Abstract
The malthinine fauna of Asia is highly diverse, but comprehensive studies are lacking, and many species have not been re-examined since their original descriptions. This study mainly focuses on the genus Malthinellus Kiesenwetter, 1874, which comprises ten Asian species. A new species, Malthinellus mushanus sp. nov. is described from Taiwan (Nantou County) and compared with two known Taiwanese species from the same genus, M. crenulatus (Wittmer, 1979) and M. crenulatomimus (Wittmer, 1984). Aedeagus of Malthinellus chinensis Wittmer, 1956 (China) is illustrated and described for the first time. Malthinellus hoabinensis (Wittmer, 1995) comb. nov. (Vietnam) and Malthinellus multispinosus (Wittmer, 1995) comb. nov. (China) are transferred from the genus Malthinus. Malthinellus malickyi Wittmer, 1997 (Thailand) is transferred to a new genus Phaneromalthinus gen. nov. (type species: P. malickyi (Wittmer, 1997) comb. nov.). Additionally, illustrations and notes are provided for several species of the genera Malthinus and Mimomalthinus that resemble Malthinellus. The lectotypes of Malthinus tonkineus Pic, 1923 and Mimomalthinus niger Pic, 1931 are designated and illustrated. The generic definition of Malthinellus is revised, and the differences from Malthinus and Mimomalthinus are discussed.
References
- Biffi, G. & Geiser, M. (2020) A revision of Peltariosilis Wittmer (Coleoptera: Cantharidae), a surprisingly diverse Amazonian radiation. Papéis Avulsos De Zoologia, 60 (special), e202060(s.i.).16. https://doi.org/10.11606/1807-0205/2020.60.special-issue.16
- Brancucci, M. (1980) Morphologie comparée, évolution et systématique des Cantharidae (Insecta: Coleoptera). Entomologica Basiliensia, 5, 215–388.
- Brancucci, M. & Wittmer, W. (1984) Zur Kenntnis der indo-malaiischen Malthininae (Coleoptera, Cantharidae). Entomologica Basiliensia, 9, 183–192.
- Fanti, F. (2022) A new fossil of Mantimalthinus (Coleoptera: Cantharidae) sheds light on its tribal attribution. Studies and Reports Taxonomical Series, 18 (2), 281–286.
- Fanti, F. & Castiglione, E. (2017) Description of a new genus and species of Cantharidae from Eocene Baltic amber (Insecta, Coleoptera). Palaeodiversity, 10, 123–127. https://doi.org/10.18476/pale.v10.a8
- Fanti, F. & Damgaard, A.L. (2019) New soldier beetles (Cantharidae) from Baltic, Burmese and Dominican ambers of the Anders Damgaard amber collection. Baltic Journal of Coleopterology, 19 (2), 101–125.
- Kazantsev, S.V. (2006) Discovery of the monotypic oriental genus Mimomalthinus Pic, 1931 (Coleoptera: Cantharidae) in the Russian Far East. Russian Entomological Journal, 16, 207–210.
- Kazantsev, S.V. & Brancucci, M. (2007) Cantharidae. In: Löbl, I. & Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera, 4. Elateroidea—Derodontoidea—Bostrichoidea—Lymexyloidea—Cleroidea—Cucujoidea. Apollo Books, Stenstrup, pp. 234–298.
- von Kiesenwetter, H. (1874) Die Malacodermen Japans nach dem Ergebnisse der Sammlungen des Herrn G. Lewis während der Jahre 1869–1871. Berliner Entomologische Zeitschrift, 18, 241–288. https://doi.org/10.1002/mmnd.18740180302
- Kim, J.-I. & Kang, T.-H. (2000) Taxonomic study of Korean Cantharidae (Coleoptera) I. Silinae, Malthininae, and Chauliognathinae. Insecta Koreana, 17, 111–120.
- Kiriyama, I. (2003) Collected records of cantharid beetles in Gifu Prefecture, Honshu, Japan (1). Keichitsu, 13 (35), 1–10. [in Japanese, with English title]
- Latreille, P.A. (1806) Genera Crustaceorum et Insectorum secundum ordinem naturalem in familias disposita, iconibus exemplisque plurimus explicata. Vol. 1. Amand Koenig, Parisiis, XVIII + 302 pp., 16 pls. https://doi.org/10.5962/bhl.title.65741
- Motyka, M., Kusy, D., Biffi, G., Geiser, M., Kazantsev, S.V., Bilkova, R., Jahodarova, E., Vogler, A.P. & Bocak, L. (2023) Untangling the evolution of soldier beetles (Coleoptera: Cantharidae) and the evaluation of the morphological phylogenetic signal in a soft-bodied elateroid lineage. Cladistics, 39, 548–570. https://doi.org/10.1111/cla.12555
- Nakamura, R. (2023) New record of Malthinellus exiguus N. Takahashi from Zamami-jima Island, the Ryukyus, Japan. Sayabane, New Series, 51, 32. [in Japanese]
- Nakamura, R. (2025) New records of four cantharid species from Yoro-shima Island, the Amami Islands, the Ryukyus, Japan. Sayabane, New Series, 58, 71. [in Japanese]
- Nakamura, R., Hayashi, M. & Okushima, Y. (2025) Fauna of Cantharidae (Coleoptera) of the Oki Islands, Shimane Prefecture, Japan. Bulletin of the Hoshizaki Green Foundation, 28, 219–243. [in Japanese, with English title and abstract.]
- Pic, M. (1931) Malacodermes exotiques. L'Echange, Revue Linneenne, Tors-Texte, 47 (443–445), 93–96 + 97–100 + 101–104.
- Satô, M. (1986) Taxonomic notes on Cantharoidea of Japan, I. Coleopterists' News, Tokyo, 72, 1–3. [in Japanese, with English title]
- Satô, M., Okushima, Y., Takahashi, N., Li, C.-L., Yang, X.-Y. & Hsiao, Y. (2014) Checklist of the Cantharidae of Taiwan (Coleoptera: Elateroidea). Collection and Research, 27, 43–69.
- Takahashi, K. (1992) The fauna of family Cantharidae (Coleoptera) in Kanagawa Prefecture, Japan. Kanagawa-Chûhô, 100, 71–124. [in Japanese, with English title]
- Takahashi, K. (2016) A new species of the genus Malthinellus (Coleoptera, Cantharidae) from Iriomote Island of the Ryukyu Islands, Southwest Japan. Elytra, Tokyo, New Series, 6, 295–298.
- Takahashi, K. (2017) Notes on the group of Malthinus mucoreus (Coleoptera, Cantharidae, Malthininae) from the Ryukyu Islands, southwestern Japan. Elytra, Tokyo, New Series, 7, 287–295.
- Takahashi, K., Nakamura, R. & Kubota, K. (2022) The genus Malthinellus Kiesenwetter, 1874 (Coleoptera, Cantharidae, Malthininae) in Japan. Japanese Journal of Systematic Entomology, 28, 305–310.
- Takahashi, K. & Takahashi, N. (2007) A new species of the genus Malthinellus (Coleoptera, Cantharidae) from Okinawa Island, Southwest Japan, with notes on the genus Malthinellus. Elytra, Tokyo, 35, 142–148.
- Takahashi, N. (2018) A new soldier beetle Malthinellus exiguus sp. nov. (Coleoptera: Cantharidae) from the Ryukyus in southwestern Japan, with a key to Japanese species. Zootaxa, 4403 (2), 395–400. https://doi.org/10.11646/zootaxa.4403.2.11
- Wittmer, W. (1956) Neue Malacodermata aus der Sammlung der California Academy of Sciences (16. Beitrag zur Kenntnis der palaearktischen Malacodermata, Col.). Mitteilungen der Schweizerischen Entomologischen Gesellschaft, 29, 303–313.
- Wittmer, W. (1974) Zur Kenntnis der Gattung Malthinus Latr. II (Col Cantharidae) (60. Beitrag zur Kenntnis der paläarktischen Cantharidae). Entomologische Arbeiten aus dem Museum G. Frey Tutzing bei München, 25, 358–427.
- Wittmer, W. (1979) 64. Beitrag zur Kenntnis der palaearktischen Cantharidae, Phengodidae und Malachiidae (Col.). Entomologica Basiliensia, 4, 327–346.
- Wittmer, W. (1984) Die Familie Cantharidae (Col.) auf Taiwan (3. Teil). Entomological Review of Japan, 39, 141–166, pls. 4–9.
- Wittmer, W. (1988) Zur Kenntnis der Cantharidae (Coleoptera) Chinas und der angrenzenden Länder. Entomologica Basiliensia, 12, 343–372.
- Wittmer, W. (1993) 79. Beitrag zur Kenntnis der palaearktischen Fauna (Coleoptera-Cantharidae). Entomologica Basiliensia, 16, 279–305.
- Wittmer, W. (1995) Neue Cantharidae (Col.) aus dem indo-malaiischen und palaearktischen Faunengebiet mit Mutationen. Entomologica Basiliensia, 18, 109–169.
- Wittmer, W. (1997) Neue Cantharidae (Col.) aus dem indo-malaiischen und palaearktischen Faunengebiet mit Mutationen. 2. Beitrag. Entomologica Basiliensia, 20, 223–365.
