TIAN, CHUNPO, Guangdong Key Laboratory of Animal Conservation and Resource Utilization, Guangdong Public Laboratory of Wild Animal Conservation and Utilization, Institute of Zoology, Guangdong Academy of Sciences, Guangzhou, China. College of Life Sciences, Shaanxi Normal University, 620 West Chang’an Street, Chang’an District, Xi’an City, 710119, Shaanxi Province, China., China