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Issue: Vol. 5756 No. 1: 2 Feb. 2026
Type: Monograph
Published: 2026-02-02
DOI: 10.11646/zootaxa.5756.1.1
Page range: 1-77
Abstract views: 53
PDF downloaded: 2

A taxonomic revision of the genus Dichoxenus Horn, 1876 (Coleoptera: Curculionidae: Entiminae: Byrsopagini)

Beaty Centre for Species Discovery; Canadian Museum of Nature; 1740 Chemin Pink Gatineau; PQ; J9J 3N7; Canada
Coleoptera species discovery biodiversity endemism sky islands Mexico

Abstract

The genus Dichoxenus Horn, 1876 is revised with 25 species, including 17 new species as follows: Dichoxenus charlesi Anderson, new species (Mexico, Nuevo León), Dichoxenus chiapaneca Anderson, new species (Mexico, Chiapas), Dichoxenus conejo Anderson, new species (Mexico, Veracruz), Dichoxenus cupreus Anderson, new species (U.S.A., Texas), Dichoxenus howdenorum Anderson, new species (Mexico, Nuevo León), Dichoxenus jonesi Anderson, new species (Mexico, Querétaro), Dichoxenus loisae Anderson, new species (Mexico, Nuevo León), Dichoxenus manzanae Anderson, new species (Mexico, Coahuila), Dichoxenus mcclevei Anderson, new species (U.S.A., Arizona), Dichoxenus reddelli Anderson, new species (U.S.A., Texas), Dichoxenus sandersi Anderson, new species (U.S.A., Texas), Dichoxenus schwartzi Anderson, new species (U.S.A., Texas), Dichoxenus sculpturatus Anderson, new species (Mexico, Nuevo León), Dichoxenus spatulirostris Anderson, new species (U.S.A., Alabama), Dichoxenus sulcirostris Anderson, new species (U.S.A., Texas), Dichoxenus sulcithorax Anderson, new species (U.S.A., Arizona) and Dichoxenus tamaulipensis Anderson, new species (Mexico, Tamaulipas). Anametis Horn, 1876 is here considered a new junior subjective synonym of Dichoxenus and the following are proposed as new combinations: Dichoxenus granulatus (Say, 1832), Dichoxenus griseus (Horn, 1876), and Dichoxenus subfuscus (Fall, 1907). Amphidees longulus Sharp, 1891 is transferred to Dichoxenus as Dichoxenus longulus (Sharp, 1891), new combination. Anametis setosus Blatchley, 1916 and Dichoxenus setiger arkansasensis Sleeper, 1956 are new junior subjective synonyms of Dichoxenus granulatus (Say, 1832). Lectotypes are designated for Amphidees longulus Sharp, 1891, Barynotus granulatus Say, 1832 and Anametis grisea Horn, 1876. Most species of Dichoxenus are collected on vegetation, especially at night, but some have been collected in leaf litter and three species are suspected or known only from caves. One of these species, D. sandersi, shows more extreme character states typical of cave life, such as reduced pigmentation, long slender appendages and reduced eye size. A key to the 25 species of Dichoxenus as well as habitus images and images of male genitalia are presented to facilitate identification. All distributions are mapped. A key is also given to the genera of Byrsopagini of North America lacking a metanepisternal suture.

 

References

  1. Alonso-Zarazaga, M.A. & Lyal, C.H.C. (1999) A World Catalogue of Families and Genera of Curculionoidea (Insecta: Coleoptera) (Excepting Scolytidae and Platypodidae). Entomopraxis, S.C.P., Barcelona, 315 pp.
  2. Anderson, R.S. (2002) Chapter 131. Curculionidae. In: Arnett, R.H. Jr., Thomas, M.C., Skelley, P. & Frank, J.H. (Eds.), American Beetles. Vol. 2. CRC Press, Gainesville, Florida, pp. 722–815.
  3. Anderson, R.S. (2018) A revision of the weevil genus Phyxelis Schoenherr 1843 (Coleoptera: Curculionidae; Entiminae; Tropiphorini) in North America. Insect Systematics and Diversity, 2 (5), 1–11. https://doi.org/10.1093/isd/ixy011
  4. Anderson, R.S. (2019) Taxonomic revision of the genus Paranametis Burke 1960 (Coleoptera: Curculionidae; Entiminae; Byrsopagini). The Coleopterists Bulletin, 73, 966–974. https://doi.org/10.1649/0010-065X-73.4.966
  5. Anderson, R.S. (2025a) A new species of Phyxelis Schoenherr from Arkansas with new synonymy of Paranametis Burke with Phyxelis (Coleoptera: Curculionidae: Entiminae: Byrsopagini). The Coleopterists Bulletin, 79 (4), 688–690.
  6. Anderson, R.S. (2025b) Classification changes in Nearctic Byrsopagini (Coleoptera: Curculionidae: Entiminae). Insecta Mundi, 1160, 1–12. https://doi.org/10.64338/im.1160.308m4
  7. Blatchley, W.S. (1916) In: Blatchley, W.S. & Leng, C.W., Rhynchophora or weevils of North Eastern America. The Nature Publishing Co., Indianapolis, Indiana, 682 pp. https://doi.org/10.5962/bhl.title.122480
  8. Bright, D.E. & Bouchard, P.B. (2008) The Insects and Arachnids of Canada. Part 25. Coleoptera, Curculionidae, Entiminae. Weevils of Canada and Alaska. Vol. 2. NRC Research Press, Ottawa, xiv + 327 pp.
  9. Buchanan, L.L. (1927) Synonymical notes on several Otiorhynchid weevils (Coleoptera). The Canadian Entomologist, 59 (8), 183–184. https://doi.org/10.4039/Ent59183-8
  10. Fall, H.C. (1907) In: Fall, H.C. & Cockerell, T.D.A. The Coleoptera of New Mexico. Transactions of the American Entomological Society, 33, 145–272. [https://www.biodiversitylibrary.org/page/7540873]
  11. Geiser, S.W. (1933) G. W. Belfrage’s Texas localities. Entomological News, 44, 127–130.
  12. Horn, G.H. (1876) In: LeConte, J.L. & Horn, G.H. The Rhynchophora of America, north of Mexico. Proceedings of the American Philosophical Society, 15 (96), i–xvi + 1–455. [https://www.jstor.org/stable/982412]
  13. Kissinger, D.G. (1964) Curculionidae of America North of Mexico. A key to the Genera. Taxonomic Publications, South Lancaster, Massachusetts, v + 143 pp.
  14. Muñiz Velez, R. (2008) Sinopsis de las especies mexicanas de Amphidees (Curculionidae, Entiminae, Tropiphorini). Dugesiana, 15, 117–122.
  15. O’Brien, C.W. & Wibmer, G.J. (1982) Annotated checklist of the weevils (Curculionidae sensu lato) of North America, Central America, and the West Indies (Coleoptera: Curculionoidea). Memoirs of the American Entomological Institute, 34, 1–382.
  16. Pierce, W.D. (1907) On the biologies of the Rhynchophora of North America. Studies from the Zoological Laboratory, the University of Nebraska, Lincoln, 1907, 249–319.
  17. Pierce, W.D. (1913) Miscellaneous contributions to the knowledge of the weevils of the families Attelabidae and Brachyrhinidae. Proceedings of the United States National Museum, 45, 365–426. https://doi.org/10.5479/si.00963801.1988.365
  18. Prena, J. (2015) Thomas Say’s weevils, with special reference to his Curculionites pamphlet and the extant type material. Proceedings of the American Philosophical Society, 157 (4), 393–408.
  19. Prena, J. (2018) An annotated inventory of the weevils (Coleoptera: Curculionoidea) described by Thomas Say. Bulletin of the Museum of Comparative Zoology, 161, 1–401. https://doi.org/10.3099/MCZ161-09.1
  20. Say, T.L. (1832) Descriptions of new species of Curculionites of North America, with observations on some of the species already known. Thomas May, New Harmony, Indiana, 30 pp. https://doi.org/10.5962/bhl.title.68600
  21. Schoenherr, C.J. (1842) Genera et species curculionidum, cum synonymia hujus familiae. Species novae aut hactenus minus cognitae, descriptionibus a Dom. Leonardo Gyllenhal, C.H. Boheman. et entomologis aliis illustratae. 6 (2). Roret, Paris, 495 pp. [https://www.biodiversitylibrary.org/page/4115130]
  22. Selander, R.B. & Vaurie, P. (1962) A Gazetteer to Accompany the “Insecta” Volumes of the “Biologia Centrali-Americana”. American Museum Novitates, 2099, 1–70. [http://hdl.handle.net/2246/3415]
  23. Sharp, D. (1891) Insecta. Coleoptera. Rhynchophora. Curculionidae. Attelabinae, Pterocolinae, Allocoryninae, Apioninae, Thecesterninae, Otiorhynchinae [part, Apterae]. In: Sharp, D. & Champion, G.C., Biologia Centrali-Americana, 4 (Part 3), pp. 89–136. [1899–1911, https://www.biodiversitylibrary.org/page/581470]
  24. Sleeper, E.L. (1956) The genus Dichoxenus Horn (Coleoptera, Curculionidae). The Ohio Journal of Science, 56, 165–169. [http://hdl.handle.net/1811/4359]
  25. Thomson, R.T. (1992) Observations on the morphology and classification of weevils (Coleoptera, Curculionoidea) with a key to major groups. Journal of Natural History, 26 (4), 835–891. https://doi.org/10.1080/00222939200770511
  26. Van Dyke, E.C. (1934) New species of North American weevils in the family Curculionidae, subfamily Brachyrhininae. Pan-Pacific Entomologist, 10, 175–191. [https://www.biodiversitylibrary.org/page/56446962]
  27. Van Dyke, E.C. (1935) New species of North American weevils in the family Curculionidae, subfamily Brachyrhininae, II. Pan-Pacific Entomologist, 11, 1–13. [https://www.biodiversitylibrary.org/page/53418882]
  28. Yunakov, N., Girón Duque, J.C., Brown, S., del Río, M.G., Rincón, J., Cortés-Hernández, K. & Cabras, A.A. (2025) World Entiminae Database. Version January 2025. In: Bánki, O., Roskov, Y., Döring, M., Ower, G., Hernández Robles, D.R., Plata Corredor, C.A., Stjernegaard Jeppesen, T., Örn, A., Pape, T., Hobern, D., Garnett, S., Little, H., DeWalt, R.E., Ma, K., Miller, J., Orrell, T., Aalbu, R., Abbott, J., Aedo, C., et al, Catalogue of Life. Version 17 January 2025. Catalogue of Life, Amsterdam. Available from: https://www.checklistbank.org/dataset/307664/source/1166 (accessed 24 December 2025) https://doi.org/10.48580/dgmv5-3f8

How to Cite

Anderson, R.S. (2026) A taxonomic revision of the genus Dichoxenus Horn, 1876 (Coleoptera: Curculionidae: Entiminae: Byrsopagini). Zootaxa, 5756 (1), 1–77. https://doi.org/10.11646/zootaxa.5756.1.1