Skip to main content Skip to main navigation menu Skip to site footer
Responsive image
Issue: Vol. 5810 No. 1: 14 May 2026
Type: Article
Published: 2026-05-14
DOI: 10.11646/zootaxa.5810.1.1
Page range: 1-48
Abstract views: 159
PDF downloaded: 12

Pseudagrion microcephalum (Rambur, 1842)—sifting through a putative familiar species, with erection of two new species (Odonata: Coenagrionidae)

Zoologisches Museum der CAU zu Kiel; Hegewischstr. 3; 24105 Kiel; Germany
National Museum of Natural History; Bulgarian Academy of Sciences Tsar Osvoboditel Blvd. 1; 1000 Sofia; Bulgaria
Odonata Damselfly neotype taxonomy nomenclature Pseudagrion crenatum Pseudagrion pirata Pseudagrion siamense Pseudagrion coarctatum Pseudagrion daponshanensis

Abstract

Specimens of the Pseudagrion blue-group allied to P. microcephalum (Rambur, 1842) were studied with special attention to the male caudal appendages and genital ligulae. The males of P. microcephalum are morphologically re-characterised, and the species is considered to have a limited range that includes India, Pakistan, Nepal, Bangladesh, and Sri Lanka. The status of P. siamense Fraser, 1922 is revised and considered a distinct species, with the scientific name removed from synonymity and a neotype designated and described (♂, 7.viii.1972, stream at Ang Kep Nam-Bang Phra environments, South Chon Buri, Thailand, approximately 13.205°N, 100.975°E, G. von Rosen leg., GvRosen no. 1753; deposited at the Zoological State Collection Munich, Germany). Two new species are erected and figured: P. crenatum sp. nov. from Australia (holotype ♂, 2.x.1915, Caboolture, Queensland, Australia, approximately -27.084°N, 152.951°E, R.J. Tillyard leg., no. 14110; deposited at the Forschungsinstitut und Naturmuseum Senckenberg, Frankfurt am Main, Germany); and P. pirata sp. nov. from the Philippine Islands (holotype ♂, 17.i.1992, Barili, Montayupan falls, Cebu Island, Philippines, approximately 10.101°N, 123.535°E, Th. Borromeo Jr. leg., ex. Coll. Roland A. Müller, no. 975694; deposited at the Naturalis Biodiversity Centre Leiden, the Netherlands). The range of P. coarctatum Lieftinck, 1932, a species previously known only from a few records at Jayapura and Sentani Lake in northeastern Indonesian West Papua, is extended to Sulawesi, the Moluccas and the Solomon Islands; the male is morphologically re-characterised and illustrated, and the female described. Photographs of the holotypes of P. microcephalum, P. coarctatum and P. starreanum Lieftinck, 1949 are provided. In addition, P. stainbergerorum Marinov, 2012 stat. nov. is proposed, based on distinct morphological differences from P. microcephalum. A key to males of the Pseudagrion blue-group is provided, including illustrations of the male caudal appendages and partly of the genital ligulae for all species treated. Pseudagrion daponshanensis Zhou & Zhou, 2007 is transferred to the genus Coenagrion.

 

References

  1. Asahina, S. (1968) Records and notes on Philippine Odonata. Japanese Journal of Zoology, 15 (4), 349–376, pls. 1–2.
  2. Asahina, S. (1982) A list of the Odonata from Thailand. Part 1. Agrionidae. Kontyu, 50 (3), 454–466.
  3. Bárta, D. & Dolný, A. (2013) Dragonflies of Sungai Wain. Ecological Field Guide to the Odonata of Lowland Mixed Dipterocarp Forest of Southeastern Kalimantan. Taita Publishers, Hradec Králové, 168 pp.
  4. Brauer, F. (1869) Beschreibung neuer Neuropteren aus dem Museum Godeffroy in Hamburg. Verhandlungen der kaiserlich-königlichen zoologisch-botanischen Gesellschaft in Wien, 19, 9–18.
  5. Burmeister, H. (1839) Handbuch der Entomologie 2. Theod. Chr. Fried. Enslin, Berlin, pp. 757–1050.
  6. Campion, H. (1921) A note on some dragonflies from Amboina. The Entomologist, 54 (702), 262–264.
  7. Dow, R.A. & Wilson, K.D.P (2017) Pseudagrion microcephalum. The IUCN Red List of Threatened Species, 2017, e.T167199A83376119. https://doi.org/10.2305/IUCN.UK.2017-1.RLTS.T167199A83376119.en
  8. Dow, R.A., Choong, C.Y., Grinang, J., Lupiyaningdyah, P., Ngiam, R.W.J. & Kalkman, V. (2024) Checklist of the Odonata (Insecta) of Sundaland and Wallacea (Malaysia, Singapore, Brunei, Indonesia and Timor Leste). Zootaxa, 5460 (1), 1–122. https://doi.org/10.11646/zootaxa.5460.1.1
  9. Dumont, H.J. (2004) Distinguishing between the East-Asiatic representatives of Paracercion Weekers & Dumont (Zygoptera: Coenagrionidae). Odonatologica, 33 (4), 361–370.
  10. Fraser, F.C. (1922a) New and rare Indian Odonata in the Pusa collection. Memoirs of the Department of Agriculture in India, 7, 39–77.
  11. Fraser, F.C. (1922b) A collection of dragonflies from Bangkok. The Journal of the Natural History Society of Siam, 4, 231–238, pl. 10.
  12. Fraser, F.C. (1924a) Report on a collection of dragonflies (Odonata) from the Andaman Islands. Records of the Indian Museum, 26 (5), 409–413, pl. 24. https://doi.org/10.26515/rzsi/v26/i5/1924/162658
  13. Fraser, F.C. (1924b) A survey of the odonata (Dragonfly) fauna of western India with special remarks on the genera Macromia and Idionyx and descriptions of thirty new species. Records of the Indian Museum, 26 (5), 423–522, pls. 25–27. https://doi.org/10.26515/rzsi/v26/i5/1924/162660
  14. Fraser, F.C. (1925) A preliminary report on the Dragonflies (Order Odonata) of Samoa. Transactions of the Entomological Society of London, 1924 (3/4), 429–438. https://doi.org/10.1111/j.1365-2311.1925.tb03364.x
  15. Fraser, F.C. (1927) Notes on a collection of dragonflies (Odonata) from Siam. The Journal of the Siam Society, Natural History Supplement, 7 (2), 83–92.
  16. Fraser, F.C. (1933) The fauna of British India, including Ceylon and Burma. Odonata. Vol. 1. Taylor and Francis, London, 423 pp.
  17. Garrison, R.W., von Ellenrieder, N. & Louton, J.A. (2010) Damselfly Genera of the New World. An illustrated and annotated key to the Zygoptera. Johns Hopkins University Press, Baltimore, Maryland, 490 pp.
  18. Gassmann, D. (2011) Pseudagrion lorenzi sp. nov., a new damselfly species from New Britain Island, Papua New Guinea (Odonata: Coenagrionidae). International Journal of Odonatology, 14 (2), 149–162. https://doi.org/10.1080/13887890.2011.595650
  19. Hämäläinen, M. & Müller, R.A. (1997) Synopsis of the Philippine Odonata, with lists of species recorded from forty islands. Odonatologica, 26 (3), 249–315.
  20. Hämäläinen, M. & Pinratana, B.A. (1999) Atlas of the dragonflies of Thailand. Distribution maps by provinces. Chok Chai Creation Printing Group Co., Ltd., Bangkok, 176 pp.
  21. ICZN [International Commission on Zoological Nomenclature] (1999) International Code of Zoological Nomenclature. 4th Edition. The International Trust for Zoological Nomenclature, London, xxix + 306 pp.
  22. Ishida, S., Ishida, K., Kojima, K. & Sugimura, M. (1988) Illustrated guide for identification of the Japanese Odonata. Tokai University Press, Tokyo, 328 pp.
  23. Kalkman, V.J. & Orr, A.G. (2013) Field Guide to the damselflies of New Guinea. Brachytron, 16 Supplement, 3–120.
  24. Kalkman, V.J., Babu, R., Bedjanič, M., Conniff, K., Gyeltshen, T., Khan, M.K., Subramanian, K.A., Zia, A. & Orr, A.G. (2020) Checklist of the dragonflies and damselflies (Insecta: Odonata) of Bangladesh, Bhutan, India, Nepal, Pakistan and Sri Lanka. Zootaxa, 4849 (1), 1–84. https://doi.org/10.11646/zootaxa.4849.1.1
  25. Kimmins, D.E. (1966) A list of the Odonata types described by F.C. Fraser, now in the British Museum (Natural History). Bulletin of the British Museum (Natural History), 18 (6), 175–227.
  26. Kirby, W.F. (1890) A synonymic catalogue of Neuroptera Odonata or dragonflies, with an appendix to fossil species. Gurney & Jackson, London and Friedländer & son, Berlin, 202 pp. https://doi.org/10.5962/bhl.title.5534
  27. Kirby, W.F. (1891) On some Neuroptera Odonata (Dragonflies) collected by Mr. E.E. Green in Ceylon. Proceedings of the general meetings for scientific business of the Zoological Society of London, 1891 (2), 203–206, pl. 20. https://doi.org/10.1111/j.1096-3642.1891.tb01741.x
  28. Kosterin, O.E. (2011) Odonata of the Cambodian coastal regions revisited: beginning of dry season in 2010. International Dragonfly Fund-Report, 40, 1–108.
  29. Kovács, T., Theischinger, G., Juhász, P. & Danyik, T. (2015) Odonata from Batanta (Indonesia, West Papua) with description of three new species. Folia Historico-Naturalia Meusei Matraensis, 39, 17–29.
  30. Kovács, T., Theischinger, G., & Danyik, T. (2016) Odonata from Batanta (Indonesia, West Papua) with description of two new species. Folia Historico-Naturalia Meusei Matraensis, 40, 27–37.
  31. Laidlaw, F.F. (1915) Fauna of the Chilka Lake. Order Odonata. Family Agrionidae. Memoirs of the Indian Museum, 5, 178–180.
  32. Laidlaw, F.F. (1919) A list of the dragonflies recorded from the Indian empire with special reference to the collection of the Indian Museum. Part 2. (Contd). The family Agrionidae. B. The legions Argia and Agrion. Records of the Indian Museum, 16, 169–195. https://doi.org/10.5962/bhl.part.25920
  33. Laidlaw, F.F. (1920) Contributions to a study of the dragonfly fauna of Borneo. Part IV. A list of species known to occur in the island. Proceedings of the general meetings for scientific business of the Zoological Society of London, 1920 (3), 311–342. https://doi.org/10.1111/j.1469-7998.1920.tb07074.x
  34. Lieftinck, M.A. (1932a) The dragonflies (Odonata) of New Guinea and neighbouring islands. Part 1. Descriptions of new genera and species of the families Lestidae and Agrionidae. Nova Guinea, 15, 485–602.
  35. Lieftinck, M.A. (1932b) An annotated list of the Odonata of Java, with notes on their distribution, habits and life-history. Treubia, 14, 377–462. https://doi.org/10.14203/treubia.v14i4.2454
  36. Lieftinck, M.A. (1934) Descriptions of five new species of Agrionidae from Java (Odonata). Stylops, 3 (1), 6–17. https://doi.org/10.1111/j.1365-3113.1934.tb01521.x
  37. Lieftinck, M.A. (1935) The dragonflies (Odonata) of New Guinea and neighbouring islands. Part 3. Descriptions of new and little known species of the families Megapodagrionidae, Agrionidae and Libellulidae (Genera Podopteryx, Argiolestes, Papuagrion, Teinobasis, Huonia, Synthemis, and Procordulia). Nova Guinea, 17, 203–300.
  38. Lieftinck, M.A. (1936a) On certain types among the Odonata. 1. Pseudagrion australasiae Selys 1876. Konowia, 15, 167–170.
  39. Lieftinck, M.A. (1936b) Die Odonaten der Kleinen Sunda-Indeln. Revue Suisse de Zoologie, 43 (5), 99–160. https://doi.org/10.5962/bhl.part.145462
  40. Lieftinck, M.A. (1937) Descriptions and records of South-east Asiatic Odonata. Treubia, 16 (1), 55–119. https://doi.org/10.14203/treubia.v16i1.2504
  41. Lieftinck, M.A. (1949a) The dragonflies (Odonata) of New Guinea and neighbouring Islands. Part VII. Results of the third Archbold Expedition 1938-1939 and of the Le Roux Expedition 1939 to Netherlands New Guinea (II. Zygoptera). Nova Guinea, New Series, 5, 1–271.
  42. Lieftinck, M.A. (1949b) Synopsis of the Odonate fauna of the Bismarck Archipelago and the Solomon Islands. Treubia, 20 (2), 319–374. https://doi.org/10.14203/treubia.v20i2.2632
  43. Lieftinck, M.A. (1953a) The Odonata of the island Sumba with a survey of the dragonfly fauna of the Lesser Sunda Islands. Verhandlungen der Naturforschenden Gesellschaft in Basel, 64, 118–228.
  44. Lieftinck, M.A. (1953b) Notes on some Odonata from Panaitan Island, Sunda strait (West Java). Idea, 9, 51–56.
  45. Lieftinck, M.A. (1959) Synonymic notes on Brauer1397969521s types of Odonata in the Hamburg Museum. Entomologische Mitteilungen aus dem Zoologischen Staatsinstitut u. Zoologischen Museum Hamburg, 21, 41–48.
  46. Lieftinck, M.A. (1962) Insects of Micronesia. Odonata. Insects of Micronesia, 5 (1), 1–95.
  47. Lieftinck, M.A. (1968) The Odonata of Rennell Island, Solomon Archipelago. The natural history of Rennell Islands, British Solomon Islands, 5, 67–74.
  48. Marinov, M. (2012) Odonata of the Kingdom of Tonga, with a description of Pseudagrion microcephalum stainbergerorum ssp. nov. (Zygoptera: Coenagrionidae). Odonatologica, 41 (3), 225–243.
  49. Marinov, M. (2013) Contribution to the Odonata of the Kingdom of Tonga. Faunistic Studies in South-East Asian and Pacific Island Odonata, 1, 1–18.
  50. Marinov, M., Schmaedick, M., Polhemus, D., Stirnemann, R.L., Enoka, F., Fa1397969521aumu, P.S. & Uili, M. (2015) Faunistic and taxonomic investigations on the Odonata fauna of the Samoan archipelago with particular focus on taxonomic ambiguities in the 1396984945Ischnurine complex1396986481. International Dragonfly Fund-Report, 91, 1–56.
  51. Marinov, M., Bybee, S., Doscher, C. & Kalfatakmolis, D. (2019) Faunistic studies on Odonata of the Republic of Vanuatu (Insecta: Odonata). Faunistic Studies in Southeast Asian and Pacific Island Odonata, 26, 1–46.
  52. Marinov, M., Krieg-Jacquier, R., Duvernay, J.M., Fleck, G., Vincent-Gorlier, J., Mary, N., Doscher, C., Le Bail, F., Jourdan, H. & Theuerkauf, J. (2021) An update of the Odonata fauna of Wallis & Futuna (Insecta: Odonata). Faunistic Studies in Southeast Asian and Pacific Island Odonata, 35, 35–67.
  53. Martin, R. (1901) Les Odonates du continent Australien. Mémoires de la Société Zoologique de France, 14, 220–248.
  54. Martin, R. (1904) Liste des Néuroptères de l1397967985Indo-Chine. In: Pavie, A. (1904) Mission Pavie Indo-Chine 1879-1895. Études diverses III. Recherches sur l1397969521histoire naturelle de l1397969521Inde-Chine orientale. Ernest Leraux, Paris, 204–222.
  55. Michalski, J. (2012) A Manual for the Identification of the Dragonflies & Damselflies of New Guinea, Maluku, & the Solomon Islands. Kanduanum Books, Morristown, 561 pp.
  56. Mitra, T.R. & Babu, R. (2009) Previously unrecorded Odonata from salt Ranges and Sind in Pakistan. Notulae odonatologicae, 7 (4), 37–44.
  57. Morton, K.J. (1907) Odonata collected by Lt.-Colonel Nurse, chiefly in North-Western India. The Transactions of the Entomological Society of London, 1907 (2), 303–308, pl. 24. https://doi.org/10.1111/j.1365-2311.1907.tb01766.x
  58. Needham, J.G. (1930) A Manual of the dragonflies of China. A monographic study of the Chinese Odonata. Zoologia Sinica, Series A, Invertebrates of China, 11 (1), 1–304, pls. 1–20.
  59. Needham, J.G. & Gyger, M.K. (1939) The Odonata of the Philippines, 2. Suborder Zygoptera. The Philippine Journal of Science, 70, 239–314, pls. 11–22.
  60. Orr, A.G. (2001) An annotated checklist of the Odonata of Brunei with ecological notes and descriptions of hitherto unknown males and larvae. International Journal of Odonatology, 4 (2), 167–220. https://doi.org/10.1080/13887890.2001.9748168
  61. Orr, A.G. & van Tol, J. (2001) Pseudagrion lalakense spec. nov. from Borneo with notes on its ecology (Odonata: Coenagrionidae). International Journal of Odonatology, 4 (1), 51–56. https://doi.org/10.1080/13887890.2001.9748158
  62. Paulson, D., Schorr, M., Abbott, J., Bota-Sierra, C., Deliry, C., Dijkstra, K.-D. & Lozano, F. (Coordinators) (2026) World Odonata list. Last revision: 2 March 2026. Available from: https://www.odonatacentral.org/app/#/wol/ (accessed 26 March 2026)
  63. Prasad, M., Ram, R. & Ghosh, S.K. (1987) Notes on the Indian species of the genus Pseudagrion Selys (Coenagrionidae: Odonata). Bulletin of the zoological Survey of India, 8 (1–3), 313–321.
  64. Ram, R., Srivastava, V.D. & Prasad, M. (1982) Odonata (Insecta) fauna of Calcutta and surroundings. Records of the zoological Survey of India, 80, 169–196. https://doi.org/10.26515/rzsi/v80/i1-2/1982/161723
  65. Ram, R., Chandra, K. & Yadav, K. (2000) Studies on the Odonata fauna of Andaman and Nicobar Islands. Records of the zoological Survey of India, 98 (3), 25–60. https://doi.org/10.26515/rzsi/v98/i3/2000/159659
  66. Rambur, P. (1842) Histoire Naturelle des Insectes: Névroptères. Libraire Encyclopédique de Roret, Paris, 534 pp., pls. 1–12.
  67. Ris, F. (1900) Libellen vom Bismarck-Archipel, gesammelt durch Prof. Friedr. Dahl. Archiv für Naturgeschichte, 66, 175–204, pls. 9–10. https://doi.org/10.5962/bhl.part.29035
  68. Ris, F. (1915) Fauna simalurensis. Odonata. Tijdschrift voor Entomologie, 58, 5–21.
  69. Ris, F. (1916) H. Sauter1397967985s Formosa-Ausbeute. Odonata (mit Notizen über andere ostasiatische Odonaten). Supplementa Entomologica, 5, 1–81, pls. 1–3.
  70. Ris, F. (1930) Drei Notizen über ostasiatische Agrioniden (Odonata). Arkiv för Zoologi, 21 (1), 1–32.
  71. Saito, Y. & Ogata, S. (1995) [Records of Hong Kong dragonflies, collected from June 1994 to October 1995. In Japanese]. Bohso no Konchu, 15, 25–47.
  72. Schmidt, E. (1935) Odonata der Deutschen Limnologischen Sunda-Expedition. 1. Imagines. Mit Beschreibungen zweier neuer Ictinus aus Celebes und Neu-Guinea. Archiv für Hydrobiologie Supplement, 13, 316–397, pls. 14–17.
  73. Seehausen, M., Kalkman, V.J. & Bedjanič, M. (2025) Revision of species of the Pseudagrion red-group occurring west of Lydekker1397969521s line, with description of four new species (Odonata: Coenagrionidae). Zootaxa, 5587 (1), 1–66. https://doi.org/10.11646/zootaxa.5587.1.1
  74. Selys Longchamps, E. de (1876) Synopsis des Agrionines (suite de la 5me Légion: Agrion). Bulletins de l1397969521académy royale des sciences, des lettres et des beaux-arts de Belgique, Série 2, 42 (or 45), 247–322 + 496–539.
  75. Selys Longchamps, E. de (1891) Viaggio di Leonardo Fea in Birmania e regioni Vicine XXXII. Odonates. Annali del Museo Civico di Storia Naturale di Genova, Série 2, 10, 433–518.
  76. Theischinger, G. (1997) The Pseudagrion ignifer complex from Australia (Odonata: Zygoptera: Coenagrionidae). Linzer biologische Beiträge, 29 (2), 799–805.
  77. Theischinger, G., Mitchell, A., Polhemus, D. A. & Richards, S. (2026) Systematics of the Pseudagrion red-group from the Bismarck and Solomon archipelagoes (Odonata: Zygoptera: Coenagrionidae). International Dragonfly Fund-Report, 197, 1–38.
  78. Tillyard, R.J. (1906) New Australian species of the family Agrionidae (Neuroptera: Odonata). The Proceedings of the Linnaean Society of New South Wales, 31, 177–194, pl. 17.
  79. Tillyard, R.J. (1913) On some new and rare Australian Agrionidae (Odonata). The Proceedings of the Linnaean Society of New South Wales, 37, 404–479, pls. 44–49.
  80. Tillyard, R.J. (1924) The Dragonflies (Order Odonata) of Fiji, with special reference to a collection made by Mr. H.W. Simmonds, F.E.S., on the Island of Viti Levu. The Transactions of the Entomological Society of London, 1923 (3/4), 305–346.
  81. Tillyard, R.J. (1926) On a collection of Papuan dragonflies (Odonata) made by the late Mr. Allan R. McCulloch in 1922–3, with descriptions of new species. Records of the Australian Museum, 15 (2), 157–166. https://doi.org/10.3853/j.0067-1975.15.1926.806
  82. Tillyard, R.J. & Fraser, F.C. (1938) A reclassification of the order Odonata based on some new interpretations of the venation of the dragonfly wing. With notes, preface and completion thereof. The Australian Zoologist, 9, 125–169.
  83. Trueman, J.W.H. & Rowe, R.J. (2019) The wing venation of Odonata. International Journal of Odonatology, 22 (1), 73–88. https://doi.org/10.1080/13887890.2019.1570876
  84. Vick, G.S. (1985) Odonata collected by the Shiplake College Trekking Society Expedition to Nepal in 1984. Notulae odonatologicae, 2 (5), 80–82.
  85. Villanueva, R.J.T. (2009) Odonata of Dinagat Island, the Philippines: updated species list and notes on conservation of species and habitats. Notulae odonatologicae, 7 (3), 27–35.
  86. Villanueva, R.J.T. (2011) Odonata of Siargao and Bucas Grande islands, The Philippines. International Dragonfly Fund-Report, 34, 1–25.
  87. Villanueva, R.J.T. & Gil, J.R.S. (2011) Odonata Fauna of Catanduanes Island, Philippines. International Dragonfly Fund-Report, 39, 1–38.
  88. Villanueva, R.J.T. & Cahilog, H. (2012) Notes on a small Odonata collection from Tawi-Tawi, Sanga-Sanga and Jolo islands, Philippines. International Dragonfly Fund-Report, 55, 1–32.
  89. Villanueva, R.J.T. & Cahilog, H. (2013) Odonata Fauna of Balabac Island, Philippines with descriptions of two new species. International Dragonfly Fund-Report, 60, 1–34.
  90. Villanueva, R.J.T., Cahilog, H., Jose, E. & van Beijnen, J. (2018) A brief odonatological survey in Palawan and in Cuyo Island, the Philippines. International Dragonfly Fund-Report, 119, 1–12.
  91. Villanueva, R.J.T. (2018) Paracercion luzonicum (amended version of 2009 assessment). The IUCN Red List of Threatened Species, 2018, e.T169275A127510665. https://doi.org/10.2305/IUCN.UK.2009-2.RLTS.T169275A127510665.en
  92. Watson, J.A.L., Theischinger, G. & Abbey, H.M. (1991) The Australian dragonflies. A guide to identification, distributions and habitats of Australian Odonata. CSIRO, Canberra and Melbourne, 278 pp. https://doi.org/10.1071/9780643100701
  93. Weekers, P.H.H. & Dumont, H.J. (2004) A molecular study of the relationship between the coenagrionid genera Erythromma and Cercion, with the creation of Paracercion gen. nov. for the east asiatic 1396984945Cercion1396986481 (Zygoptera: Coenagrionidae). Odonatologica, 33 (2), 181–188.
  94. Wilson, K.D.P. & Reels, G.T. (2001) Odonata of Hainan, China. Odonatologica, 30 (2), 145–208.
  95. Wilson, K.D.P. & Reels, G.T. (2003) Odonata of Guangxi Zhuang Autonomous Region, China, Part 1: Zygoptera. Odonatologica, 32 (3), 237–279.
  96. Yokoi, N. & Souphanthong, V. (2014) A list of Lao dragonflies. Kyoei Printing, Koriyama, 110 pp.
  97. Zhang, H. (2010) The Superfamily Calopterygoidea in South China: taxonomy and distribution. Progress Report for 2009 surveys. International Dragonfly Fund-Report, 26, 1–36.
  98. Zhang, H. (2019) Dragonflies and damselflies of China. 2 Vols. Chongqing University Press, 1460 pp.
  99. Zhang, H., Ning, X., Yu, X. & Bu, W.-J. (2021) Integrative species delimitation based on COI, ITS, and morphological evidence illustrates a unique evolutionary history of the genus Paracercion (Odonata: Coenagrionidae). PeerJ, 9, e11459. https://doi.org/10.7717/peerj.11459
  100. Zhou, X. & Zhou, W.-B. (2007) A new species of Protoneuridae and a new species of Coenagrionidae (Odonata) from China. Entomotaxonomia, 29, 40–44. [in Chinese, English summary]

How to Cite

Seehausen, M. & Marinov, M. (2026) Pseudagrion microcephalum (Rambur, 1842)—sifting through a putative familiar species, with erection of two new species (Odonata: Coenagrionidae). Zootaxa, 5810 (1), 1–48. https://doi.org/10.11646/zootaxa.5810.1.1