XU, KUIDONG, Laboratory of Marine Organism Taxonomy and Phylogeny, Institute of Oceanology, Chinese Academy of Sciences, Qingdao 266071, China 2Laboratory for Marine Biology and Biotechnology, Qingdao National Laboratory for Marine Science and Technology, Qingdao 266237, China. University of Chinese Academy of Sciences, Beijing 100039, China 5Center for Ocean Mega-Science, Chinese Academy of Sciences, Qingdao 266071, China, China