Abstract
The first critically revised catalogue of the centipede fauna of the Paiko, Pinovo, Tzena and Voras mountains in northern Greece is presented. A total of 34 centipede species are listed, belonging to 13 genera, six families and four orders. The highest diversity was recorded within the order Lithobiomorpha (20 species), dominated by species of the genus Lithobius Leach, 1814 (14 species), followed by the order Geophilomorpha (10 species). The species Lithobius karamani Verhoeff, 1937 and Clinopodes skopljensis (Verhoeff, 1938) are here reported for the first time from Greece. The genera Scutigera, Cryptops, Himantarium, Bothriogaster, Henia and Geophilus were recorded for the first time in the fauna of the study area. Twenty-one out of the 34 species are newly reported from the studied mountainous area, including Scutigera coleoptrata, Eupolybothrus zeus, E. werneri, Lithobius forficatus, L. latro, L. lucifugus, L. muticus, L. peregrinus, L. crassipes, L. microps, Cryptops hortensis, C. parisi, Scolopendra cingulata, Himantarium gabrielis, Bothriogaster signata, Henia devia, H. illyrica, Clinopodes carinthiacus and Geophilus carpophagus. South European, European, Balkan, and endemic faunal elements predominate among the identified taxa. For each species, information is provided on distribution within the northern Greek mountain range, habitat preferences, altitudinal range, and relevant taxonomic notes. In addition, we assessed whether the sites where the species were recorded are included in the Natura 2000 Network; the conservation status of all species is reported, together with distribution maps for northwestern Greece.
References
- Andersson, G., Meidell, B.A., Scheller, U., Winqvist, J.-Å., Osterkamp Madsen, M., Djursvoll, P., Budd, G. & Gärdenfors, U. (2005) Nationalnyckeln till Sveriges flora och fauna. Mångfotingar. Myriapoda. ArtDatabanken. SLU, Uppsala.
- Aravanopoulos, F.A. (2010) The importance of the biodiversity of Hellenic forest and land-based ecosystems for Europe and the Mediterranean. Conference: Aristotle University Environmental Council Conference, 1–11. Thessaloniki, Greece.
- Attems, C.G. (1902) Myriapoden von Kreta, nebst Beiträgen zur allgemeinen Kenntnis einiger Gattungen. Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, Abteilung, I, 111 (1), 527–614.
- Attems, C.G. (1929) Myriapoda. In: Beier, M. (Ed.), Zoologische Forschungsreise nach den Ionischen Inseln und dem Peloponnes. Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften, Mathematisch-naturwissenschaftliche Klasse, Wien, 228, 403–470.
- Attems, C. (1947) Neue Geophilomorpha des Wiener Museums. Annalen des Naturhistorischen Museums in Wien, 55, 50–149.
- Attems, C. (1959) Die Myriopoden der Höhlen der Balkanhalbinsel. Nach dem Material der “Biospeologica balcanica”. Annalen des Naturhistorischen Museums in Wien, 63, 281–406.
- Baba, Ș.C., Giurginca, A., Plăiașu, R., Opran, R., Mîrleneanu, A. & Pacioglu, O. (2025) The troglophilic population of Lithobius peregrinus Latzel, 1880 (Chilopoda, Lithobiomorpha) from Gaura cu Muscă Cave – first record of the species in Romania: morphological variations, asymmetry, teratology and behaviour. Subterranean Biology, 52, 69–102. https://doi.org/10.3897/subtbiol.52.154132
- Barber, A.D. (2009) Centipedes: Keys and notes for the identification of British species (Synopses of the British Fauna, New Series No. 58). Field Studies Council, Shrewsbury, UK.
- Bonato, L. & Minelli, A. (2011) Geophilus arenarius, a long-misunderstood species in the still unresolved carpophagus species-complex (Chilopoda: Geophilidae). Zootaxa, 3114, 40–49. https://doi.org/10.11646/zootaxa.3114.1.4
- Bonato, L. & Zapparoli, M. (2011) Chilopoda—Geographical Distribution. In: Minelli, A. (Ed.), The Myriapoda. Vol. 1. Treatise on Zoology - Anatomy, Taxonomy, Biology. Brill, Leiden, pp. 327–337. https://doi.org/10.1163/9789004188266_017
- Bonato, L., Edgecombe, G.D., Lewis, J.G.E., Minelli, A., Pereira, L.A., Shelley, R.M. & Zapparoli, M. (2010) A common terminology for the external anatomy of centipedes (Chilopoda). ZooKeys, 69, 17–51. https://doi.org/10.3897/zookeys.69.737
- Bonato, L., Iorio, É. & Minelli, A. (2011) The centipede genus Clinopodes C. L. Koch, 1847 (Chilopoda, Geophilomorpha, Geophilidae): reassessment of species diversity and distribution, with a new species from the Maritime Alps (France). Zoosystema, 33 (2), 175–205. https://doi.org/10.5252/z2011n2a3
- Bonato, L., Dányi, L., Socci, A.A. & Minelli, A. (2012) Species diversity of Strigamia (Chilopoda: Linotaeniidae): a preliminary synthesis. Zootaxa, 3593, 1–39. https://doi.org/10.11646/zootaxa.3593.1.1
- Bonato, L., Minelli, A., Lopresti, M. & Cerretti, P. (2014) ChiloKey, an interactive identification tool for the geophilomorph centipedes of Europe (Chilopoda, Geophilomorpha). ZooKeys, 443, 1–9. https://doi.org/10.3897/zookeys.443.7530
- Bonato, L., Chagas Junior, A., Edgecombe, G.D., Lewis, J.G.E., Minelli, A., Pereira, L.A., Shelley, R.M., Stoev, P. & Zapparoli, M. (2016) ChiloBase 2.0 - A World Catalogue of Centipedes (Chilopoda). Available from: https://chilobase.biologia.unipd.it (accessed 30 July 2025) https://doi.org/10.3897/zookeys.69.737.figures41-48
- Bonato, L., Bortolin, F., De Zen, G., Decker, P., Lindner, E.N., Orlando, M., Spelda, J., Voigtländer, K. & Wesener, T. (2023) Towards elucidating species diversity of European inland Strigamia (Chilopoda: Geophilomorpha): a first reassessment integrating multiple lines of evidence. Zoological Journal of the Linnean Society, 199, 945–966. https://doi.org/10.1093/zoolinnean/zlad070
- Brölemann, H.W. (1909) À propos d’un système des géophilomorphes. Archives de Zoologie expérimentale et générale, (5) 3, 303–340.
- Dafis, S., Papastergiadou, E., Georgiou, K., Babalonas, D., Georgiadis, T., Papageorgiou, M., Lazaridou, E. & Tsiaousi, V. (2014) The Greek Habitat Project (Directive 92/43/EEC) NATURA 2000: An overview. LIFE contract B4-3200/94/756.
- Dányi, L. (2006) On the occurrence of Lithobius erythrocephalus C. L. Koch, 1847, and Lithobius schuleri Verhoeff, 1925 (Myriapoda: Chilopoda) in Hungary. Folia historico-naturalia Musei Matraensis, 30, 105–113. https://doi.org/10.69595/folianatmatra
- Eason, E.H. (1964) Centipedes of the British Isles. Warne, London, 294 pp.
- Eason, E.H. (1970) A redescription of the species of Eupolybothrus Verhoeff s. str. preserved in the British Museum (Natural History) and the Hope departement of Zoology, Oxford (Chilopoda Lithobiomorpha). Bulletin of the British Museum (Natural History), Zoology, 19, 289–310. https://doi.org/10.5962/p.117867
- Eason, E.H. (1972) The type specimens and identity of the species described in the genus Lithobius by C.L. Koch and L. Koch from 1841 to 1878 (Chilopoda: Lithobiomorpha). Bulletin of the British Museum (Natural History), Zoology, 22, 103–150.
- Eason, E.H. (1982) A review of the north-west European species of Lithobiomorpha. Zoological Journal of the Linnean Society, 74, 9–112.
- Edgecombe, G.D. (2011) Chilopoda – the fossil history. In: Minelli, A. (Ed.), Treatise on zoology – anatomy, taxonomy, biology. The Myriapoda. Vol. 1. Brill Academic Publishers, Leiden, pp. 355–361. https://doi.org/10.1163/9789004188266_019
- Edgecombe, G.D. & Giribet, G. (2006) A century later – a total evidence re-evaluation of the phylogeny of scutigeromorph centipedes (Myriapoda: Chilopoda). Invertebrate Systematics, 20 (5), 503–525. https://doi.org/10.1071/IS05044
- European Commission (1992) Council Directive 92/43/EEC of 21 May 1992 on the conservation of natural habitats and of wild fauna and flora. Official Journal of the European Union, 206, 7–50.
- European Commission (2009) Directive 2009/147/EC of the European Parliament and of the Council of 30 November 2009 on the conservation of wild birds. Official Journal of the European Union, 20, 7–25.
- European Commission (2020) Natura 2000. Available from: https://ec.europa.eu/environment/nature/natura2000/index_en.htm (accessed 11 May 2026)
- Foddai, D., Minelli, A. & Zapparoli, M. (1996) I chilopodi delle isole circumsarde nel contesto del popolamento insulare dell’area tirrenica s.l. Biogeographia. The Journal of Integrative Biogeography, 18, 357–376. https://doi.org/10.21426/b618110444
- Georgiakakis, P., Kafkaletou Diez, A., Salvarina, I., Benda, P., Billington, G., Dietz, C., Billington, J., Cove, D., Davison, S., Cooke, M. & Papadatou, E. (2023) The Bats of Greece: An Updated Review of Their Distribution, Ecology and Conservation. Animals, 13 (15), 2529. https://doi.org/10.3390/ani13152529
- Georgopoulou, E., Kougioumoutzis, K. & Simaiakis, S.M. (2025) The Impact of Climate and Land Use Change on Greek Centipede Biodiversity and Conservation. Land, 14 (8), 1685. https://doi.org/10.3390/land14081685
- Gregnanin, L. & Bonato, L. (2024) A comprehensive dataset of the geophilid centipedes of the south-eastern Alps (Chilopoda, Geophilomorpha, Geophilidae s.l.). Biodiversity Data Journal, 12, e122144. https://doi.org/10.3897/BDJ.12.e122144
- Hellenic Ornithological Society (2023) Setting conservation objectives for avifauna species of community importance per Special Protection Area of the Natura 2000 Network. LIFE-IP 4 NATURA: Integrated actions for the conservation and management of Natura 2000 sites, species, habitats and ecosystems in Greece (LIFE16 IPE/GR/000002). Deliverable for Action C.7: Establishment of conservation objectives for SACs and SPAs. Athens, 19 pp.
- Iorio, É. (2014) Catalogue biogéographique et taxonomique des chilopodes (Chilopoda) de France métropolitaine. Mémoires de la Société Linnéenne de Bordeaux, 15, 1–372.
- Iorio, É. & Jacquemin, G. (2025) Les Chilopodes d’Auvergne-Rhône-Alpes et de Provence-Alpes-Côte d’Azur (Chilopoda): guide pratique d’identification. Ouvrage finalisé pour le Pôle Invertébrés d’Auvergne-Rhône-Alpes, 172 pp.
- Iorio, É., Carnet, M., Cherpitel, T., Desmots, D., Geoffroy, J.-J., Jacquemin, G., Quindroit, C. & Racine, A. (2023) Les Chilopodes de France métropolitaine (Myriapoda, Chilopoda): liste commentée des espèces avec état des connaissances et proposition de noms français. Naturae, 1, 1–20. https://doi.org/10.5852/naturae2023a1
- Jeekel, C.A.W. (1964) Beitrag zur Kenntnis der Systematik und Okologie der Hundertfusser (Chilopoda) Nordwestdeutschlands. Abhandlungen und Verhandlungen des Naturwissenschaftlichen Vereins in Hamburg, 8, 111–153.
- Jeekel, C.A.W. (1967) On two Italian Lithobius species described by Silvestri, with taxonomic notes on the genus Eupolybothrus Verhoeff (Chilopoda, Lithobiida). Beaufortia, 14 (175), 165–175.
- Kaltsas, D. & Simaiakis, S.M. (2012) Seasonal patterns of activity of Scolopendra cretica and S. cingulata (Chilopoda: Scolopendromorpha) in East Mediterranean maquis ecosystems. International Journal of Myriapodology, 7, 1–14. https://doi.org/10.3897/ijm.7.2133
- Koch, L. (1867) Zur Arachniden und Myriapoden-fauna Süd-Europa’s. Verhandlungen der Kaiserlich-Königlichen Zoologisch-Botanischen Gesellschaft in Wien, 17, 857–900.
- Kos, I. (1992) A review of the European species of Strigamia. Acta Zoologica Hungarica, 38, 1–22.
- Kos, A., Delić, T., Kos, I., Kozel, P., Polak, S. & Zagmajster, M. (2023) The overview of lithobiomorph centipedes (Chilopoda, Lithobiomorpha) from caves of Slovenia. Subterranean Biology, 45, 165–185. https://doi.org/10.3897/subtbiol.45.101430
- Latzel, R. (1880) Die Myriopoden der Österreichisch-Ungarischen Monarchie. A. Holder, Wien, 1–414.
- Legakis, A., Constantinidis, T. & Petrakis, P.V. (2018) Biodiversity in Greece. In: Pullaiah, T. (Ed.), Global biodiversity. Apple Academic Press Inc., London, pp. 71–124. https://doi.org/10.1201/9780429487750-4
- Legakis, A. & Maragkou, P. (2009) The red book of threatened species. Hellenic Zoological Society, Athens.
- Lewis, J.G.E. (2010) A key and annotated list of the Scolopendra species of the Old World with a reappraisal of Arthrorhabdus (Chilopoda: Scolopendromorpha: Scolopendridae). International Journal of Myriapodology, 3, 83–122. https://doi.org/10.1163/187525410X12578602960380
- Lewis, J.G.E. (2011) A review of the species of the genus Cryptops Leach, 1815 from the Old World related to Cryptops (Cryptops) hortensis (Donovan, 1810) (Chilopoda, Scolopendromorpha). International Journal of Myriapodology, 4, 11–50. https://doi.org/10.3897/ijm.4.1116
- Matic, Z. (1966) Clasa Chilopoda, subclasa Anamorpha. Fauna Republicii Socialiste România, 6, 1–272.
- Matic, Z. (1980) Chilopodes collected on the island of Crete, Greece. Acta Zoologica Bulgarica, 15, 99–102.
- Meinert, F. (1868) Danmarks scolopender og lithobier. Naturhistorisk Tidsskrift, 3rd series, 5, 241–268.
- Minelli, A. (1982) Contributo alla revisione dei chilopodi geofilomorfi finora riferiti ai generi Henia e Chaetechelyne (Chilopoda, Geophilomorpha). Memorie della Società Entomologica Italiana, 60, 253–268.
- Minelli, A. & Bonato, L. (2014) Chilopoda Geophilomorpha of Europe: a revised list of species, with taxonomic and nomenclatorial notes. Zootaxa, 3770 (1), 1–136. https://doi.org/10.11646/zootaxa.3770.1.1
- Mitić, B.M. (2002) On the diversity of centipedes (Chilopoda, Myriapoda) in Serbia. Part two. Archives of Biological Sciences, 54 (1–2), 13P–14P.
- Mitić, B.M., Ilić, B.S. & Makarov, S.E. (2012) First record of Cryptops parisi Brolemann, 1920 (Chilopoda: Scolopendromorpha: Cryptopidae) in Serbia. Kragujevac Journal of Science, 34, 97–100.
- Mitić, B.M., Živić, N.V., Stojanović, D.Z., Antić, D.Ž., Ilić, B.S. & Makarov, S.E. (2014) The centipedes (Chilopoda) of Serbia (including Kosovo and Metohija). In: Tuf, I.H. & Tajovský, K. (Eds.), 16th International Congress of Myriapodology, 20–25 July 2014, Olomouc, Czech Republic. Book of Abstracts, pp. 56. https://doi.org/10.3897/zookeys.510.10176.figure1
- Mountrakis, D. (2010) Geology and Geotectonic Evolution of Greece. University Studio Press, Thessaloniki. [in Greek]
- Mucina, L., Bültmann, H., Dierßen, K., Theurillat, J.-P., Raus, T., Čarni, A., Šumberová, K., Willner, W., Dengler, J., García, R.G., Chytrý, M., Hájek, M., Di Pietro, R., Iakushenko, D., Pallas, J., Daniëls, F.J.A., Bergmeier, E., Santos Guerra, A., Ermakov, N., Valachovič, M., Schaminée, J.H.J., Lysenko, T., Didukh, Y.P., Pignatti, S., Rodwell, J.S., Capelo, J., Weber, H.E., Solomeshch, A., Dimopoulos, P., Aguiar, C., Hennekens, S.M. & Tichý, L. (2016) Vegetation of Europe: Hierarchical floristic classification system of vascular plant, bryophyte, lichen, and algal communities. Applied Vegetation Science, 19 (1), 3–264. https://doi.org/10.1111/avsc.12257
- Natural Environment and Climate Change Agency (NECCA) (2024) The Greek Red List of Threatened Species. Athens, Greece. Available from: redlist.necca.gov.gr (accessed 30 August 2025)
- Negrea, Ș. (2006) A catalogue to the Lithobiida, Scutigerida and Scolopendrida species (Myriapoda: Chilopoda) of Romania. Travaux du Muséum National d’Histoire Naturelle “Grigore Antipa”, 49, 93–118.
- Papp, B. & Tzakiri, E. (2017) Contributions to the bryophyte flora of the Paiko, Tzena and Pinovo mountains in Greece. Studia Botanica Hungarica, 48 (1), 33–49. https://doi.org/10.17110/studbot.2017.48.1.33
- Paragamian, K., Poulinakis, M., Paragkamian, S. & Nikoloudakis, I. (2025) Cave fauna of Greece database - Hellenic Institute of Speleological Research. Available from: https://database.inspee.gr/ (accessed 27 August 2025)
- Paragamian, K. (2025) Eupolybothrus macedonicus (Verhoeff, 1943). In: Paragamian, K., Poulinakis, M., Paragamian, S. & Nikoloudakis, I., Cave fauna of Greece database - Hellenic Institute of Speleological Research. Species data updated 01.06.2021 (accessed 27 August 2025).
- Paragamian, K. (2025) Lithobius auritus (Verhoeff, 1943). In: Paragamian, K., Poulinakis, M., Paragamian, S. & Nikoloudakis, I., Cave fauna of Greece database - Hellenic Institute of Speleological Research. Species data updated 16.02.2018 (accessed 27 August 2025).
- Paragamian, K. (2025) Lithobius temnensis Verhoeff, 1943. In: Paragamian, K., Poulinakis, M., Paragamian, S. & Nikoloudakis, I., Cave fauna of Greece database - Hellenic Institute of Speleological Research. Species data updated 04.11.2020 (accessed 27 August 2025).
- Peretti, E., Cecchin, C., Fusco, G., Gregnanin, L., Kos, I. & Bonato, L. (2022) Shedding light on species boundaries in small endogeic animals through an integrative approach: species delimitation in the centipede Clinopodes carinthiacus (Chilopoda: Geophilidae) in the south-eastern Alps. Zoological Journal of the Linnean Society, 196, 902–923. https://doi.org/10.1093/zoolinnean/zlac008
- Popovici, G. (2024) A review of the genus Pachymerium C. L. Koch, 1847 (Geophilomorpha: Geophilidae) from Romania. Zootaxa, 5406 (4), 588–600. https://doi.org/10.11646/zootaxa.5406.4.7
- Rémy, P. (1951) Description de grottes de Macédoine grecque. Annales de Spéléologie, 6 (2–3), 107–118.
- Sfenthourakis, S., Pafilis, P., Parmakelis, A., Poulakaelis, P. & Triantis, K. (2018) Biogeography and biodiversity of the Aegean: in honour of Prof. Moysis Mylonas. Broken Hill Publishers Ltd, Nicosia.
- Simaiakis, S., Minelli, A. & Mylonas, M. (2004) The centipede fauna (Chilopoda) of Crete and its satellite islands (Greece, Eastern Mediterranean). Israel Journal of Zoology, 50, 367–418. https://doi.org/10.1560/hbe1-qjer-ydkf-bd8q
- Simaiakis, S., Minelli, A. & Mylonas, M. (2005) The Centipede Fauna (Chilopoda) of the South Aegean Archilpelgo (Greece, Eastern Mediterranean). Israel Journal of Zoology, 51 (4), 241–307. https://doi.org/10.1560/43yf-y0jl-j13p-4520
- Simaiakis, S. & Mylonas, M. (2006) Intraspecific variation in segment number in Pachymerium ferrugineum (C. L. Koch, 1835) (Chilopoda: Geophilomorpha) in the south Aegean Archipelago (NE Mediterranean, Greece). Biological Journal of the Linnaean Society, 88, 533–539. https://doi.org/10.1111/j.1095-8312.2006.00635.x
- Simaiakis, S., Voulgaris, G. & Mylonas, M. (2007) A revised catalogue of the centipedes (Chilopoda) of the North Aegean Archipelago with particular reference to the islands of Híos, Límnos and Skyros. Belgian Journal of Zoology, 137 (2), 243–246.
- Simaiakis, S. & Mylonas, M. (2008) The Scolopendra species (Chilopoda: Scolopendromorpha: Scolopendridae) of Greece (E-Mediterranean): a theoretical approach on the effect of geography and palaeogeography on their distribution. Zootaxa, 1792, 39–53. https://doi.org/10.11646/zootaxa.1792.1.3
- Simaiakis, S.M., Dimopoulou, A., Mitrakos, A., Mylonas, M. & Parmakelis, A. (2012) The evolutionary history of the Mediterranean centipede Scolopendra cingulata (Latreille, 1829) (Chilopoda: Scolopendridae) across the Aegean archipelago. Biological Journal of the Linnean Society, 105 (3), 507–527. https://doi.org/10.1111/j.1095-8312.2011.01813.x
- Simaiakis, S.M., Zapparoli, M., Minelli, A. & Bonato, L. (2013) The centipede fauna (Chilopoda) of the island of Cyprus, with one new lithobiomorph species. Zootaxa, 3647 (2), 279–306. https://doi.org/10.11646/zootaxa.3647.2.3
- Simaiakis, S.M. & Strona, G. (2015) Patterns and processes in the distribution of European centipedes (Chilopoda). Journal of Biogeography, 42 (6), 1018–1028. https://doi.org/10.1111/jbi.12463
- Simaiakis, S., Akkari, N. & Zapparoli, M. (2016) The centipedes of Peloponnisos and first records of genus Eurygeophilus in the East Mediterranean. Zootaxa, 4061 (4), 301–346. https://doi.org/10.11646/zootaxa.4061.4.1
- Spiliopoulou, K., Dimitrakopoulos, P.G., Brooks, T.M., Kelaidi, G., Paragamian, K., Kati, V., Oikonomou, A., Vavylis, D., Trigas, P., Lymberakis, P., Darwall, W., Stoumboudi, M.T. & Triantis, K. (2021) The Natura 2000 network and the ranges of threatened species in Greece. Biodiversity and Conservation, 30, 945–961. https://doi.org/10.1007/s10531-021-02125-7
- Stagl, V. & Zapparoli, M. (2006) Type specimens of the Lithobiomorpha (Chilpoda) in the natural History Museum in Vienna. Kataloge der Wissenschaftlichen Sammlungen des Naturhistorischen Museums, Wien, 21 (3), 49 pp.
- Stoev, P. (1997) The centipede fauna of Albania (Chilopoda). Historia Naturalis Bulgarica, 8, 101–117.
- Stoev, P. (2000) On centipedes (Chilopoda) of Albania, 2. Arthropoda Selecta, 9, 199–206.
- Stoev, P. (2001a) On the centipedes (Chilopoda) of the Republic of Macedonia. Historia Naturalis Bulgarica, 13, 93–107.
- Stoev, P. (2001b) A synopsis of the Bulgarian cave centipedes (Chilopoda). Arthropoda Selecta, 10 (1), 31–54.
- Stoev, P. (2002) A catalogue and key to the centipedes (Chilopoda) of Bulgaria (Series Faunistica No. 25). Pensoft Publishers, Sofia–Moscow.
- Stoev, P. (2005) On the identity of some poorly known lithobiid centipedes described by Karl Verhoeff (Chilopoda: Lithobiomorpha). Zootaxa, 796, 1–12. https://doi.org/10.11646/zootaxa.796.1.1
- Stoev, P., Akkari, N., Zapparoli, M., Porco, D., Enghoff, H., Edgecombe, D., Georgiev, T. & Penev, L. (2010) The centipede genus Eupolybothrus Verhoeff, 1907 (Chilopoda: Lithobiomorpha: Lithobiidae) in North Africa, a cybertaxonomic revision, with a key to all species in the genus and the first use of DNA barcoding for the group. ZooKeys, 50, 29–77. https://doi.org/10.3897/zookeys.50.504
- Stojanović, D., Stankovic, M., Šćiban, M., Šević, M. & Antić, D. (2022) The centipedes fauna (Arthropoda: Myriapoda) of the “Zasavica” Special Nature Reserve. In: Naučno-stručni skup o biodiverzitetu i drugim vrednostima rezervata Zasavica “Zasavica 2022”, pp. 133–144.
- Tsipidou, O., Leinemann, L., Korakis, G., Finkeldey, R., Gailing, O. & Papageorgiou, A.C. (2021) Fine-Scale Spatial Patterns of the Genetic Diversity of European Beech (Fagus sylvatica L.) around a Mountainous Glacial Refugium in the SW Balkans. Forests, 12 (6), 725. https://doi.org/10.3390/f12060725
- Tuf, I.H. & Tajovský, K. (2016) An annotated checklist of the centipedes (Chilopoda) recorded in the Czech Republic. Acta Societatis Zoologicae Bohemicae, 80, 45–50.
- Vesović, N., Deltshev, C., Mitov, P., Antić, D., Stojanović, D.Z., Stojanović, D.V., Stojanović, K., Božanić, M., Ignjatović-Ćupina, A. & Ćurčić, S. (2024) The Diversity of Subterranean Terrestrial Arthropods in Resava Cave (Eastern Serbia). Diversity, 16 (4), 234. https://doi.org/10.3390/d16040234
- Verhoeff, K.W. (1899) Beiträge zur Kenntnis paläarktischer Myriopoden. XI. Aufsatz: Neue und wenig bekannte Lithobiiden. Verhandlungen der Zoologisch-Botanischen Gesellschaft in Wien, 49, 451–459. https://doi.org/10.5962/bhl.part.24105
- Verhoeff, K.W. (1901) Beiträge zur Kenntnis der Geophiliden. Zoologischer Anzeiger, 24, 417–454.
- Verhoeff, K.W. (1928) Geophilomorphen-Beiträge und eine Lithobius-Form. Mitteilungen aus dem Zoologischen Museum, Berlin, 14, 229–286. https://doi.org/10.1002/mmnz.4830140202
- Verhoeff, K.W. (1937) Zur Kenntnis der Lithobiiden. Archiv für Naturgeschichte, N.F. 6, 171–257.
- Verhoeff, K.W. (1938) Chilopoden-Studien, zur Kenntnis der Epimorphen. Zoologische Jahrbücher, Abteilung für Systematik, Ökologie und Geographie der Tiere, 71, 339–388.
- Verhoeff, K.W. (1943) Über neue und wenig bekannte Myriopoden aus Palästina. Zoologischer Anzeiger, 114, 89–104.
- Vigna Taglianti, A., Audisio, P., Biondi, M., Bologna, M.A., Carpaneto, G.M., De Biase, A., Fattorini, S., Piattella, E., Sindaco, R., Venchi, A. & Zapparoli, M. (1999) A proposal for a chorotype classification of the Near East fauna, in the framework of the Western Palearctic region. Biogeographia, Lavori della Società Italiana di Biogeografia, 20, 31–59. https://doi.org/10.21426/b6110172
- Voigtländer, K. (2011) Chilopoda – ecology. In: Minelli, A. (Ed.), Treatise on zoology – anatomy, taxonomy, biology. The Myriapoda. Vol. 1. Brill Academic Publishers, Leiden, pp. 309–325. https://doi.org/10.1163/9789004188266_016
- Voigtländer, K., Reip, H.S., Decker, P. & Spelda, J. (2011) Critical reflections on German Red Lists of endangered myriapod species (Chilopoda, Diplopoda) (with species list for Germany). International Journal of Myriapodology, 6, 85–105. https://doi.org/10.3897/ijm.6.2175
- Vougioukalakis, G. (2002) Petrological, geochemical, volcanological study of the Almopias Pliocene volcanic formations and their correlation with the geothermal manifestations in the area. Ph.D Thesis, Aristotle University of Thessaloniki, 303 pp.
- Zapparoli, M. (1984) Note su alcune specie del gen. Eupolybothrus della fauna di Grecia (Chilopoda, Lithobiomorpha). Fragmenta Entomologica, 17 (2), 195–209.
- Zapparoli, M. (1993) Note faunistiche sui Chilopodi della Grecia. Fragmenta Entomologica, 25, 93–102.
- Zapparoli, M. (1994) Chilopodi di alcune località della Grecia. Fragmenta Entomologica, 26, 13–22.
- Zapparoli, M. (1996) Distribution patterns and qualitative composition of the centipede fauna in forestal habitats of mainland Greece. In: Geoffroy, J.-J., Mauriès, J.-P. & Nguyen Duy-Jacquemin, M. (Eds.), Acta Myriapodologica. Mémoires du Muséum National d’Histoire Naturelle, 169, 599–605.
- Zapparoli, M. (2002) A catalogue of the centipedes (Chilopoda) of Greece. Fragmenta Entomologica, 34, 1–118.
- Zapparoli, M. (2003) The present knowledge on the European fauna of Lithobiomorpha (Chilopoda). Bulletin of the British Myriapod and Isopod Group, 19, 20–41.
- Zapparoli, M. (2006) A catalogue of the centipedes (Chilopoda) of the Central Apennines (Italy). Bollettino del Museo Civico di Storia Naturale di Verona (Botanica Zoologia), 30, 165–274.
- Zapparoli, M. (2009) An annotated catalogue of the epigeic and cave centipedes (Chilopoda) of Sardinia. Zootaxa, 2318, 56–168. https://doi.org/10.11646/zootaxa.2318.1.6
