingense sp. n., new species of eutardigrades from China 
X.C. LI, L.N. SU & D. YU (P.R. China)
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Zootaxa 638: 1-7 (13 September 2004) 2 plates; 19 references                        Accepted: 2 Sept. 2004
Macrobiotus dariae sp. n., a new species of eutardigrade (Eutardigrada, Macrobiotidae) from Cyprus
G. PILATO, R. BERTOLANI
(Italy)
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Zootaxa 576: 1-6 (19 July 2004)  1 plate; 6 references                                   Accepted: 9 July 2004
Notes on some tardigrades from South Africa, with the description of Diphascon (Diphascon) zaniewi sp. nov. (Eutardigrada: Hypsibiidae)
L. KACZMAREK & L. MICHALCZYK (Poland)

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Zootaxa 545: 1-7 (21 June 2004)  2 plates, 9 references                               Accepted: 9 June 2004
Doryphoribius neglectus sp. n. and Parascon nichollsae sp. n., new species of eutardigrades from Australia
G. PILATO & O. LISI
  (Italy)

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Zootaxa 406: 1-8 (16 January 2004) 4 plates; 5 references                Accepted: 17 December 2003
Macrobiotus kovalevi, a new species of Tardigrada from New Zealand (Eutadigrada, Macrobiotidae)
D. V. TUMANOV (Russia)

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2003

Zootaxa 331: 1-24 (21 October 2003) 21  plates; 7 references                   Accepted: 10 October 2003
A description of the new tardigrade Macrobiotus reinhardti (Eutardigrada: Macrobiotidae, harmsworthi group) with some remarks on the oral cavity armature within the genus Macrobiotus Schultze
L. MICHALCZYK & L. KACZMAREK (Poland)

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Zootaxa 314: 1-4 (1 October 2003) 1 plate; 18 references      Accepted: 26 September 2003
Thulinius, new generic name substituting for Thulinia Bertolani, 1981 (Tardigrada, Eutardigrada)
R. BERTOLANI (Italy)

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Zootaxa 306: 1-6 (24 September 2003) 1 plate, 7 references            Accepted: 16 September 2003
An opinion on the genus Fujiscon Ito, 1991 (Eutardigrada, Hypsibiidae)
G. PILATO & M.G. BINDA
(Italy)
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Zootaxa 258: 1-8 (4 August 2003) 5 plates, 5 references                           Accepted: 26 July 2003
Macrobiotus polonicus, a new species of Tardigrada from Poland (Eutardigrada: Macrobiotidae, ‘hufelandi group’)
G. PILATO (Italy), L. KACZMAREK (Poland), L. MICHALCZYK (Poland) & O. LISI  (Italy )
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Zootaxa 242: 1-24  (16 July 2003) 20 plates; 17 references                             Accepted: 1 July 2003
Water Bears (Tardigrada) of the Bieszczady Mountains (Poland, West Carpathians)
L. MICHALCZYK & L. KACZMAREK  (Poland)
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Zootaxa 241: 1-7  (16 July 2003) 1 plate; 10 references                             Accepted: 11 July 2003
Notes on some tardigrades from Central Africa, with the description of a new species of Hypsibiidae
G. PILATO, M.G. BINDA & O. LISI  (USA)
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Zootaxa 195: 1-8 (14 May 2003) 2 plates, 17 references                             Accepted: 2 May 2003
Remarks on some species of tardigrades from South America with the description of Minibiotus sidereus n. sp.
G. PILATO, M. G. BINDA & O. LISI (Italy)

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Zootaxa 177: 1-4 (3 April 2003) 5 references                          Accepted: 26 March 2003
New records of and key to Tardigrada from Costa Rica
L KACZMAREK (Poland)

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Zootaxa 140: 1-6 (31 January 2003) 2 plates; 6 9 references               Accepted: 24 January 2003
Hexapodibius christenberryae, a new species of tardigrade from North America (Eutardigrada, Calohypsibiidae)
G. PILATO & M.G. BINDA (Italy)

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Zootaxa 132: 1-8 (17 January 2003) 3 plates; 5 references            Accepted: 3 January 2003
Platicrista horribilis, a new species of Tardigrada (Eutardigrada:
Hypsibiidae) from Mongolia
L KACZMAREK  & L. MICHALCZYK (Poland)
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2002

Zootaxa 53: 1-4 (19 July 2002) 1 plate; 3 references                                        Accepted: 10 July 2002
Echiniscus barbarae, a new species of tardigrade from Cuba Island (Tardigrada: Heterotardigrada, Echiniscidae, ‘arctomys group’)
L. KACZMAREK & L. MICHALCZYK (Poland)

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Published on 3 Jun. 2009