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Issue: Vol. 5715 No. 1: 28 Oct. 2025
Type: Article
Published: 2025-10-28
DOI: 10.11646/zootaxa.5715.1.7
Page range: 53-105
Abstract views: 181
PDF downloaded: 10

A study on the taxonomy and distribution of Brahmaea certhia species group (Lepidoptera: Brahmaeidae) using an integrative approach and websites data

Federal Scientific Center of the East Asia Terrestrial Biodiversity; Far Eastern Branch of the Russian Academy of Sciences; pr-t 100-let Vladivostoka; 159; Vladivostok; 690022; Russia.
Zoological Institute of Russian Academy of Sciences; Universitetskaya embankment; 1; St. Petersburg; 199034; Russia
Lepidoptera Brahmaea certhia Brahmaea tancrei

Abstract

This paper reviews the taxonomy and distribution of East Asian moths from Brahmaea certhia species group (Lepidoptera: Brahmaeidae), using an integrative approach and extensive involvement of websites data. A review of the taxonomic history of this group is given. Based on the correlation of morphological and genetic data on the ‘barcode’ fragment of COI gene, the presence of two species in this group is substantiated, Brahmaea certhia (Fabricius, 1793) and Brahmaea tancrei Austaut, 1896, the second of which is represented by two subspecies, Brahmaea tancrei tancrei Austaut, 1896 and Brahmaea tancrei diastemata (Zhang & Yang, 1993). The morphological differences between these taxa are analysed and keys to the species and subspecies are given. The distribution of each taxon is specified and their brief ecological characteristics are reviewed. Some comments on the systematics of the genus Brahmaea Walker, 1855 as a whole are made. The following nomenclature acts are proposed: Brahmaea certhia (Fabricius, 1793) = Saturnia lunulata Bremer & Grey, 1853, stat. resurr., Saturnia undulata Bremer & Grey, 1853, stat. resurr., Brahmaea carpenteri Butler, 1883, stat. resurr., Brahmaea bicolor Matsumura, 1921, stat. resurr., Brahmaea separata Yang & Zhang, 1994, syn. nov., Brahmaea recta Yang & Zhang, 1994, syn. nov., Brahmaea goniata Zhang & Yang, 1994, syn. nov.; Brahmaea tancrei tancrei Austaut, 1896, stat. resurr. = Brahmaea magnificentia Bryk, stat. resurr., Brahmaea jilinensis Zhang, 1988, syn. nov.; Brahmaea tancrei diastemata (Zhang & Yang, 1993), stat. nov.

 

References

  1. Aarvik, L., Bengtsson, B.Å., Elven, H., Ivinskis, P., Jürivete, U., Karsholt, O., Mutanen, M. & Savenkov, N. (2017) Nordic-Baltic Checklist of Lepidoptera. Norwegian Journal of Entomology Supplement, 3, 1–236.
  2. Antoshin, D.A. & Zolotuhin, V.V. (2011) Lemonia strigata Rougeot et Viette, 1978, stat. n., and L. ballioni (Christoph, 1888) (Lepidoptera: Lemoniidae) in a fauna of Europe with taxonomic notes on related species and description of a new subspecies. Eversmannia, 25–26, 9–24. [in Russian]
  3. Antoshin, D.A. & Zolotuhin, V.V. (2013) Taxonomic remarks on the Lemoniidae (Lepidoptera) with description of a new species from Iran. Tinea, 22, 168–174.
  4. Austaut, J.L. (1896) Notice sur le Brahmaea Lunulata [sic] Bremer [sic] et sur une variété nouvelle de cette espèce (Br. Lunulata Bremer, variété Tancrei, Austaut). Le Naturaliste. Revue Illustrée Sciences Naturelles, 18, 98. [in French]
  5. Barma, A.Yu. (2012) Family Brahmaeidae – brahmin moths. In: Averin, A.A., Kalinin, A.Yu., Malikova, E.I., Osipov, P.E., Rubtsova, T.A. & Streltsov, A.N. (Eds.), Fauna of Bastak Nature Reserve. BSPU Press, Blagoveshchensk, pp. 1–242. [in Russian]
  6. Bidzilya, A.V. & Klyuchko, Z.F. (1994) Heterocerous Lepidoptera of the Lazovsky Reserve (An annotated checklist of species). Taras Shevchenko University of Kyiv Publ., Kyiv, 44 pp. [in Russian]
  7. BOLD (2025) Brahmaea. In: Barcode of Life Data System. Available from: https://v3.boldsystems.org/index.php/TaxBrowser_Taxonpage?taxid=95157 (accessed 6 May 2025)
  8. Bouyer, T. (2002) Description de nouveaux Dactyloceras Mell, 1927 (Lepidoptera, Brahmaeidae). Entomologia Africana, 7 (1), 15–24 + 26–27, pls. 2–3. [in French]
  9. Bremer, O. & Grey, W. (1853a) [1852] Diagnoses de Lépidoptères nouveaux, trouvés par MM. Tatarinoff et Gaschkewitsch aux environs de Pekin. In: Motschulsky, V. de (Ed.), Études Entomologiques. Vol. 1. Imprimerie de la Société de Littérature Finnoise, Helsingfors, pp. 58–80. [in Latin and French]
  10. Bremer, O. & Grey, W. (1853b) Beiträge zur Schmetterlings-Fauna des noerdlichen China’s. Druckerei von J. Johnson, St. Petersburg, 23 рp., pls. 4 + 5. [in German]
  11. Bryk, F. (1949) [1948] Zur Kenntnis der Großschmetterlinge von Korea. Pars II. Macrofrenatae II (finis): Notodontidae, Lymantriidae, Saturniidae, Brahmaeidae, Drepanidae, Epiplemidae, Lasiocampidae, Arctiidae, Thyatiridae, Noctuidae, Geometridae. Heteroneura: Fam.: Cochlifiidae, Zygaenidae. Tineoidea: Fam.: Cossidae, Psychidae. Arkiv för Zoologi, 41A (1), 1–225. [in German]
  12. Butler, A.G. (1866a) Note on the Genus Brahmaea of Walker. Proceedings of the zoological Society of London, 1866, 118–121.
  13. Butler, A.G. (1866b) 12. Corrections and addenda to certain papers on lepidoptera published during the years 1865–66; with additional notes on some of the species described. Proceedings of the zoological Society of London, 1866, 451–458.
  14. Butler, A.G. (1873) Description of a new species of Brahmaea, in the Collection of the British Museum. The Entomologist's Monthly Magazinei, 10, 56–57.
  15. Butler, A.G. (1880) Note on the genus Brahmaea of Walker. The Annals and magazine of natural history; zoology, botany, and geology, 5 (5), 188–189. https://doi.org/10.1080/00222938009459401
  16. Butler, A.G. (1883) XII.—On Lepidoptera from Manchuria and the Corea. The Annals and Magazine of Natural History; including Zoology, Botany, and Geology, Series 5, 11 (62), 109–117. https://doi.org/10.1080/00222938309459105
  17. Chu, H.-F. & Wang, L.-Ya. (1977) The Brahmaeidae of China (Lepidoptera). Acta Entomologica Sinica, 20 (1), 83–84. [in Chinese and English]
  18. Conte, A. (1911) 7e Fascicule, Bombycides (Extrait des rapports du Laboratoire d’Études de la Soie). Essai de Classification des Lépidoptères producteurs de Soie, 14, [5]–90, pls. 1–15. [in French]
  19. De Prins, J. & De Prins, W. (2011–2023) Brahmaeidae Swinhoe, 1892. In: Afromoths, online database of Afrotropical moth species (Lepidoptera). Available from: https://www.afromoths.net/moth/159 (accessed 6 May 2025)
  20. Dubatolov, V.V. (2009) Macroheterocera excluding Geometridae and Noctuidae s. lat. (Insecta, Lepidoptera) of Lower Amur. Amurian Zoological Journal, 1 (3), 221–252. [in Russian with English Summary] https://doi.org/10.33910/1999-4079-2009-1-3-221-252
  21. Dubatolov, V.V. (2020) Macromoths (Insecta, Lepidoptera, Macroheterocera) of Anyuysky National Park (Khabarovsk Krai). Amurian Zoological Journal, 12 (4), 490–512. [in Russian with English Summary] https://doi.org/10.33910/2686-9519-2020-12-4-490-512
  22. Dubatolov, V.V. & Dolgikh, A.M. (2007) Macroheterocera (excluding Geometridae and Noctuidae) of the Bolshekhekhtsirskii Nature Reserve (the Khabarovsk suburbs). In: Streltsov, A.N. (Ed.), Fauna of the Far East: collection of research papers. Vol. 6. BSPU Press, Blagoveshchensk, pp. 105–127. [in Russian with English Summary]
  23. Dubatolov, V.V. & Dolgikh, A.M. (2011) New findings of macromoths (Insecta, Lepidoptera, Macroheterocera) in the Nature Reserve Bolshekhekhtsyrskii (the Khabarovsk suburbs) in 2010. Amurian Zoological Journal, 3 (2), 188–195. [in Russian with English Summary] https://doi.org/10.33910/1999-4079-2011-3-2-188-195
  24. Eolingbul (2019) Mountain King Wave Moth. In: Animals/Animals (Mammals, Insects, Reptiles, Birds, etc.). Available from: https://lambsppk.tistory.com/5865480 (accessed 6 May 2025) [in Korean]
  25. Fabricius, J.Ch. (1793) Entomologia Systematica emendata et aucta. Secundum classes, ordines, genera, species adjectis synonimis, locis, observationibus, descriptionibus. Tom. III Pars I. C.G. Proft, fil., Hafniae [Copenhagen], 487 + [1] pp. [in Latin] https://doi.org/10.5962/bhl.title.122153
  26. Fixsen, C. (1887) Lepidoptera aus Korea. In: Romanoff, N.M. (Ed.), Mémoires sur les Lépidoptères. Vol. 3. Imprimerie de M. M. Stassuléwitch, St.-Pétersbourg, pp. 233–356, pls. 13–15. [in German]
  27. GBIF (2023) Brahmaea Walker, 1855. In: Global Biodiversity Information Facility. GBIF Secretariat. GBIF Backbone Taxonomy. Checklist dataset. GBIF, Copenhagen. Available from: https://www.gbif.org/species/1869358 (accessed 6 May 2025)
  28. Graeser, L. (1888) Beiträge zur Kenntniss der Lepidopteren-Fauna des Amurlandes. Berliner Entomologische Zeitschrift, 32 (1), 33–153. [in German] https://doi.org/10.1002/mmnd.47918880407
  29. Gray, J.E. (1831) The Zoological Miscellany. Treuttel, London, 86 pp., 4 pls.
  30. Gromyko, M.N. (1982) Invertebrates. In: Vasilyev, P.G. & Matyushkin, E.I. (Eds.), Flora and fauna of the Sikhote-Alin State Nature Reserve. Nauka, Moscow, pp. 174–186. [in Russian]
  31. Hamilton, C.A., St Laurent, R.A., Dexter, K., Kitching, I.J., Breinholt, J.W., Zwick, A., Timmermans, M.J.T.N., Barber, J.R. & Kawahara, A.Y. (2019) Phylogenomics resolves major relationships and reveals significant diversification rate shifts in the evolution of silk moths and relatives. BMC Evolutionary Biology, 19 (182), 1–13. https://doi.org/10.1186/s12862-019-1505-1
  32. Hampson, G.F. (1893) [1892] The fauna of British India, including Ceylon and Burma. Moths. Vol. 1. Taylor and Francis, London, 527 pp.
  33. Hampson, G.F. (1901) A Classification of a new Family of the Lepidoptera. The Transactions of the Entomological Society of London for the Year 1901, [49] (2), 187–192. https://doi.org/10.1111/j.1365-2311.1901.tb02384.x
  34. Hao, M., Jin, Q., Meng, G., Yang, C., Yang, Sh., Shi, Zh., Tang, M., Liu, Sh., Li, Yi., Li, J., Zhang, D., Su, X., Shih, Ch., Sun, Yi., Wilson, J.-J., Zhou, X. & Zhang, A. (2020) Using full-length metabarcoding and DNA barcoding to infer community assembly for speciose taxonomic groups: a case study. Evolutionary Ecology, 34, 1063–1088. https://doi.org/10.1007/s10682-020-10072-y
  35. Hebert, P.D.N., Cywinska, A., Ball, S.L. & deWaard, J.R. (2003) Biological Identifications through DNA Barcodes. Proceedings of the Royal Society B: Biological Sciences, 270, 313–321. https://doi.org/10.1098/rspb.2002.2218
  36. Heikkilä, M., Mutanen, M., Wahlberg, N., Sihvonen, P. & Kaila, L. (2015) Elusive ditrysian phylogeny: an account of combining systematized morphology with molecular data (Lepidoptera). BMC Evolutionary Biology, 15 (260), 1–27. https://doi.org/10.1186/s12862-015-0520-0
  37. Heo, U.-H. (2012) Guidebook of Moths Larvae. 1. Nature and Ecology, Seoul, 520 pp.
  38. Höge, R. & Renner, F. (2023) Hybriden mit palaearktischen Arten der Gattung Brahmaea Walker, 1855 (Lepidoptera, Brahmaeidae). Neue Entomologische Nachrichten, 81, 1–67. [in German with English Summary]
  39. Holloway, J.D. (2011) The Moths of Borneo (part 2): Families Phaudidae, Himantopteridae and Zygaenidae; revised and annotated checklist. Malayan Nature Journal, 63 (1), 1–601.
  40. Hübner, J. (1816–1826) Verzeichnis bekannter Schmettlinge [sic]. J. Huebner, Augsburg, 431 pp. https://doi.org/10.5962/bhl.title.48607
  41. ICZN (1991) Case 2737. Nassig, W.A. & Nye, W.B. Brahmaea Walker, 1855 (Insecta, Lepidoptera): proposed confirmation of Bombyx certhia Fabricius, 1793 as the type species. Bulletin of Zoological Nomenclature, 48 (2), 137–139. https://doi.org/10.5962/bhl.part.697
  42. ICZN (1992) Opinion 1698. Brahmaea Walker, 1855 (Insecta, Lepidoptera): Bombyx certhia Fabricius, 1793 confirmed as the type species. Bulletin of Zoological Nomenclature, 49 (4), 294.
  43. ICZN (1999) International Code of Zoological Nomenclature. 4th Edition. International Trust for Zoological Nomenclature, London, xxix + 306 pp.
  44. Ignatenko, E.V. (2000) Rare species of insects in the Khingansky State Nature Reserve and surrounding areas. In: Shlotgauer, S.D., Batalov, A.S. & Andronov, V.A. (Eds.), Scientific research in the nature reserves of the Amur region. Dalʹnauka, Vladivostok-Khabarovsk, pp. 86–90. [in Russian with English Summary]
  45. iNaturalist (2025) Brahmin Moths (Genus Brahmaea). In: iNaturalist. Available from: https://www.inaturalist.org/taxa/179911-Brahmaea (accessed 6 May 2025)
  46. Insecta.pro (2007–2025) Genus Brahmaea. In: Insecta.pro. Insects (Insecta) of the World. Available from: https://insecta.pro/taxonomy/565 (accessed 6 May 2025)
  47. Jarvis, Ch.E. & Oswald, Ph.H. (2015) The collecting activities of James Cuninghame FRS on the voyage of Tuscan to China (Amoy) between 1697 and 1699. Notes and Records, 69, 135–153. https://doi.org/10.1098/rsnr.2014.0043
  48. Jpmoth (2024) 68. List of Brahmaea (Brahmaeidae). In: Niwakagamania, Jinbo, U. & Moth Love (Eds.), Jpmoth: An Identification Guide of Japanese Moths Compiled by Everyone. Available from: http://www.jpmoth.org/Brahmaeidae/Brahmaea.html (assessed 23 June 2025) [in Japanese]
  49. Kang, T.H., Kim, S., Hong, K.-J. & Lee, H.S. (2019) DNA barcoding in quarantine inspection: a case study on quarantine insect monitoring for Lepidoptera obtained through quarantine inspection on foreign vessels. Mitochondrial DNA Part B: Resources, 4 (1), 43–48. https://doi.org/10.1080/23802359.2018.1536447
  50. Kirby, W.F. (1892) A Synonymic Catalogue of Lepidoptera Heterocera. (Moths). Vol. I. Sphinges and Bombyces. Gurney & Jackson, London; R. Friedländer & Son., Berlin, XII + 951 pp. https://doi.org/10.5962/bhl.title.9152
  51. Kitching, I.J., Rougerie, R., Zwick, A., Hamilton, C.A., St Laurent, R.A., Naumann, S., Ballesteros, M.L. & Kawahara, A.Y. (2018) A global checklist of the Bombycoidea (Insecta: Lepidoptera). Biodiversity Data Journal, 6, e22236. https://doi.org/10.3897/BDJ.6.e22236
  52. Klots, A.B. (1970) Lepidoptera. In: Tuxen, S.L. (Ed.), Taxonomist’s Glossary of genitalia in Insects. 2nd Edition. Munksgaard, Copenhagen, pp. 115–130. https://doi.org/10.1163/9789004631663
  53. Koshkin, E.S. (2023) Additions to the fauna of Lepidoptera of the Bastak Nature Reserve (Russian Far East): Spring and early summer aspects. Amurian Zoological Journal, 15 (1), 185–204. [in Russian with English Summary] https://doi.org/10.33910/2686-9519-2023-15-1-185-204
  54. Kozlov, M.V. (1985) Morphology and biology of Brahmaea certhia F. (Lepidoptera, Brahmaeidae). In: Zolotarenko, G.S. (Ed.), Arthropods of Siberia and the Far East. Nauka, Novosibirsk, pp. 105–112. [in Russian]
  55. Kristensen, N.P. (2003) Skeleton and muscles: adults. In: Kristensen, N.P. (Ed.), Lepidoptera: Moths and Butterflies 2. Morphology, Physiology, and Development. Handbuch der Zoologie/Handbook of Zoology. IV/36. Walter de Gruyter, Berlin and New York, pp. 39–131. https://doi.org/10.1515/9783110893724.39
  56. Kumar, S., Stecher, G., Suleski, M., Sanderford, M., Sharma, S. & Tamura, K. (2024) MEGA12: Molecular Evolutionary Genetics Analysis Version 12 for adaptive and green computing. Molecular Biology and Evolution, 41, 1–9. https://doi.org/10.1093/molbev/msae263
  57. Kurentsov, A.I. (1939) Injurious moths of the trees and shrubs in the Ussuri Territory. Proceedings of the Gornotayozhnaya Station of the Far Eastern Branch of the USSR Academy of Sciences, 3, 107–210. [in Russian]
  58. Kurentsov, A.I. (1965) The Zoogeography of the Amur Region. Nauka, Moscow-Leningrad, 155 pp.
  59. Kuznetzov, V.I. & Stekolnikov, A.A. (1985) Comparative and functional morphology of the male genitalia of the Bombycoid moths (Lepidoptera, Papilionomorpha: Lasiocampoidea, Sphingoidea, Bombycoidea) and their systematic position. Proceedings of the Zoological Institute USSR Academy of Sciences, 134, 3–48. [in Russian]
  60. Kuznetzov, V.I. & Stekolnikov, A.A. (2001) New approaches to the system of Lepidoptera of the world fauna (based on the functional morphology of abdomen). Nauka, St. Petersburg, 462 pp. [in Russian]
  61. Lee, B.-Y. (2018) Biodiversity of the Five West Sea Islands Baeknyeongdo, Daecheongdo, Socheongdo, Yeonpyeongdo, Soyeonpyeongdo. National Institute of Biological Resources, Incheon, 271 pp.
  62. Lee, B.-W., Lim, J.-S., Park, S.-Y. & Jo, D.-G. (2011) Insect Fauna of Mt. Jang-san, Yeongwol-gun, Gangwon-do, Korea. Journal of Korean Nature, 4 (3), 173–184. https://doi.org/10.7229/jkn.2011.4.3.173
  63. Lee, K.-W. (2015) Caterpillars of Moths in Korea I. National Institute of Biological Resources, Incheon, 303 pp. [In Korean]
  64. Lee, W., Koh, S.-H., Choi, W.I., Jung, Ch.S., Kim, I.-K., Byun, B.-K., Lee, B.-W., Kim, Y.-S., Lim, J., Kim, S., Akimoto, Sh.-I. & Lee, S. (2012) Barcoding forest insect pests in South Korea: Constructing a basic endemic species dataset. Journal of Asia-Pacific Entomology, 15, 363–368. https://doi.org/10.1016/j.aspen.2012.01.008
  65. Leech, J.H. (1889) On the Lepidoptera of Japan and Corea. Part II. Heterocera, Sect. I. Proceedings of the Scientific Meetings of the Zoological Society of London for the Year 1888, 1889, 580–655, pls. 30–32.
  66. Leech, J.H. (1898) XV. Lepidoptera Heterocera from Northern China, Japan, and Corea. The Transactions of the Entomological Society of London for the Year 1898, [46] 3, 261–379. https://doi.org/10.1111/j.1365-2311.1898.tb02377.x
  67. Lemaire, C. & Minet, J. (1998) The Bombycoidea and their relatives. In: Fischer, M. (Ed.), Handbook of Zoology 4. Arthropoda Insecta. Vol. 4. Walter de Gruyter, Berlin and New York, pp. 353. https://doi.org/10.1515/9783110804744.321
  68. Lepiforum (2023) Brahmaea Walker, 1855. In: Lepiforum e.V. Bestimmung von Schmetterlingen und ihren Präimaginalstadien. Available from: https://lepiforum.org/wiki/taxonomy/Bombycoidea/Brahmaeidae/Brahmaea?view=1&regions (accessed 6 May 2025) [in German]
  69. Liu, X.F., Yang, C.H., Han, H.L., Ward, R.D. & Zhang, A.B. (2014) Identifying species of moths (Lepidoptera) from Baihua Mountain, Beijing, China, using DNA barcodes. Ecology and Evolution, 4 (12), 2472–2487. https://doi.org/10.1002/ece3.1110
  70. Macroclub.ru (2025) Brahmeidae. In: Macroid.ru – The MacroClub project. The qualifier-determinant of insects, spiders & etc. Available from: https://macroid.ru/showgallery.php?cat=13611 (accessed 6 May 2025)
  71. Matsumura, S. (1921) Thousand Insects of Japan [Shin Nihon senchu zukai]. Additamenta IV. Keiseisha, Tokyo, pp. 743–962, pls. 54–71. [in Japanese and English]
  72. Mell, R. (1928) 15. Family: Brahmaeidae. In: Seitz, A. (Ed.), Die Groß-Schmetterlinge der Erde. 10. Band, Die Indo-Australischen Spinner und Schwärmer. Alfred Kernen, Stuttgart, pp. 521–522, pl. 56. [in German]
  73. Mell, R. (1930) Beiträge zur Fauna Sinica. V. Die Brahmaeiden und Eupterotiden Chinas. Deutsche Entomologische Zeitschrift, 1929 (5), 337–494, pls. 3–12 [in German] https://doi.org/10.1002/mmnd.48019290502
  74. Mell, R. (1937) Beiträge zur Fauna sinica. XIV. Ergänzungen zur Sphingiden-, Brahmaeiden- und Eupterotidenfauna Chinas (Lep.). Deutsche Entomologische Zeitschrift, 1937 (1/2), 1–19, 1 pl. https://doi.org/10.1002/mmnd.48019370101
  75. Ménétriès, E. (1859) Lépidoptères de la Siberie orientale et en particulier des rives de l'Amour. In: Shrenk, L. (Ed.), Dr. L.V. Schrenck’s Reisen und Forschungen im Amur-Lande. Band 2. 1. Lieferung. Lepidopteren. Kaiserlichen Akademie der Wissenschaften, St.-Pétersbourg, pp. 1–75, 5 pl. [in French] https://doi.org/10.5962/bhl.title.15761
  76. Ménétriès, E. (1863) Descriptions de nouvelles espèces de Lépidoptères de la collection de l'Académie impériale des Sciences. In: Ménétriès, E. (Ed.), Enumeratio corporum animalium Musei imperialis Academiae scientiarum Petropolitanae. Classis insectorum, ordo lepidopterorum. IIIme et dernière partie. Academiae Scientiarum Imperialis, Petropoli, pp. 145–161, pls. 15–18. [in French] https://doi.org/10.5962/bhl.title.12490
  77. Moltrecht, A. (1929) Über die geographische Verbreitung der Macrolepidopteren des Ussuri- und Amur-Gebietes. Zapiski Vladivostokskogo otdela Gosudarstvennogo Russkogo Geograficheskogo obstshestva, 3 (20/2), 5–70. [in Russian and German]
  78. Nässig, W.A. & Paukstadt, U. (1990) The Brahmaeidae of Sumatra. Heterocera Sumatrana, Göttingen, 6, 117–136.
  79. Nässig, W.A. (1980) Ein Beitrag zur Kenntnis der Saturniidae und Brahmaeidae des Iran und der Türkei (Lepidoptera) Teil 1: Brahmaeidae. Nachrichten des Entomologischen Vereins Apollo, N.F., 1 (3/4), 77–91. [in German with English Abstract]
  80. Nässig, W.A. (1994) Vorläufige Anmerkungen zur Saturniiden- und Brahmaeidenfauna von Vietnam mit Neubeschreibungen (Lepidoptera). Nachrichten des entomologischen Vereins Apollo, N.F., 15 (3), 339–358. [in German and English]
  81. Naumann, S. (2000) Brahmaea cf. certhia Fabricius, 1793 [sic] from China. Entomologische Zeitschrift mit Insekten-Börse, Stuttgart, 110 (4), cover illustration.
  82. Naumann, S. (2009) Notes on the Chinese Calliprogonos miraculosa Mell, 1937 (Lepidoptera: Brahmaeidae). Nachrichten des Entomologischen Vereins Apollo, N.F., 30 (1/2), 5–8.
  83. Naumann, S. & Brosch, U. (2005) Zur verbreitung der hearseyi-Gruppe (Lepidoptera: Brahmaeidae: Brahmaea Walker, 1855) in Indonesien und auf den Philippinen. Galathea, 21 (2), 97–104. [in German]
  84. Novomodnyi, E.V. (2018) The activity of the agents employed by Rudolf Tancre tasked with collecting natural samples in the Russian Far East. A.I. Kurentsov's Annual Memorial Meetings, 29, 5–22. [in Russian with English Summary] https://doi.org/10.25221/kurentzov.29.1
  85. Novomodnyi, E.V. & Shergalin, J.E. (2019) The collectors-naturalist’s of Rückbeil family dynasty: human fates and fates of their collected materials. A.I. Kurentsov's Annual Memorial Meetings, 30, 19–38. [in Russian with English Summary] https://doi.org/10.25221/kurentzov.30.02
  86. Oberprieler, R.G. & Duke, N.J. (1994) The life history and immature stages of Spiramiopsis comma Hampson, 1901 (Lepidoptera: Bombycoidea), with comments on its taxonomic position and on preimaginal characters of the Bombycoidea. Nachrichten des Entomologischen Vereins Apollo, N.F., 15 (3), 199–244.
  87. Owada, M. (1987) Brahmaeidae. In: Sugi, S. (Ed.), Larvae of Larger Moths in Japan. Kodansha, Tokyo, pp. 120–122, pl. 51. [in Japanese]
  88. Park, K.T., Kim, S.S., Tshistjakov, Yu.A. & Kwon, Y.D. (1999) Illustrated Catalogue of Moths in Korea (I). In: Park, K.T. (Ed.), Insects of Korea. Series 4. Korea Research Institute of Bioscience and Biotechnology & Center for Insect Systematics, Chuncheon, pp. 1–358, 23 pls.
  89. Park, Ch.-W. (2017) King Wave Moth. In: Flowering of the Moonflower. Ecology/Insects. Available from: https://ecochulwoo.tistory.com/2010 (accessed 6 May 2025) [in Korean]
  90. Park, K.T., Ronkay, L., Przybylowicz, L., Kun, A. & Peregovits, L. (2001) Moths of North Korea (Lepidoptera, Heterocera, Macrolepidoptera – parts). In: Park, K.T. (Ed.), Insects of Korea. Series 7. Korea Research Institute of Bioscience and Biotechnology & Center for Insect Systematics, Chuncheon, pp. 1–443.
  91. Paukstadt, U. & Aussem, B. (2021) An interesting aberration of Brahmaea (Brahmaea) certhia White, 1862 [“1861”] of the hearseyi-group of the subgenus Brahmophthalma Mell, 1928 (Lepidoptera: Brahmaeidae). Beiträge zur Kenntnis der wilden Seidenspinner, 19 (15), 654–656. [in German with English Summary]
  92. Paukstadt, U. & Paukstadt, L.H. (2017a) Die Präimaginalstadien von Brahmaea (Brahmaea) certhia (Fabricius, 1793) von Jiangsu, P. R. China (Lepidoptera: Brahmaeidae). Beiträge zur Kenntnis der wilden Seidenspinner, 15 (2), 47–72. [in German with English Summary]
  93. Paukstadt, U. & Paukstadt, L.H. (2017b) Die Präimaginalstadien von Brahmaea (Brahmaea) tancrei Austaut, 1896 von Russland (Lepidoptera: Brahmaeidae). Beiträge zur Kenntnis der wilden Seidenspinner, 15 (2), 73–88. [in German with English Summary]
  94. Paukstadt, U. & Paukstadt, L.H. (2021a) A Preliminary Annotated Checklist of the Brahmaeidae of the World – Part IIA. Brahmaea hearseyi White, 1862 [“1861”] of the hearseyi-group of the subgenus Brahmophthalma Mell, 1928 (Lepidoptera: Brahmaeidae). Beiträge zur Kenntnis der wilden Seidenspinner, 19 (4), 127–168.
  95. Paukstadt, U. & Paukstadt, L.H. (2021b) A preliminary annotated checklist of the Brahmaeidae of the world – part IIIA. B. (Brahmaea) certhia (Fabricius, 1793) and its synonyms of the certhia-group of the subgenus Brahmaea Walker, 1855 (Lepidoptera: Brahmaeidae). Beiträge zur Kenntnis der wilden Seidenspinner, 19 (7), 255–294.
  96. Paukstadt, U. & Paukstadt, L.H. (2021c) A preliminary annotated checklist of the Brahmaeidae of the world – part IIIB. Further taxa of the certhia-group of the subgenus Brahmaea Walker, 1855 (Lepidoptera: Brahmaeidae). Beiträge zur Kenntnis der wilden Seidenspinner, 19 (8), 299–348.
  97. Paukstadt, U. & Paukstadt, L.H. (2021dc) A Preliminary Annotated Checklist of the Brahmaeidae of the World – Part IVA. Brahmaea (Brahmaea) europaea Hartig, 1963 (Lepidoptera: Brahmaeidae). Beiträge zur Kenntnis der wilden Seidenspinner, 19 (9), 351–374.
  98. Paukstadt, U. & Paukstadt, L.H. (2021e) A preliminary annotated checklist of the Brahmaeidae of the world – part VI. Systematics and literature (Lepidoptera: Brahmaeidae). Beiträge zur Kenntnis der wilden Seidenspinner, 19 (16), 659–704.
  99. Paukstadt, U. & Paukstadt, L.H. (2022) A preliminary annotated checklist of the Brahmaeidae of the world – part VI – 1st supplement: updated checklist of the names in the family Brahmaeidae Swinhoe, 1892 (Lepidoptera). Beiträge zur Kenntnis der wilden Seidenspinner, 20 (3), 71–84.
  100. Paukstadt, U. & Ragus, G. (1990) Ein Beitrag zur Kenntnis von Brahmaea (Brahmaea) tancrei Austaut, 1896 aus südkoreanischen Populationen (Lepidoptera: Brahmaeidae). Entomologische Zeitschrift, Essen, 100 (1/2), 11–30. [in German with English Summary]
  101. Paukstadt, U. & Weritz, U. (2021a) Interessante Zeichnungs- und Farbaberrationen bei Arten der Untergattung Brahmaea Walker, 1855 (Lepidoptera: Brahmaeidae). Beiträge zur Kenntnis der wilden Seidenspinner, 19 (15), 637–641. [in German with English Abstract]
  102. Paukstadt, U. & Weritz, U. (2021b) Die Brahmaea-Hybriden certhia (Fabricius, 1793) x lunulata carpenteri Butler, 1883 und umgekehrt (Lepidoptera: Brahmaeidae). Beiträge zur Kenntnis der wilden Seidenspinner, 19 (15), 642–645. [in German with English Abstract]
  103. Peters, H. (1998) Zuchtbericht von Brahmaea tancrei Austaut, 1896 aus den Amurlanden (Ostsibirien). Mitteilungen des Internationalen Entomologischen Vereins, 22 (3–4), 173–177. [in German]
  104. Petiver, J. (1702–1706) Gazophylacii naturae & artis decades. [Decas prima] – Decas quinta. Ex Officina Christ. Bateman, Londini, 96 pp. [in Latin]
  105. Petiver, J. (1709) Catalogus classicus & topicus, omnium rerum figuratarum in V. decadibus, seu primo volumine Gazophylacii naturæ & artis; singulis ad proprias tabulas & numeros relatis. Ex Officina Christ. Bateman, Londini, 4 pp., 50 pl. [in Latin]
  106. Pittaway, T. (2007) Brahmaea certhia (Sino-Korean owl moth, male). In: Flickr. The best place to be a photographer online. Available from: https://www.flickr.com/photos/16504650@N00/345875321/ (accessed 6 May 2025)
  107. Prozorov, A.M., Prozorova, T.A., Volkova, J.S., Yakovlev, R.V., Saldaitis, A., Ortiz, A.S., Bianco, G., Schneider, T., Revay, E.E. & Müller, G.C. (2022a) Description of a new Middle Eastern Lemonia species, comments on the Lemonia philopalus complex and Lemonia syriensis (Lepidoptera: Brahmaeidae: Lemoniinae). Ecologica Montenegrina, 57, 77–97. https://doi.org/10.37828/em.2022.57.8
  108. Prozorov, A.M., Prozorova, T.A., Volkova, J.S., Yakovlev, R.V., Nedoshivina, S.V., Pinzari, M., Scalercio, S., Bianco, G., Saldaitis, A., Hausmann, A., Revay, E.E. & Müller, G.C. (2022b) Revision of the Lemonia taraxaci complex, with a description of a new species from Italy and clarification of the status of Lemonia strigata (Lepidoptera: Brahmaeidae: Lemoniinae). Zootaxa, 5195 (4), 337–360. https://doi.org/10.11646/zootaxa.5195.4.2
  109. Rajaei, H., Aarvik, L., Arnscheid, W.R., Baldizzone, G., Bartsch, D., Bengtsson, B.Å., Bidzilya, O., Buchner, P., Buchsbaum, U., Buszko, J., Dubatolov, V.V., Erlacher, S., Esfandiari, M., Freina, J.J. De, Gaedike, R., Gyulai, P., Hausmann, A., Haxaire, J., Hobern, D., Hofmann, A., Ignatev, N., Kaila, L., Kallies, A., Keil, T., Kiss, Á., Kitching, I.J., Kun, A., László, G.M., Leraut, G., Mally, R., Matov, A., Meineke, J.-U., Melichar, T., Mey, W., Mironov, V., Müller, B., Naderi, A., Nässig, W.A., Naumann, S., Nazari, V., Nieukerken, E.J. Van, Nuss, M., Pöll, N., Prozorov, A.M., Rabieh, M.M., Rákosy, L., Rindoš, M., Rota, J., Rougerie, R., Schintlmeister, A., Shirvani, A., Sihvonen, P., Simonsen, T.J., Sinev, S.Yu., Skou, P., Sobczyk, T., Sohn, J.-C., Tabell, J., Tarmann, G., Tokár, Z., Trusch, R., Varga, Z., Volynkin, A.V., Wanke, D., Yakovlev, R.V., Zahiri, R., Zehzad, P., Zeller, H.C. & Karsholt, O. (2023) Catalogue of the Lepidoptera of Iran. In: Rajaei, H. & Karsholt, O. (Eds.), Lepidoptera Iranica. Integrative Systematics, 6 (Special Issue), pp. 121–459. https://doi.org/10.18476/2023.997558.7
  110. Regier, J.C., Mitter, C., Zwick, A., Bazinet, A.L., Cummings, M.P., Kawahara, A.Y., Sohn, J.-C., Zwickl, D.J., Cho, S., Davis, D.R., Baixeras, J.L., Brown, J., Parr, C., Weller, S., Lees, D.C. & Mitter, K.T. (2013) A large-scale, higher-level, molecular phylogenetic study of the insect order Lepidoptera (moths and butterflies). PLoS ONE, 8, e58568. https://doi.org/10.1371/journal.pone.0058568
  111. Rogenhofer, A. & Mann, J. (1873) Neue Lepidopteren gesammelt von Herrn J. Haberhauer. Verhandlungen der kaiserlich-königlichen zoologisch-botanischen Gesellschaft in Wien, 23, 569–574. [in German]
  112. Sauter, W. (1986) Zur Morphologie von Acanthobrahmaea europaea (Hartig, 1963) und zur systematischen Gliederung der Brahmaeidae (Lepidoptera): Dactyloceratinae, subfam. n. Nota Lepidopterologica, 9 (3–4), 262–271. [in German with English Summary]
  113. Seitz, A. (1911) Brahmaeidae. In: Seitz, A. (Ed.), The Macrolepidoptera of the world: a systematic account of all the known Macrolepidoptera. Vol. 2. The Palearctic Bombyces & Sphinges. Alfred Kernen, Stuttgart, pp. 227–228, pl. 35: b, c. [1909–1913]
  114. Shchetkin, Yu.L. (1984) Brameya dal'nevostochnaya Brahmaea certia (Fabricius, 1793). In: Borodin, A.M. (Chief Ed.), Red Data Book of USSR. Rare and endangered species of animals and plants. Vol. 1. Lesnaya Promyshlennost Publishers, Moscow, pp. 280–281. [in Russian]
  115. Sinev, S.Yu. (2019) Catalogue of the Lepidoptera of Russia. 2nd Edition. Zoological Institute RAS, St. Petersburg, 447 pp. [in Russian]
  116. Staudinger, O. (1878/1879) Lepidopteren-Fauna Kleinasien’s. Horae Societatis Entomologicae Rossicae, 14, 176–320 (1878), 321–482 (1879). [in German]
  117. Staudinger, O. (1892) Die Macrolepidopteren des Amurgebietes. I. Theil. Rhopalocera, Sphinges, Bombyces, Noctuae. In: Romanoff, N.M. (Ed.), Mémoires sur les Lépidoptères. Vol. 6. Imprimerie de M. M. Stassuléwitch, St.-Pétersbourg, pp. 83–658, pls. 9–14. [in German] https://doi.org/10.5962/bhl.title.10956
  118. Staudinger, O. & Rebel, H. (1901) I Theil. Famil. Papilionidae—Hepialidae. In: Catalog der Lepidopteren des palaearctischen Faunengebietes. R. Friedländer & Sohn, Berlin, pp. 1–256. [in German] https://doi.org/10.5962/bhl.title.120482
  119. Staudinger, O. & Wocke, M. (1871) Catalog der Lepidopteren des europäischen Faunengebiets. Dr. O. Staudinger und in der Königl. Hofbuchhandlung von Hermann Burdach, Dresden, xxxviii + 426 pp. https://doi.org/10.5962/bhl.title.9475
  120. Strand, E. (1913) Brahmaeidae. In: Wagner, H. (Ed.), Lepidopterorum Catalogus. Pars 16. W. Junk, Berlin, pp. 3–6. [in German]
  121. Sugi, Sh. (1987) A summary: general views on immature stages and life-histories of larger moths in Japan. In: Sugi, Sh. (Ed.), Larvae of Larger Moths in Japan. Kodansha, Tokyo, pp. 267–301.
  122. Tschistjakov, Yu.A. (1992) Family Brahmaeidae. In: Tschistjakov, Yu.A. (Ed.), Insects of the Khingansky Reserve. Part 2. Dal’nauka, Vladivostok, pp. 139–140. [in Russian]
  123. Tschistjakov, Yu.A. (1999) Family Brahmaeidae – Brahmaeas, or Wavy Peacock-eyes. In: Lehr, P.A. (Ed.), Key to the insects of Russian Far East. Vol. 5. Trichoptera and Lepidoptera. Part 2. Dal’nauka, Vladivostok, pp. 628–630. [in Russian]
  124. Tschistjakov, Yu.A. (2009) Family Brahmaeidae. In: Storozhenko, S.Yu. (Ed.), Insects of Lazovsky Nature Reserve. Dal’nauka, Vladivostok, pp. 203. [in Russian]
  125. Walker, F. (1855) Lepidoptera Heterocera. In: List of the Specimens of Lepidopterous Insects in the Collection of the British Museum. Part VI. The Trustees of the British Museum (Natural History), London, pp. 1259–1507. https://doi.org/10.5962/bhl.title.58221
  126. Walker, F. (1865) Supplement. Part 2. In: List of the Specimens of Lepidopterous Insects in the Collection of the British Museum. Part XXXII. The Trustees of the British Museum (Natural History), London, pp. 323–706.
  127. Wang, J.-Sh. (2014) Brahmaea porphyrio. In: Flickr. The best place to be a photographer online. Available from: https://www.flickr.com/photos/105015136@N07/15422753818 (accessed 6 May 2025)
  128. Wu, V.-Y. (2017) The Marvelous Moths of China. Henan Science and Technology Press, Zhengzhou, 403 pp.
  129. Wootton, R.J. (1979) Function, homology and terminology in insect wings. Systematic Entomology, 4 (1), 81–93. https://doi.org/10.1111/j.1365-3113.1979.tb00614.x
  130. Yang, J.-K. (1978) Brahmaeidae. In: Yang, J.-K. (Ed.), Moths of North China. Vol. 2. Beijing Agricultural University Press, Beijing, pp. 424–427.
  131. Yoon, I.-H. (1985) On two Korean species of Brahmaea. Abstracts of the general lecture at the 31st General Meeting of the Lepidopterological Society of Japan (1984, Tokyo), date of lecture: November 18. Transactions of the Lepidopterological Society of Japan (Tyô To Ga), 36 (1), 36–37. [in Japanese]
  132. Yoon, I.-H. (1991) Life history of Brahamaeidae from Korea. Journal of the Amateur Lepidopterists Society of Korea, 4, 29–31. [in Korean]
  133. Yun, Z.-N. (2014) Brachygnatha christophi ? ?. In: Flickr. The best place to be a photographer online. Available from: https://www.flickr.com/photos/stray_bird726/13930952444 (accessed 6 May 2025) [in Chinese]
  134. Zhang, X., Hao, H. & Yang, J. (1999) Genus and Distribution of Brahmaeidae (Lepidoptera) from China. Journal of China Agricultural University, 4 (5), 37–41. [in Chinese with English summary]
  135. Zhang, X.-R. & Yang, C.-K. (1993) A new genus and species of Brahmaeidae (Lepidoptera). Entomotaxonomia, 15 (1), 48–52. [in Chinese and English]
  136. Zhang, X.-R. & Yang, C.-K. (1994a) Two new species of Brahmaea (Lepidoptera: Brahmaeidae) from China. Entomotaxonomia, 16 (2), 111–114. [in Chinese]
  137. Zhang, X.-R. & Yang, C.-K. (1994b) A new species of Brahmaea (Lepidoptera: Brahmaeidae) from China. Entomotaxonomia, 16 (4), 275–277. [in Chinese and English]
  138. Zhang, X.-R. (1988) A new species of Brahmaea from China (Lepidoptera: Brahmaeidae). Entomotaxonomia, 10 (3–4), 229–23. [in Chinese and English]
  139. Zhu, H.-G. [Chu, H.-F.] & Wang, L. (1983) Brahmaeidae. In: Wang, P., Wang, L., Fang, C., Pai, K., Zhu, H. [Chu, H.F.], Liu, Y., Liu, X., Chen, Y., Shen, G., Zhang, B., Zhao, Z., Hou, T. & Cai, R. (Eds.), Iconographia Heterocerorum Sinicorum, IV. Science Press, Beijing, pp. 414–415, col.-figs. 2985–2991. [in Chinese]
  140. Zolotuhin, V.V. (2008) Brahmaeidae. In: Sinev, S.Yu. (Ed.), Catalogue of the Lepidoptera of Russia. KMK Scientific Press Ltd., St. Petersburg-Moscow, pp. 230 + 340. [in Russian]
  141. Zolotuhin, V.V. (2016) To the nomenclature of the certhia Fabricius, 1793 species group of the genus Brahmaea Walker, 1855 (Lepidoptera: Brahmaeidae). Eversmannia, 47–48, 4–10. [in Russian with English Summary]
  142. Zolotuhin, V.V. (2019) Brahmaeidae. In: Sinev, S.Yu. (Ed.), Catalogue of the Lepidoptera of Russia. 2nd Edition. Zoological Institute RAS, St. Petersburg, pp. 284 + 388. [in Russian]
  143. Zwick, A. (2008) Molecular phylogeny of Anthelidae and other bombycoid taxa (Lepidoptera: Bombycoidea). Systematic Entomology, 33, 190–209. https://doi.org/10.1111/j.1365-3113.2007.00410.x

How to Cite

Beljaev, E.A. & Makhov, I.A. (2025) A study on the taxonomy and distribution of Brahmaea certhia species group (Lepidoptera: Brahmaeidae) using an integrative approach and websites data. Zootaxa, 5715 (1), 53–105. https://doi.org/10.11646/zootaxa.5715.1.7