Skip to main content Skip to main navigation menu Skip to site footer
Responsive image
Issue: Vol. 5782 No. 2: 24 Mar. 2026
Type: Article
Published: 2026-03-24
DOI: 10.11646/zootaxa.5782.2.1
Page range: 201-248
Abstract views: 71
PDF downloaded: 9

Synopsis of the pinhole borers of West Malaysia (Coleoptera: Curculionidae: Platypodinae)

161/2 Mu 5; Soi Wat Pranon; T. Donkaew; A. Maerim; Chiangmai 50180; Thailand.
Department of Entomology; Michigan State University; 288 Farm Lane; 243 Natural Science Bldg.; East Lansing; MI 48824; USA.; University of Florida School of Forest; Fisheries; & Geomatics Sciences; 136 Newins-Ziegler Hall PO Box 110410 Gainesville; FL 32611; USA
Forest Research Institute Malaysia (FRIM); 52109 Kepong; Selangor; Malaysia.
Department of Entomology; Michigan State University; 288 Farm Lane; 243 Natural Science Bldg.; East Lansing; MI 48824; USA.
Coleoptera ambrosia beetles faunal synopsis new records doubtful records Peninsular Malaysia Malaya

Abstract

A synopsis of the West (Peninsular) Malaysian fauna of platypodine pinhole borer beetles is provided with information on local and world distribution, host trees, biology and taxonomy. One hundred and thirty-eight species are recorded from West Malaysia, eight for the first time. Diapus borneensis Browne comb. res. and Diapus pendleburyi Schedl comb. res. are restored to Diapus Chapuis from Genyocerus Motschulsky. Platypus philippinensis (Schedl) comb. res. is restored to Platypus Herbst from Dendroplatypus Browne. Attention is drawn to the presence of wax glands on the elytra as a possible synapomorphy defining the genus Genyocerus, and to the probable polyphyly of Peroplatypus Wood and Treptoplatypus Schedl. We also list 16 species for which ‘Malaya’ is included in the distribution in various catalogues, but for which we have been unable to find published records or see West Malaysian specimens. These records require confirmation.

 

References

  1. Alonso-Zarazaga, M.A. & Lyal, C.H.C. (2009) A catalogue of family and genus group names in Scolytinae and Platypodinae with nomenclatural remarks (Coleoptera: Curculionidae). Zootaxa, 2258 (1), 1–134. https://doi.org/10.11646/zootaxa.2258.1.1
  2. B.E.S. (1987) Francis George Browne. The Commonwealth Forestry Review, 66 (1), 12–13. [https://www.jstor.org/stable/42608138]
  3. Beaver, R.A. (1977) Bark and ambrosia beetles in tropical forests. BIOTROP Special Publication, 2, 133–147.
  4. Beaver, R.A. (1979) Non-equilibrium ‘island’ communities: a guild of tropical bark beetles. Journal of Animal Ecology, 48, 987–1002. https://doi.org/10.2307/4209
  5. Beaver, R.A. (1989) Insect-fungus relationships in the bark and ambrosia beetles. In: Wilding, N., Collins, N.M., Hammond, P.M. & Webber, J.F. (Eds.), Insect-fungus interactions. Academic Press, London, pp. 121–143. https://doi.org/10.1016/b978-0-12-751800-8.50011-2
  6. Beaver, R.A. (1995) New synonymy and taxonomic changes in Oriental and Australasian Scolytidae and Platypodidae (Insecta: Coleoptera). Annalen des Naturhistorischen Museums in Wien, 97B, 197–204.
  7. Beaver, R.A. (1998a) New synonymy, new combinations and taxonomic notes on Scolytidae and Platypodidae (Insecta, Coleoptera). Annalen des Naturhistorischen Museums in Wien, 100B, 179–192.
  8. Beaver, R.A. (1998b) The genus Platytarsulus Schedl (Coleoptera, Platypodidae). Deutsche Entomologische Zeitschrift, 45, 65–71.
  9. https://doi.org/10.1002/mmnd.4810450108
  10. Beaver, R.A. (1998c) Three new Spathidicerus Chapuis (Col., Platypodidae) with a key to species. Entomologist’s Monthly Magazine, 134, 285–292.
  11. Beaver, R.A. (2000a) Ambrosia beetles (Coleoptera: Platypodidae) of the South Pacific. The Canadian Entomologist, 132, 55–63. https://doi.org/10.4039/Ent132755-6
  12. Beaver, R.A. (2000b) Studies on the genus Diapus Chapuis (Coleoptera: Platypodidae): new species and new synonymy. Serangga, 5, 247–260.
  13. Beaver, R.A. (2002) Studies on the genus Diapus Chapuis (Coleoptera: Platypodidae): keys to males and females, a new species and a new synonym, Serangga, 7, 245–260.
  14. Beaver, R.A. (2007) A new Oriental genus of pinhole borer (Coleoptera: Curculionidae: Platypodinae). Entomologist’s Monthly Magazine, 143, 17–25.
  15. Beaver, R.A. (2013) The invasive neotropical ambrosia beetle Euplatypus parallelus (Fabricius, 1801) in the Oriental region and its pest status (Coleoptera: Curculionidae, Platypodinae). Entomologist’s Monthly Magazine, 149, 143–154.
  16. Beaver, R.A. (2016) The Platypodine ambrosia beetles of Laos (Coleoptera: Curculionidae: Platypodinae). Entomologica Basiliensia et Collectionis Frey, 35, 487–504.
  17. Beaver, R.A. & Browne, F.G. (1979) The Scolytidae and Platypodidae (Coleoptera) of Penang, Malaysia. Oriental Insects, 12 (1978), 575–624. https://doi.org/10.1080/00305316.1978.10432538
  18. Beaver, R.A. & Liu, L-Y. (2007) A review of the genus Genyocerus Motschulsky (Coleoptera: Curculionidae: Platypodinae), with new synonyms and keys to species. Zootaxa, 1576 (1), 25–56.
  19. https://doi.org/10.11646/zootaxa.1576.1.3
  20. Beaver, R.A. & Liu, L-Y. (2013) A synopsis of the pin-hole borers of Thailand (Coleoptera: Curculionidae: Platypodinae). Zootaxa, 3646 (4), 447–486.
  21. https://doi.org/10.11646/zootaxa.3646.4.7
  22. Beaver, R.A. & Liu, L-Y. (2018) A review of the genus Baiocis Browne, 1962 (Coleoptera: Curculionidae: Platypodinae), with new species, new synonymy and a key to males. Zootaxa, 4434 (3), 481–501.
  23. https://doi.org/10.11646/zootaxa.4434.3.5
  24. Beaver, R.A. & Sanguansub, S. (2015) A review of the genus Carchesiopygus Schedl (Coleoptera: Curculionidae: Platypodinae), with keys to species. Zootaxa, 3931 (3), 401–412.
  25. https://doi.org/10.11646/zootaxa.3931.3.4
  26. Beaver, R.A. & Sanguansub, S. (2020) New synonymy and taxonomic changes in Australian and Oriental pin-hole borers (Coleoptera: Curculionidae, Platypodinae). Entomologist’s Monthly Magazine, 156, 79–86. https://doi.org/10.31184/M00138908.1562.4024
  27. Beeson, C.F.C. (1917) The life-history of Diapus furtivus, Sampson. The Indian Forest Records, 6 (1), 1–29, pls. I–II.
  28. Beeson, C.F.C. (1937) New Crossotarsus (Platypodidae, Col.). Indian Forest Records, New Series, Entomology, 3, 49–103.
  29. Beeson, C.F.C. (1961) The ecology and control of the forest insects of India and the neighbouring countries. Government of India, New Delhi, 1007 pp., 36 pls.
  30. Bigger, M. (1988) The insect pests of forest plantation trees in the Solomon Islands. Solomon Islands’ Forest Record, 4, 1–190.
  31. Blandford, W.F.H. (1895) A list of the Scolytidae collected in Ceylon by Mr. George Lewis, with descriptions of new species. Annals and Magazine of Natural History, Series 6, 15 (88), 315–328.
  32. https://doi.org/10.1080/00222939508677888
  33. Blandford, W.F.H. (1896) Descriptions of new Scolytidae from the Indo-Malayan and Austro-Malayan regions. Transactions of the Entomological Society of London, 1896, 191–228.
  34. https://doi.org/10.1111/j.1365-2311.1896.tb00962.x
  35. Browne, F.G. (1935) Biological notes on some Malayan ambrosia beetles. Malayan Forester, 4, 90–95.
  36. Browne, F.G. (1936a) Biological notes on Malayan ambrosia beetles. Malayan Forester, 5, 120–127 + Plates 1–2.
  37. Browne, F.G. (1936b) On the meranti shot-hole borer (Crossotarsus impar Schedl, Coleoptera, Platypodidae). Imperial Forestry Institute Paper, 3, 1–13.
  38. Browne, F.G. (1937) An entomological survey of tapped jelutong trees. Malayan Forester, 6, 240–254.
  39. Browne, F.G. (1938) Biological notes on Malayan Scolytidae. Malayan Forester, 7, 23–30.
  40. Browne, F.G. (1949) Notes on Malayan Scolytoidea (Coleoptera) with descriptions of new species. Annals and Magazine of Natural History, Series 12, 1 (12), 892–912.
  41. https://doi.org/10.1080/00222934808653953
  42. Browne, F.G. (1950) New Scolytidae and Platypodidae (Coleoptera) from Malaya. Annals and Magazine of Natural History, Series 12, 3 (32), 641–650.
  43. https://doi.org/10.1080/00222935008654090
  44. Browne, F.G. (1955) Synonymy and descriptions of some Oriental Scolytidae and Platypodidae (Coleoptera). Sarawak Museum Journal, 6, 343–373.
  45. Browne, F.G. (1961a) The biology of Malayan Scolytidae and Platypodidae. Malayan Forest Records, 22, 1–255.
  46. Browne, F.G. (1961b) Borer beetles from Bako National Park. Sarawak Museum Journal, 10, 300–318.
  47. Browne, F.G. (1962a) Some Scolytidae and Platypodidae (Coleoptera) from the Oriental region. Philippine Journal of Science, 89 (1960), 201–220.
  48. Browne, F.G. (1962b) Taxonomic notes on Platypodidae (Coleoptera). Annals and Magazine of Natural History, Series 13, 4 (47), 641–656.
  49. https://doi.org/10.1080/00222936108651189
  50. Browne, F.G. (1964) On some Platypodidae (Coleoptera) in the collection of the British Museum. Annals and Magazine of Natural History, Series 13, 6 (72), 753–760.
  51. https://doi.org/10.1080/00222936308651426
  52. Browne, F.G. (1968) Pests and Diseases of Forest Plantation Trees. Clarendon Press, Oxford, 1330 pp.
  53. Browne, F.G. (1972) Some Oriental Scolytidae and Platypodidae (Coleoptera). Oriental Insects, 6, 19–31.
  54. https://doi.org/10.1080/00305316.1972.10434049
  55. Browne, F.G. (1977) Beeson’s undescribed Diapodinae and some other species (Col. Platypodidae. Entomologist’s Monthly Magazine, 112, 97–103.
  56. Browne, F.G. (1980a) Bark beetles and ambrosia beetles (Coleoptera, Scolytidae and Platypodidae) intercepted at Japanese ports, with descriptions of new species. I. Kontyû, 48, 370–379.
  57. Browne, F.G. (1980b) Bark beetles and ambrosia beetles (Coleoptera, Scolytidae and Platypodidae) intercepted at Japanese ports, with descriptions of new species. II. Kontyû, 48, 380–389.
  58. Browne, F.G. (1980c) Bark beetles and ambrosia beetles (Coleoptera, Scolytidae and Platypodidae) intercepted at Japanese ports, with descriptions of new species. III. Kontyû, 48, 482–489.
  59. Browne, F.G. (1980d) Bark beetles and ambrosia beetles (Coleoptera, Scolytidae and Platypodidae) intercepted at Japanese ports, with descriptions of new species. IV. Kontyû, 48, 490–500.
  60. Browne, F.G. (1981a) Bark beetles and ambrosia beetles (Coleoptera, Scolytidae and Platypodidae) intercepted at Japanese ports, with descriptions of new species. V. Kontyû, 49, 125–136.
  61. Browne, F.G. (1981b) Bark beetles and ambrosia beetles (Coleoptera, Scolytidae and Platypodidae) intercepted at Japanese ports, with descriptions of new species. VI. Kontyû, 49, 597–606.
  62. Browne, F.G. (1984) Bark beetles and ambrosia beetles (Coleoptera, Scolytidae and Platypodidae) intercepted at Japanese ports, with descriptions of new species. X. Kontyû, 52, 448–457.
  63. Browne, F.G. (1986a) Bark beetles and ambrosia beetles (Coleoptera, Scolytidae and Platypodidae) intercepted at Japanese ports, with descriptions of new species. XIV. Kontyû, 54, 333–343.
  64. Browne, F.G. (1986b) Bark beetles and ambrosia beetles (Coleoptera, Scolytidae and Platypodidae) intercepted at Japanese ports, with descriptions of new species. XV. Kontyû, 54, 661–671.
  65. Bumrungsri, S., Beaver, R., Phongpaichit, S. & Sittichaya, W. (2008) The infestation by an exotic ambrosia beetle, Euplatypus parallelus (F.) (Coleoptera: Curculionidae: Platypodinae) of Angsana trees (Pterocarpus indicus Willd.) in southern Thailand. Songklanakarin Journal of Science and Technology, 30, 579–582.
  66. Chapuis, F. (1865) Monographie des Platypides. H. Dessain, Liège, 344 pp. https://doi.org/10.5962/bhl.title.9204
  67. Chey, V-K. (2002) Major insect pests and their management in forest plantations in Sabah, Malaysia. In: Baig, S.A.S., Varley, J.G. & Chin, J.S. (Eds.), Forest Health in Central and South and Southeast Asia. FORSPA Publication. No. 30. Food and Agriculture Organization of the United Nations (FAO) / Regional Office for Asia and the Pacific, Bangkok, pp. 19–23.
  68. Fabricius, J.C. (1801) Systema eleutheratorum secundum ordines, genera, species: adiectis synonymis, locis, observationibus, descriptionibus. Vol. 2. Bibliopoli Academici Novi, Kiliae, 687 pp.
  69. Fairmaire, L. (1850) Essai sur les Coléoptères de la Polynésie. Revue et Magasin de Zoologie Pure et Appliquée, Série 2, 2 (1850), 50–64 + 115–122 + 181–185.
  70. Gray, B & Wylie, F.R. (1974) Forest tree and timber pests in Papua New Guinea. Pacific Insects, 16, 67–115.
  71. Ho, Y.F. (1993) Platypus parallelus, a common ambrosia beetle of timbers. FRIM Technical Information, 43, 1–4.
  72. Hulcr, J. & Stelinski. L.L. (2017) The ambrosia symbiosis: from evolutionary ecology to practical management. Annual Review of Entomology, 62, 285–303.
  73. https://doi.org/10.1146/annurev-ento-031616-035105
  74. Hulcr, J., Dole, S.A., Beaver, R.A. & Cognato, A.I. (2007) Cladistic review of generic taxonomic characters in Xyleborini (Coleoptera: Curculionidae: Scolytinae). Systematic Entomology, 32, 568–584. https://doi.org/10.1111/j.1365-3113.2007.00386.x
  75. Hulcr, J., Atkinson, T.H., Cognato, A.I., Jordal, B.H. & McKenna, D.D. (2015) Morphology, taxonomy, and phylogenetics of bark beetles. In: Vega, F.E. & Hofstetter, R.W. (Eds.), Bark Beetles. Biology and Ecology of Native and Invasive Species. Academic Press, London, pp. 41–84. https://doi.org/10.1016/B978-0-12-417156-5.00002-2
  76. Jayanthi, P.D.K., Suma, A., Sane, A., Sreedevi, K. & Veena, N.V. (2023) First report of Euplatypus parallelus (Fabricius) (Coleoptera: Curculionidae: Platypodinae) on pomegranate (Punica granatum L.), symptoms of infestation, its attraction to ethanol-baited traps, and niche partitioning with scolytines. The Coleopterists Bulletin, 76, 70–72.
  77. https://doi.org/10.1649/0010-065X-76.1.70
  78. Jordal, B.H. (2015) Molecular phylogeny and biogeography of the weevil subfamily Platypodinae reveals evolutionarily conserved range patterns. Molecular Phylogenetics and Evolution, 93, 294–307.
  79. https://doi.org/10.1016/j.ympev.2015.05.028
  80. Kalshoven, L.G.E. (1960) Studies on the biology of Indonesian Scolytoidea. 7. Data on the habits of Platypodidae. Tijdschrift voor Entomologie, 103, 31–50.
  81. Kangkamanee, T., Sittichaya, W., Ngampongsai, A., Permkam, S. & Beaver, R.A. (2011) Wood-boring beetles (Coleoptera: Bostrichidae, Curculionidae: Platypodinae and Scolytinae) infesting rubberwood sawn timber in Southern Thailand. Journal of Forest Research, 16, 302–308.
  82. https://doi.org/10.1007/s10310-010-0224-7
  83. Kent, D.S. & Simpson, J.A. (1992) Eusociality in the beetle Austroplatypus incompertus (Coleoptera: Curculionidae). Naturwissenschaften, 79 (2), 86–87. https://doi.org/10.1007/bf01131810
  84. Kirkendall, L.R., Biedermann, P.H.W. & Jordal, B.H. (2015) Evolution and diversity of bark and ambrosia beetles. In: Vega, F.E. & Hoffstetter, R.W. (Eds.), Bark Beetles: Biology and Ecology of Native and Invasive Species, Academic Press, New York, New York, pp. 85–156. https://doi.org/10.1016/B978-0-12-417156-5.00003-4
  85. Kirkendall, L.R. & Atkinson, T.H. (2024) What we do and don’t know about New World pinhole borers (Coleoptera, Curculionidae, Platypodinae). Norwegian Journal of Entomology Supplement, 4, 25–92.
  86. Maetô, K. & Fukuyama, K. (2003) Vertical stratification of ambrosia beetle assemblage in a lowland rain forest at Pasoh, Peninsular Malaysia. In: Okuda, T., Manokaran, N., Matsumoto, Y., Niiyama, K., Thomas, S.C. & Ashton, P.S. (Eds.), Pasoh. Springer, Tokyo, pp. 325–336. https://doi.org/10.1007/978-4-431-67008-7_24
  87. Manohara, T.M., Balakrishna, S.M. & Shetteppanavar, V.S. (2023) Infestation of pinhole borer (Euplatypus parallelus (F.)) on Dalbergia latifolia Roxb. Plant Science Today, 10, 75–78.
  88. https://doi.org/10.14719/pst.2074
  89. Mayers, C.G., Harrington, T.C. & Biedermann, P.H.W. (2022) Mycangia define the diverse ambrosia beetle-fungus symbioses. In: Schultz, T.R., Gawne, R. & Peregrine, P.N. (Eds.), The Convergent Evolution of Agriculture in Humans and Insects. MIT Press, Cambridge, Massachusetts, pp. 105–142.
  90. https://doi.org/10.7551/mitpress/13600.003.0013
  91. Motschulsky, V. de (1858) Insectes des Indes orientales. Études Entomologiques, 7, 20–122
  92. Motschulsky, V. de (1863) Essai d’un catalogue des insectes de l’Île Ceylan. Bulletin de la Société Impériale des Naturalistes de Moscou, 36, 421–532.
  93. Murphy, D.H. & Meepol, W. (1990) Timber beetles of the Ranong mangrove forests. Mangrove Ecosystems Occasional Paper, 7, 5–8.
  94. Nikulina, T., Mandelshtam, M., Petrov, A., Nazarenko, V. & Yunakov, N. (2015) A survey of the weevils of Ukraine. Bark and ambrosia beetles (Coleoptera: Curculionidae: Platypodinae and Scolytinae). Zootaxa, 3912 (1), 1–61. https://doi.org/10.11646/zootaxa.3912.1.1
  95. Nobuchi, A. (1973) The Platypodidae of Japan. Bulletin of the Government Forest Experiment Station, Tokyo, 256, 1–22.
  96. Nunberg, M. (1939) Neue Arten der Gattung Platypus Hrbst. aus Peru (Coleoptera). Annales Musei Zoologici Polonici, 13, 221–244, tabs. XVI–XXII.
  97. Ohno, S. (1990) The Scolytidae and Platypodidae (Coleoptera) from Borneo found in logs at Nagoya port. 2. Research Bulletin of the Plant Protection Service of Japan, 26, 95–103.
  98. Ohno, S., Yoneyama, K. & Nakazawa, H. (1987a) The Scolytidae and Platypodidae (Coleoptera) from Philippines, found in logs at Nagoya port. Research Bulletin of the Plant Protection Service of Japan, 23, 87–91.
  99. Ohno, S., Yoneyama, K. & Nakazawa, H. (1987b) The Scolytidae and Platypodidae (Coleoptera) from Moluccas Islands, found in logs at Nagoya port. Research Bulletin of the Plant Protection Service of Japan, 23, 93–97.
  100. Ohno, S., Yoshioka, K., Uchida, N., Yoneyama, K. & Tsukamoto, K. (1989) The Scolytidae and Platypodidae (Coleoptera) from Bismarck Archipelago found in logs at Nagoya port. Research Bulletin of the Plant Protection Service of Japan, 25, 59–69.
  101. Parsons, A.D. (1963) Ambrosia beetles and their importance in the Sarawak peat swamp forests. Commonwealth Forestry Review, 42, 347–354.
  102. Philip, E. (1999) Wilt disease of angsana (Pterocarpus indicus) in Peninsular Malaysia and its possible control. Journal of Tropical Forest Science, 11, 519–527.
  103. Prathapan, K.D. (1991) Platypus andrewesi Strohmeyer (Coleoptera: Platypodidae) as a pest of coconut palm in Kerala. Proceedings of the Kerala Science Congress, February–March 1991, 69.
  104. Roberts, H. (1969) Forest insects of Nigeria with notes on their biology and distribution. Commonwealth Forestry Institute Papers, 44, 1–206.
  105. Roberts, H. (1989) New Crossotarsus from Papua New Guinea (Platypodidae, Coleoptera). Koleopterologische Rundschau, 59, 87–93.
  106. Roberts, H. (1993) The tribe Diapodini (Coleoptera: Platypodidae) of New Guinea. Bishop Museum Occasional Papers, 35, 1–39.
  107. Sampson, F.W. (1912) Some new species of Ipidae and Platypodidae in the British Museum. Annals and Magazine of Natural History, Series 8, 10 (56), 245–250.
  108. https://doi.org/10.1080/00222931208693228
  109. Sampson, F.W. (1913) Some hitherto undescribed Ipidae and Platypodidae from India and Burma. Annals and Magazine of Natural History, Series 8, 12 (71), 443–452.
  110. https://doi.org/10.1080/00222931308693422
  111. Sampson, F.W. (1922) Previously undescribed Scolytidae and Platypodidae from the Indian area. Annals and Magazine of Natural History, Series 9, 10 (55), 145–152.
  112. https://doi.org/10.1080/00222932208632753
  113. Sampson, F.W. (1923) Some new Burmese Platypodidae. Annali del Museo Civico di Storia Naturale di Genova, 51, 71–75.
  114. Sampson, F.W. (1928) Platypodidae from Borneo. Sarawak Museum Journal, 3, 389–394.
  115. Sanderson, F.R., Fong, Y.K., Yik, C.P., Ong, K.H. & Anuar, S. (1997) A Fusarium wilt (Fusarium oxysporum) of angsana (Pterocarpus indicus) in Singapore. I. Epidemiology and identification of the causal organism. Arboricultral Journal, 21, 187–204. https://doi.org/10.1080/03071375.1997.9747165
  116. Schedl, K.E. (1935a) Scolytidae and Platypodidae: new species from the Philippines and Formosa. Philippine Journal of Science, 57, 479–489.
  117. Schedl, K.E. (1935b) Some new Platypodidae from Borneo and Malaya. Journal of the Federated Malay States Museums, 17, 632–642.
  118. Schedl, K.E. (1935c) Fauna Philippinensis (Platypodidae et Scolytidae). III. Philippine Journal of Science, 56, 395–403.
  119. Schedl, K.E. (1935d) Scolytidae und Platypodidae. 30. Beitrag. Miscellanea Zoologica Sumatrana, 94, 1–4.
  120. Schedl, K.E. (1936a) Notes on Malaysian Scolytidae and Platypodidae and descriptions of some new species. Journal of the Federated Malay States Museums, 18, 1–18.
  121. Schedl, K.E. (1936b) Some new Scolytidae and Platypodidae from the Malay peninsula. Journal of the Federated Malay States Museums, 18, 19–35.
  122. Schedl, K.E. (1936c) Platypodidae des Museo Civico di Storia Naturale di Genova. Scolytidae und Platypodidae. 27 Beitrag. Annali del Museo Civico di Storia Naturale di Genova, 59, 43–62.
  123. Schedl, K.E. (1936d) Scolytidae und Platypodidae. Fauna Philippinensis, IV. Philippine Journal of Science, 60, 59–69.
  124. Schedl, K.E. (1937) Platypodiden des Berliner Zoologisches Museums. Entomologische Blätter, 33, 33–44.
  125. Schedl, K.E. (1938) Scolytidae und Platypodidae. Fauna Philippinensis V. Philippine Journal of Science, 67, 421–429.
  126. Schedl, K.E. (1939a) Malaysian Scolytidae and Platypodidae (IV). Journal of the Federated Malay States Museums, 18, 327–364.
  127. Schedl, K.E. (1939b) Die Einteilung und geographische Verbreitung der Platypodidae. Verhandlungen, VII Internationale Kongresse fur Entomologie, Berlin, 1, 377–410. [1938]
  128. Schedl, K.E. (1941a) Die Variations-Breite in den Platypi cupulati Chap. Archiven für Naturgeschichte, Neue Folge, 10, 416–426.
  129. Schedl, K.E. (1941b) Javanische Platypodiden. 74 Beitrag. Entomologische Berichten, 10, 354–366.
  130. Schedl, K.E. (1942a) Zur Scolytoiden-Fauna der Malayischen Halbinsel. V. Kolonialforstliche Mitteilungen, 5, 169–218.
  131. Schedl, K.E. (1950) Fauna Indo-Malayensis. II. 104 Contribution. Annals and Magazine of Natural History, Series 12, 3 (34), 892–900.
  132. https://doi.org/10.1080/00222935008654721
  133. Schedl, K.E. (1951) Fauna Indo-malayensis I. 91 Beitrag. Tijdschrift voor Entomologie, 93 (1950), 41–98.
  134. Schedl, K.E. (1951/52) Zur Synonymie der Borkenkäfer I. 121 Beitrag. Entomologische Blätter, 47–48, 158–164.
  135. Schedl, K.E. (1952) Scolytoidea nouveaux du Congo Belge. 134e Contribution. Annales du Musée Royal du Congo Belge, Tervuren, Série 8vo, Sciences Zoologiques, 13, 1–62.
  136. Schedl, K.E. (1954) Fauna indomalayensis IV. 141 Beitrag. Philippine Journal of Science, 83, 137–159.
  137. Schedl, K.E. (1958a) Bark and timber beetles from Malaya. Malayan Forester, 21, 99–105.
  138. Schedl, K.E. (1958b) Scolytidea [sic] from Borneo – II. Sarawak Museum Journal, 8, 498–499.
  139. Schedl, K.E. (1960) Zur Synonymie der Borkenkäfer, V. 181 Beitrag. Entomologische Blätter, 56, 103–112.
  140. Schedl, K.E. (1962a) Zur Synonymie der Borkenkäfer, VI. 203 Beitrag. Entomologische Blätter, 58, 201–211.
  141. Schedl, K.E. (1962b) Indian bark and timber beetles. IV. 201 Contribution. Indian Forest Records, New Series, Entomology, 10 (8), 163–169. [1960]
  142. Schedl, K.E. (1964a) Neue und interessante Scolytoidea von den Sunda-Inseln, Neu Guinea und Australien. 220 Beitrag. Tijdschrift voor Entomologie, 107, 297–306.
  143. Schedl, K.E. (1964b) Scolytoidea from Borneo III. 185 Contribution. Reichenbachia, 4, 241–254.
  144. Schedl, K.E. (1964c) Three new species of Scolytidae from Australia, and some introduced Coleoptera. Proceedings of the Linnaean Society of New South Wales, 89, 246–249.
  145. Schedl, K. E. (1965) Scolytidae und Platypodidae Afrikas. Band 3. Platypodidae. Revista de Entomologia de Moçambique, 5, 595–1352.
  146. Schedl, K.E. (1966) Checklist of the Scolytidae and Platypodidae from the Philippine Islands. 196 Contribution. Entomologische Abhandlungen aus dem Staatliches Museum für Tierkunde, Dresden, 35, 1–122.
  147. Schedl, K.E. (1967) Zur Synonymie der Borkenkäfer XVI. Entomologisk Tidskrift, 88, 147–163.
  148. Schedl, K.E. (1969a) Bark beetles and pin-hole borers (Scolytidae and Platypodidae) intercepted from imported logs in Japanese ports III. Kontyû, 37, 202–219.
  149. Schedl, K.E. (1969b) Indian bark and timber beetles. V. Oriental Insects, 3, 47–70. https://doi.org/10.1080/00305316.1969.10433895
  150. Schedl, K.E. (1970) Bark beetles and pinhole borers (Scolytidae and Platypodidae) intercepted from imported logs in Japanese ports IV. 274 Contribution. Kontyû, 38, 353–370.
  151. Schedl, K.E. (1971) Indomalayan bark and timber beetles. 273 Contribution. Oriental Insects, 5, 361–399.
  152. https://doi.org/10.1080/00305316.1971.10434023
  153. Schedl, K.E. (1972a) Monographie der Familie Platypodidae Coleoptera. W. Junk, Den Haag, 322 pp.
  154. Schedl, K.E. (1972b) Zur Synonymie der Borkenkäfer. XXI. 281 Beitrag. Entomologische Arbeiten aus dem Museum G. Frey, 23, 150–161.
  155. Schedl, K.E. (1975a) New Scolytidae and Platypodidae from Papua and New Guinea. 317 Contribution. Annalen des Naturhistorischen Museums in Wien, 79, 337–399.
  156. Schedl, K.E. (1975b) Bark and timber beetles of the Oriental region. Oriental Insects, 9, 451–460. https://doi.org/10.1080/00305316.1975.10434514
  157. Schedl, K.E. (1977) Die Scolytidae und Platypodidae Madagaskars und einiger naheliegender Inselgruppen. Mitteilungen der Forstlichen Bundes-Versuchsanstalt, Wien, 119, 1–326.
  158. Schedl, K.E. (1978) Die Typen der Sammlung Schedl. Familie Platypodidae (Coleoptera). 324 Beitrag. Kataloge der wissenschaftlichen Sammlungen des Naturhistorischen Museums in Wien, Entomologie, 1, 1–82.
  159. Schedl, K.E. (1979) Zur Synonymie der Borkenkäfer XXIX. 345 Beitrag. Entomologische Arbeiten aus dem Museum G. Frey, 28, 119–131.
  160. Shuckard, W.E. (1840) The British Coleoptera delineated, consisting of figures of all the genera of British Beetles, drawn in outline by W. Spry, W. Crofts, London, VIII + 84 pp., 94 pls.
  161. https://doi.org/10.5962/bhl.title.59532
  162. Smith, J.H. (1935) The pinhole borer of north Queensland cabinet woods. Queensland Division of Entomology and Plant Pathology Bulletin, New Series, 12, 1–38.
  163. Smith, S.M., Beaver, R.A., Ong, S.P. & Cognato, A.I. (2025) A checklist of West Malaysian bark and ambrosia beetles (Coleoptera: Curculionidae: Scolytinae). Zootaxa, 5720 (1), 1–114. https://doi.org/10.11646/zootaxa.5720.1.1
  164. Stebbing, E.P. (1914) Indian Forest Insects of Economic Importance. Eyre & Spottiswoode, London, 648 pp. https://doi.org/10.5962/bhl.title.9203
  165. Strohmeyer, H. (1910) Neue Borkenkäfer aus Abessynien, Madagaskar, Indien und Tasmania. Entomologische Blätter, 6, 126–132.
  166. Strohmeyer, H. (1912) ‘H. Sauter’s Formosa-ausbeute’, Ipidae und Platypodidae (Col.). Entomologische Mitteilungen, 1, 38–42.
  167. https://doi.org/10.5962/bhl.part.25902
  168. Strohmeyer, H. (1913) Neue Platypodiden. Entomologische Blätter, 9, 161–165.
  169. Strohmeyer, H. (1914a) Neue Platypodiden aus der Stettiner Museum. Stettiner Entomologische Zeitung, 75, 3–8.
  170. Strohmeyer, H. (1914b) Coleoptera. Fam. Platypodidae. In: Wytsman, P.A.G. (Ed.), Genera Insectorum, 163, 1–55 + pl. 1–12.
  171. Sukartana, P. (1987) Tunneling and breeding habits of ambrosia beetle, Platypus trepanatus in ramin log. Jurnal Penelitian Hasil Hutan, 4 (2), 30–35.
  172. Thomson, J. (1857) Description de trois espèces de Coléoptères. Archives Entomologiques, 1, 341–344.
  173. Thube, S.H., Mohan, C., Pandian, R.T.P., Saneera, E.K., Sannagoudra, H.M., Hegde, V. & Chowdappa, P. (2018) First record of the invasive neotropical ambrosia beetle Euplatypus parallelus (Fabricius, 1801) (Coleoptera: Curculionidae: Platypodinae) infesting arecanut in Karnataka, India. The Coleopterists Bulletin, 72, 713–716.
  174. https://doi.org/10.1649/0010-065X-72.4.713
  175. Whitmore, T.C. (1984) Tropical Rain Forests of the Far East. Clarendon Press, Oxford, 352 pp.
  176. Wood, S.L. (1960) Insects of Micronesia, Coleoptera: Platypodidae and Scolytidae. Insects of Micronesia, 18 (1), 1–73.
  177. Wood, S.L. (1969) New synonymy and records of Platypodidae and Scolytidae (Coleoptera). Great Basin Naturalist, 29, 113–128.
  178. https://doi.org/10.5962/bhl.part.17054
  179. Wood, S.L. (1986) A reclassification of the genera of Scolytidae (Coleoptera). Great Basin Naturalist Memoirs, 10, 1–126.
  180. Wood, S.L. (1993) Revision of the genera of Platypodidae (Coleoptera). Great Basin Naturalist, 53, 259–281.
  181. https://doi.org/10.5962/bhl.part.16605
  182. Wood, S.L. & Bright, D.E. (1992) A catalog of Scolytidae and Platypodidae (Coleoptera), Part 2: Taxonomic index. Great Basin Naturalist Memoirs, 13, 1–1553.

How to Cite

Beaver, R.A., Smith, S.M., Ong, S.P. & Cognato, A.I. (2026) Synopsis of the pinhole borers of West Malaysia (Coleoptera: Curculionidae: Platypodinae). Zootaxa, 5782 (2), 201–248. https://doi.org/10.11646/zootaxa.5782.2.1