Abstract
Two new species belonging to the Empis subgenus Empis Linnaeus, 1758 and one new species belonging to the Empis subgenus Coptophlebia Bezzi, 1909 are described and illustrated, all of them from localities in France: Empis (Empis) inaequalis sp. nov., from two sites in the southern Alps (National nature reserve of Ristolas-Mont-Viso, Regional nature reserve of Partias—05); Empis (Empis) pinstoni sp. nov. from four sites in Bourgogne-Franche-Comté (National nature reserve of Sabot de Frotey—70, National nature reserve of Val de Loire—58, National nature reserve of Bois du parc—89, Commune of Solutré-Pouilly—71526); Empis (Coptophlebia) tissoti sp. nov., from Burgundy, commune of Monéteau (89263).
References
- Bahid, F.Z., Kettani, K. & Daugeron, C. (2017) Two new species of the Empis (Leptempis) rustica-group of the Mediterranean basin (Diptera: Empididae: Empidinae). Annales de la Société Entomologique de France, 53 (6), 422–427. https://doi.org/10.1080/00379271.2017.1399085
- Bezzi, M. (1909) Einige neue paläarktische Empis-Arten. Deutsche Entomologische Zeitschrift, 1909 (Beiheft), 85–103.
- Chvála, M. (1994) The Empidoidea (Diptera) of Fennoscandia and Denmark. III—Genus Empis. Fauna entomologica Scandinavica, 29, 1–192. https://doi.org/10.1163/9789004273559_001
- Chvála, M. (2012) Revision of European species of the Empis (s. str.) pennipes group of species (Diptera: Empididae) with description of five new species from Mediterranean and the Caucasus. Studia dipterologica, 18, 173–193.
- Chvála, M. & Pont, A.C. (2015) Revision of the European Empis (s. str.) alpicola-group of species (Diptera: Empididae), with a new synonymy of Rhamphomyia subgenus Aclonempis Collin with the subgenus Empis Linnaeus s. str. Studia dipterologica, 21 (1), 53–68.
- Chvála, M. & Pont, A.C. (2016) Revision of the European Empis (s. str.) simulium-group of species (Diptera: Empididae), with description of four new species from the European mountains. Studia dipterologica, 22 (1), 89–110.
- Collin, J.E. (1961) British Flies. Vol. VI. Empididae. Part II. Hybotinae and Empidinae (except Hilara). Cambridge University Press, Cambridge, 329 pp. [pp. 223–551]
- Daugeron, C. (1997) Découverte du sous-genre Xanthempis Bezzi en Afrique du Nord et description de trois espèces nouvelles (Diptera, Empididae). Annales de la Société Entomologique de France, New Series, 33, 155–164. https://doi.org/10.1080/21686351.1997.12277746
- Daugeron, C. (1999) Monophyly of the genus Leptempis, and description of seven new species of the Empis (Leptempis) rustica-group (Diptera: Empididae). European Journal of Entomology, 96, 439–449.
- Daugeron, C. (2000) The subgenus Xanthempis: new species and taxonomical data (Diptera: Empididae). Annales de la Société Entomologique de France, New Series, 36 (4), 371–388.
- Daugeron, C. (2009) Two new species of Xanthempis Bezzi (Diptera, Empididae, Empidinae) endemic to the Pyrenees. Zootaxa, 2087 (1), 59–64. https://doi.org/10.11646/zootaxa.2087.1.4
- Daugeron, C. & Grootaert, P. (2005) Phylogenetic systematics of the Empis (Coptophlebia) hyalea-group (Insecta: Diptera: Empididae). Zoological Journal of the Linnean Society of London, 145, 339–391. https://doi.org/10.1111/j.1096-3642.2005.00189.x
- Daugeron, C., Plant, A., Shamshev, I., Stark, A. & Grootaert, P. (2011) Phylogenetic reappraisal and taxonomic review of the Empis (Coptophlebia) hyalipennis-group (Diptera: Empididae: Empidinae). Invertebrate Systematics, 25, 254–271.
- Daugeron, C. & Lefebvre, V. (2015) Descriptions of two new species of Empidinae Schiner, 1862 (Diptera: Empididae) from the Mercantour National Park, France. Zoosystema, 37 (4), 605–309. https://doi.org/10.5252/z2015n4a6
- Engel, E.O. & Frey, R. (1956) Empididae. In: Lindner, E. (Ed.), Die Fliegen der palaearktischen Region, Abtheil 4, E. Schweizerbart’sche Verlagsbuchhandlung, Stuttgart, pp. 1–639.
- Kustov, S.Yu. & Shamshev, I.V. (2013) New data on Empis (Empis) chioptera Meigen species group (Diptera: Empididae) from the Caucasus, Russia. Euroasian Entomological Journal, 12 (1), 79–86.
- Kustov, S.Yu. & Mikhaylichenko, T.V. (2013) A new species of the genus Empis Linnaeus, 1758 (Diptera: Empididae) from the Caucasus. Russian Entomological Journal, 22 (1), 71–73. https://doi.org/10.31610/trudyzin/2013.317.1.45
- Kustov, S.Yu. & Shamshev, I.V. (2014a) Two new species of the Empis (s. str.) nigripes group (Diptera: Empididae) from the North-West Caucasus. Proceedings of the Zoological Institute RAS, 318 (2), 177–183. https://doi.org/10.31610/trudyzin/2014.318.2.177
- Kustov, S.Yu. & Shamshev, I.V. (2014b) A review of dance-flies of the Empis (s. str.) pennipes group of species (Diptera: Empididae) from the Caucasus, with descriptions of five new species. Caucasian Entomological Bulletin, 10 (1), 170–184.
- Langlois, D. & Daugeron, C. (2024) Two new species of the genus Empis Linnaeus, 1758 (Diptera, Empididae, Empidinae) from French nature reserves. Zoosystema, 46 (13), 319–325. https://doi.org/10.5252/zoosystema2024v46a13
- Langlois, D. (2025) Liste des Empidoidea (hors Dolichopodidae Latreille, 1809) de 53 sites remarquables de France métropolitaine dont 36 réserves naturelles (Diptera). Revue française d’entomologie Générale, 6 (8), 236–258. https://doi.org/10.5281/zenodo.14914814
- Linnaeus, C. (1758) Systema Naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. 1 (10). Impensis Direct. Laurentii Salvii. Holmiae, Stockholm, 824 pp. https://doi.org/10.5962/bhl.title.542
- McAlpine, J.F. (1981) Morphology and terminology—Adults [Chapter] 2. In: McAlpine, J.F., Peterson, B.V., Shewell, G.E., Teskey, H.J., Vockeroth, J.R. & Wood, D.M. (Co-Ords.), Manual of Nearctic Diptera. Vol. 1. Agriculture Canada Monograph, 27, 9–63.
- National Imagery and Mapping Agency (1997) Department of Defense World Geodetic System 1984, Its Definition and Relationships with Local Geodetic Systems. NIMA TR8350.2. Available from: https://gis-lab.info/docs/nima-tr8350.2-wgs84fin.pdf (accesed 12 November 2025)
- Shamshev, I.V. & Berezhnova, O.N. (2019) A new species and new records of dance flies of the subgenus Empis s. str. (Diptera, Empididae) from European Russia. Russian Entomological Journal, 28 (1), 86–90. https://doi.org/10.15298/rusentj.28.1.15
- Shamshev, I.V. & Ivković, M. (2020) The Empididae (Diptera) of Serbia: faunistic survey and description of a new Empis species. Zootaxa, 4853 (1), 79–98. https://doi.org/10.11646/zootaxa.4853.1.4
- Stuckenberg, B.R. (1999) Antennal evolution in the Brachycera (Diptera), with a reassessment of terminology relating to the flagellum. Studia dipterologica, 6, 33–48.
- Syrovátka, O. (1991a) Revision of H. Loew’s and T. Becker’s types of Empis s. str. species (Diptera, Empididae) in the Berlin and St Petersburg Museums. Mitteilungen aus dem Zoologischen Museum Berlin, 67 (2), 225–278. https://doi.org/10.1002/mmnz.19910670203
- Syrovátka, O. (1991b) Two new species of Empis s. str. (Diptera, Empididae) from central Europe. Acta Entomologica Bohemoslovaca, 88, 139–144.
- Syrovátka, O. (1995) Revision of Empis s. str. (Diptera, Empididae) described by G. Strobl. Part I. European Journal of Entomology, 92, 497–511.
- Syrovátka, O. (2000) Revision of G. Strobl’s types of Empis s. str. species (Diptera, Empididae) Part II. Acta Universitatis Carolinae Biologica, 44, 195–236.
- Syrovátka, O. & Chvála, M. (1986) Revision of J.W. Meigen’s types of Empis s. str. (Diptera, Empididae) of the Paris Museum, with appendix on Macquart’s species. Véstik Českoslonvenské Společnosti zoologické, 50, 231–239.
