HAN, TAEMAN, Applied Entomology Division, Department of Agricultural Biology, National Academy of Agricultural Science, Suin-ro 150, Gweonseon-gu, Suwon, 441-100 KOREA Division of Entomology, School of Agricultural Biotechnology, Seoul National University, San 51-1 Shillim-dong, Gwanak-gu, Seoul,151-742 KOREA