Skip to main content Skip to main navigation menu Skip to site footer
Responsive image
Issue: Vol. 5666 No. 2: 28 Jul. 2025
Type: Article
Published: 2025-07-28
DOI: 10.11646/zootaxa.5666.2.1
Page range: 151-174
Abstract views: 69
PDF downloaded: 8

Pygmy grasshoppers (Orthoptera: Tetrigidae) from Sekayu, Terengganu; with the description of a new species

Institute of Biological Sciences; Faculty of Science; Universiti Malaya; Kuala Lumpur 50603; Malaysia
Lee Kong Chian Natural History Museum; Faculty of Science; National University of Singapore; 2 Conservatory Drive; Singapore 117377; Republic of Singapore
Institute of Biological Sciences; Faculty of Science; Universiti Malaya; Kuala Lumpur 50603; Malaysia
Institute of Biological Sciences; Faculty of Science; Universiti Malaya; Kuala Lumpur 50603; Malaysia
Orthoptera distribution Eufalconius

Abstract

A recent orthopteran collection was conducted in Sekayu, Terengganu, Peninsular Malaysia. We describe a new species of Eufalconius Günther, 1938: Eufalconius sekayu Muhammad, Azizan, Abdullah & Tan, sp. nov. This new species can be easily distinguished from the monotypical species of the genus, E. pendleburyi Günther, 1938, by the slender pronotum (smaller pronotal and prozonal width); its lateral lobe curves outwards, forming an acute apex; the hindwings visible from lateral view and reaching or slightly surpassing the apex of its pronotum; and the 12th antennal segment with its basal half white. This represents the second species from the genus, and the first record of the genus outside the Titiwangsa Mountain range, where E. pendleburyi was mostly collected. We also record Potua coronata Bolívar, 1887, Systolederus cinereus Brunner von Wattenwyl, 1893, Xistrella dohrni dohrni Günther, 1939, Coptotettix nigrifemurus Deng & Jiang, 2017, Lamellitettigodes contractus (Bolívar, 1887), Lamellitettigodes sagittatus (Bolívar, 1887), Criotettix bispinosus (Dalman, 1818), Eucriotettix simulans Tan & Storozhenko, 2017, and Loxilobus insidiosus (Bolívar, 1887) in Terengganu for the first time. We also present color variations of Coptotettix nigrifemurus Deng & Jiang, 2017, Criotettix bispinosus (Dalman, 1818), and Systolederus cinereus Brunner von Wattenwyl, 1893.

 

References

  1. Abadi, M. (2024) Unravelling imaginaries of the digital divide among indigenous communities in Taman Negara Malaysia. Digital Theory, Culture & Society, 2 (2), 87–104. https://doi.org/10.61126/dtcs.v2i2.45
  2. Adžić, K. (2021) Pygmy Grasshoppers (Orthoptera: Tetrigidae) of Peninsular Malaysia. Master’s Thesis, University of Zagreb, Zagreb. [unknown pagination]
  3. Adžić, K., Deranja, M., Franjević, D. & Skejo, J. (2020) Are Scelimeninae (Orthoptera: Tetrigidae) monophyletic and why it remains a question?. Entomological News, 129 (2), 128–146. https://doi.org/10.3157/021.129.0202
  4. Benediktov, A.A. (2013) Vibrational signals of the pygmy grasshoppers males (Orthoptera: Tetrigoidea) from Laos and Southern Vietnam. Russian Entomological Journal, 22 (4), 255–257.
  5. Bey-Bienko, G.Y. (1968) Orthoperoid Insects of East Nepal. Entomological Review, Washington, 47 (1), 59–72.
  6. Bhuiyan, M.A.H., Siwar, C., Ismail, S.M. & Islam, R. (2012) Environmental ecotourism for Sustainable Development in Sekayu Recreational Forest, Malaysia: Perception from the Local Communities. Advances in Environmental Biology, 6 (9), 2553–2557.
  7. Blackith, R.E. (1990) Tetrigidae (Orthoptera) with partly exposed wings. Insecta Mundi, 4 (1–4), 87–92.
  8. Blackith, R.E. (1992) Tetrigidae (Insecta; Orthoptera) of Southeast Asia: Annotated Catalogue with Partial Translated Keys and Bibliography. JAPAGA, Wicklow, iiv + 248 pp.
  9. Blackith, R.E. & Blackith, R.M. (1987) Tridactylids and Tetrigids (Orthoptera) from Sulawesi, Indonesia. Tijdschrift voor Entomologie, 130 (1), 1–10.
  10. Bolívar, I. (1887) Essai sur les Acridiens de la tribu des Tettigidae. Annales de la Société Entomologique de Belgique, 31, 175–313.
  11. Bolívar, I. (1898) Contributions à l’étude des Acridiens espèces de la Faune indo et austro-malaisienne du Museo Civico di Storia Naturale di Genova. Annali del Museo Civico di Storia Naturale di Genova, 19 (39), 66–101. https://doi.org/10.5962/bhl.part.9541
  12. Bruner, L. (1915) Preliminary catalogue of the Orthopteroid insects of the Philippine Islands. University Studies, 15 (2), 195–281. [1–87]
  13. Brunner von Wattenwyl, C. (1893) Révision du système des Orthoptères et déscription des espèces rapportées par M. Leonardo Fea de Birmanie. Annali del Museo Civico di Storia Naturale di Genova, 13 (33), 1–230. https://doi.org/10.5962/bhl.title.5121
  14. Buzzetti, F.M. & Devriese, H. (2008) On some Oriental Orthoptera, mostly from Myanmar (Insecta: Orthoptera: Ensifera, Caelifera). Bollettino del Museo Civico di Storia Naturale di Venezia, 32, 161–169.
  15. Chopard, L. & Dreux, P. (1966) Contribution à l’étude des Orthoptéroides du Népal. Annales de la Société Entomologique de France, 2, 601–616.
  16. Cigliano, M.M., Braun, H., Eades, D.C. & Otte, D. (2025) Orthoptera Species File. Available from: http://Orthoptera.SpeciesFile.org (accessed 30 January 2025)
  17. Deng, W.-A. (2016) Taxonomic study of Tetrigoidea from China. Huazhong Agricultural University, Wuhan, 341 pp.
  18. Dalman, J.W. (1818) Veterinär Akademiens Handlingen. s.n., s.l. [unknwon pagiantion]
  19. de Haan, W. (1843) Bijdragen tot de kennis der Orthoptera. In: Temminck, C.J. (Ed.), Verhandelingen over de Natuurlijke Geschiedenis der Nederlansche Overzeesche Bezittingen: Vol. 19/20. Commissie bij S. en J. Luchtmans, en C.C. van der Hoek, Leiden, pp. 165–228.
  20. Deng, W.-A. (2016) Taxonomic study of Tetrigoidea from China. Huazhong Agricultural University, Wuhan, 341 pp.
  21. Deng, W.-A. (2021) New species and new synonyms of Criotettix and Eucriotettix from China (Orthoptera: Tetrigidae) with an updated key. Oriental Insects, 55 (4), 447–474. https://doi.org/10.1080/00305316.2020.1837690
  22. Deng, W.-A. & Jiang, G. (2017) Description of a new species of the genus Coptotettix Bolívar (Orthoptera: Tetrigoidea) from Thailand. Entomotaxonomia, 39 (2), 103–106.
  23. Deng, W.-A., Luo, J.-L., Huang, C.-M. & Lin, L.-L. (2023) Review of the genus Formosatettix Tinkham (Orthoptera: Tetrigidae) with description of new taxa, new synonymy and an updated key to species. Zootaxa, 5228 (3), 201–243. https://doi.org/10.11646/zootaxa.5228.3.1
  24. Deng, W.-A., Zheng, Z.-M. & Wei, S.-Z. (2007) Fauna of Tetrigoidea from Yunnan and Guangxi. Guangxi Science & Technology Press, Nanning, 458 pp.
  25. Deng, W.-A., Zheng, Z.-M., Li, X., Lin, M., Wei, S.-Z., Yuan, B.-D. & Lin, L.-L. (2015) The groundhopper fauna (Orthoptera: Tetrigidae) of Shiwanshan (Guangxi, China) with description of three new species. Zootaxa, 3925 (2), 151–178. https://doi.org/10.11646/zootaxa.3925.2.1
  26. Deng, W.-A., Zheng, Z.-M., Wei, S.-Z., Lin, M. & Lin, L.-L. (2015) Two new species of the genus Criotettix Bolívar (Orthoptera: Tetrigidae), with a key to the species of the genus from China. Neotropical Entomology, 44 (4), 448–456. https://doi.org/10.1007/s13744-015-0300-y
  27. Günther, K. (1935) Acrydiinen (Orth. Acrididae) aus dem mittleren Ostrorneo gesammelt von H. C. Siebers 1925. Arbeiten über morphologische und taxonomische Entomologie aus Berlin-Dahlem, 2 (4), 257–263, 7 figs.
  28. Günther, K. (1937) Acrydiinae (Orthopt. Acrididae) von Java, den Kleinen Sunda-Inseln und Nordostaustralien. Revue Suisse de Zoologie, 44, 121–142.
  29. Günther, K. (1937) Orthoptera celebica sarasiniana, Fam. Acrididae, Subfam. Acridiinae. Hortus Botanicus Bogoriensis, Java, Treubia, 16, 165–195. https://doi.org/10.14203/treubia.v16i2.2508
  30. Günther, K. (1938) Revision der Acrydiinae, I. Sectiones Tripetalocerae, Discotettigiae, Lophotettigiae, Cleostrateae, Bufonidae, Cladonotae, Scelimenae verae. Mitteilungen aus dem Zoologischen Museum in Berlin, 23, 299–437.
  31. Günther, K. (1939) Revision der Acrydiinae (Orthoptera), III. Sectio Amorphopi (Metrodorae Bol. 1887, aut.). Abhandlungen und Berichte aus den Staatlichen Museen für Tierkunde und Völkerkunde in Dresden, Band 20, Reihe A: Zoologie, N.F. Band 1, Heft 1, 16–335.
  32. Günther, K. (1940) Die von J. Klapperich, Bonn, in Fokien (Südostchina) gesammelten Dornschrecken (Orth., Acrididae, Acrydiinae). Decheniana, 98 (2), 249–254.
  33. Günther, K. (1941) Revision der Acrydiinenausbeute H. Sauters von Formosa (Orth.). Stettiner Entomologische Zeitung, 102, 145–165, 2 pls.
  34. Günther, K. (1942) Die Acrydiinae der Oxforder Universitats-Expedition nach Sarawak vom Jahre 1932. Eos, Revista española de Entomología, 18, 339–346, 6 figs.
  35. Gupta, S.K. & Chandra, K. (2017) A taxonomic study of the pygmy grasshopper (Orthoptera Tetrigidae) from India with description of a new species. Biodiversity Journal, 8 (2), 739–748.
  36. Hancock, J.L. (1907a) Orthoptera Fam. Acridiidae. Subfam. Tetriginae. In: Wytsman, P. (Ed.), Genera Insectorum. Vol. 48. V. Verteneuil & L. Desmet, Bruxelles, pp. 1–79.
  37. Hancock, J.L. (1907b) Studies of the Tetriginae (Orthoptera) in the Oxford University Museum. Transactions of the Entomological Society of London, 1907, 213–244, pl. 21. https://doi.org/10.1111/j.1365-2311.1907.tb01760.x
  38. Hancock, J.L. (1909) Further studies of the Tetriginae (Orthoptera) in the Oxford University Museum. Transactions of the Entomological Society of London, 1909, 387–426, pl. 22. https://doi.org/10.1111/j.1365-2311.1909.tb02160.x
  39. Hancock, J.L. (1910) Third paper on the Tetriginae (Orthoptera) in the Oxford University Museum. Transactions of the Entomological Society of London, 1910, 346–365, pl. 49. https://doi.org/10.1111/j.1365-2311.1910.tb01173.x
  40. Hancock, J.L. (1913) Studies of Tetriginae (Acrydinae) from Sarawak Museum, Borneo. The Sarawak Museum Journal, 1 (3), 39–54.
  41. Hancock, J.L. (1915) V. Indian Tetriginae (Acrydiinae). Records of the Indian Museum, 11, 55–137. https://doi.org/10.26515/rzsi/v11/i1/1915/163610
  42. Hussin, N.F.M. (2007) The Dragonfly Diversity (Insecta: Odonata) between Two Different Land Use Areas at Sekayu Forest, Terengganu. Undergraduate thesis, Kolej Universiti Sains dan Teknologi Malaysia, Terengganu. [unknown pagination, Universiti Malaysia Terengganu Digital Archive Collection]
  43. Ingrisch, S. & Garai, A. (2001) Orthopteroid Insects from Ganesh Himal, Nepal. Esperiana, Schwanfeld, 8, 755–770.
  44. Jiang, G. & Liang, G.-Q. (2004) Fifty-three species of Tetrigoidea (Orthoptera) from Cenwang Mountain in the western Guangxi Zhuang autonomous region, China. Entomological News, 115 (4), 201–206.
  45. Jiang, G. & Zheng, Z.-M. (1998) Grasshoppers and Locusts from Guangxi. Guangxi Normal University, Guilin, Guangxi, 390 pp.
  46. Johari, J. (2006) Composition and Abundance of Odonates (Insecta: Odonata) Community and their Microhabitat Preferences in Sekayu Recreational Forest Terengganu. Undergraduate thesis, Kolej Universiti Sains dan Teknologi Malaysia, Terengganu. [unknown pagination, Universiti Malaysia Terengganu Digital Archive Collection]
  47. Karny, H.H. (1915) H. Sauter’s Formosa-Ausbeute. Orthoptera et Oothecaria. Supplementa Entomologica, 4, 56–108.
  48. Kasalo, N., Skejo, J. & Husemann, M. (2023) DNA barcoding of pygmy hoppers—The first comprehensive overview of the BOLD Systems’ data shows promise for species identification. Diversity, 15 (6), 1–17. https://doi.org/10.3390/d15060696
  49. Kirby, W.F. (1910) A Synonymic Catalogue of Orthoptera (Orthoptera Saltatoria, Locustidae vel Acridiidae). British Museum (Natural History), London, 674 pp.
  50. Kirby, W.F. (1914) Fauna of British India, including Ceylon and Burma. Orthoptera (Acrididae). London, 276 pp. https://doi.org/10.5962/bhl.title.109305
  51. Li, W., Boonmee, S., Eungwanichayapant, P.D. & Zha, L. (2018) Notes on Gibbotettix and Austrohancockia (Tetrigidae: Cladonotinae) with description of two new species. Entomological News, 127 (4), 303–314. https://doi.org/10.3157/021.127.0401
  52. Lu, Y.-Z. & Zha, L. (2020) A new species of the genus Lamellitettigodes (Orthoptera: Tetrigidae) from PR China, with taxonomic notes on the genus. Zootaxa, 4851 (2), 338–348. https://doi.org/10.11646/zootaxa.4851.2.7
  53. Liang, G.-Q. & Zheng, Z.-M. (1998) Orthoptera Tetrigoidea. Fauna Sinica, Insecta, 12, i–x + 1–278.
  54. Liang, G.-Q. (2002) Two new species of Scelimenidae from Hainan Province (Orthoptera: Tetrigoidea). Acta Entomologica Sinica Supplement, 45, 13–15.
  55. Mahmood, K., Idris, A.B. & Salmah, Y. (2007) Tetrigidae (Orthoptera: Tetrigoidea) from Malaysia with the description of six new species. Acta Entomologica Sinica, 50 (12), 1272–1284.
  56. Muhamad, N. (2006) The Composition and Distribution of Butterfly Community in Sekayu Recreational Forest, Terengganu. Undergraduate thesis, Kolej Universiti Sains dan Teknologi Malaysia, Terengganu. [unknown pagination, Universiti Malaysia Terengganu Digital Archive Collection]
  57. Muhammad, A.A., Tan, M.K., Abdullah, N.A., Azirun, M.S., Bhaskar, D. & Skejo, J. (2018) An annotated catalogue of the pygmy grasshoppers of the tribe Scelimenini Bolívar, 1887 (Orthoptera: Tetrigidae) with two new Scelimena species from the Malay Peninsula and Sumatra. Zootaxa, 4485 (1), 1–70. https://doi.org/10.11646/zootaxa.4485.1.1
  58. Muhammad, A.A., Deranja, M., Adžić, K. & Abdullah, N.A. (2023) Towards a better understanding of the genus Scelimena (Orthoptera, Tetrigidae, Scelimeninae): New insights and notes on the taxonomy, ecology, and physiology of the genus in Peninsular Malaysia. Journal of Orthoptera Research, 32 (1), 55–62. https://doi.org/10.3897/jor.32.91153
  59. Neo, I., Tan, M.K., Cho, T.J.Y. & Yeo, D.C.J. (2023) A faunistic study and taxonomic account of species of pygmy grasshoppers (Orthoptera: Tetrigidae) from Singapore’s last freshwater swamp forest. Journal of Asia-Pacific Biodiversity, 17 (1), 87–116. https://doi.org/10.1016/j.japb.2023.09.006
  60. Otte, D. (1979) The primary types of Orthoptera (Saltatoria, Blattodea, Phasmatodea, and Mantodea) at the Academy of Natural Sciences of Philadelphia. Proceedings of the Academy of Natural Sciences, Philadelphia, 130, 26–87.
  61. Otte, D. (1997) Orthoptera Species File 6. Tetrigoidea and Tridactyloidea (Orthoptera: Caelifera) and Addenda to OSF Vols. 1–5. The Orthopterists’ Society and The Academy of Natural Sciences of Philadelphia, Philadelphia, Pennsylvania, 261 pp.
  62. Paris, M. (1994) Catálogo de tipos de ortopteroides (Insecta) de Ignacio Bolivar, I: Blattaria, Mantodea, Phasmoptera y Orthoptera (Stenopelmatoidea, Rhaphidophoroidea, Tettigonioidea, Grylloidea, Tetrigoidea). Eos, Revista española de Entomología, 69, 143–264.
  63. Rahim, N.A.A., Muhammad, A.A. & Abdullah, N.A. (2024) An insight into the distribution of pygmy grasshoppers (Orthoptera: Tetrigidae) in the Klang Valley, Malaysia. Serangga, 29 (2), 96–109. https://doi.org/10.17576/serangga-2024-2902-08
  64. Shishodia, M.S. (1991a) Taxonomy and zoogeography of the Tetrigidae (Orthoptera: Tetrigoidea) of north eastern India. Records of the Zoological Survey of India, Miscellaneous Publication, Occasional Paper, 140, 1–204.
  65. Shishodia, M.S. (1991b) Grouse locusts (Insecta: Orthoptera: Tetrigidae) of the Kanha National Park, District Mandla, Madhya Pradesh, India. Records of the Zoological Survey of India, Calcutta, 89 (1–4), 101–104. https://doi.org/10.26515/rzsi/v89/i1-4/1991/160996
  66. Shishodia, M.S., Chandra, K. & Gupta, S.K. (2010) An annotated checklist of Orthoptera (Insecta) from India. Records of the Zoological Survey of India, Miscellaneous Publication, Occasional Paper, 314, 1–366.
  67. Skejo, J. & Bertner, P. (2017) No More Dust and Exoskeletons—in Vivo Photographic Records Provide New Data on Eufalconius pendleburyi Günther, 1938 (Orthoptera: Tetrigidae) from the Titiwangsa Mts. Annales Zoologici, 67 (4), 665–672. https://doi.org/10.3161/00034541ANZ2017.67.4.003
  68. Skejo, J., Pushkar, T.I., Kasalo, N., Pavlović, M., Deranja, M., Adžić, K., Tan, M.K., Rebrina, F., Muhammad, A.A., Abdullah, N.A., Japir, R., Chung, A.Y.C. & Tumbrinck, J. (2022) Spiky pygmy devils: revision of the genus Discotettix (Orthoptera: Tetrigidae) and synonymy of Discotettiginae with Scelimeninae. Zootaxa, 5217 (1), 1–64. https://doi.org/10.11646/zootaxa.5217.1.1
  69. Steinmann, H. (1970) Check-list of the Tetricidae (Orthoptera) of the Oriental faunal region. Acta Zoologica Academiae Scientiarum Hungaricae, 16, 215–240.
  70. Storozhenko, S.Y. & Pushkar, T.I. (2017) A new genus of pygmy locusts (Orthoptera: Tetrigidae: Cladonotinae) from the Malay Peninsula. Annales Zoologici, Warszaw, 67 (1), 47–53. https://doi.org/10.3161/00034541ANZ2017.67.1.006
  71. Storozhenko, S.Y., Tan, M.K., Vu, T.T., Le, C.V.C. & Pham, H.-T. (2023) Pygmy grasshoppers (Orthoptera: Tetrigidae) from Bach Mã National Park, central Vietnam. Zootaxa, 5319 (2), 199–223. https://doi.org/10.11646/zootaxa.5319.2.3
  72. Subedi, M. (2023) Groundhoppers (Orthoptera: Tetrigidae) of the wetlands of Ramaroshan, Achham (Nepal) with description of a new species. Journal of Insect Biodiversity, 42 (1), 1–30. https://doi.org/10.12976/jib/2023.42.1.1
  73. Subedi, M. & Kasalo, N. (2023) Aryalidonta itishreea, a new genus and species of Thoradontini (Orthoptera, Tetrigidae) from Nepal honors the Emperor of Laughter. Journal of Orthoptera Research, 32 (1), 63–80. https://doi.org/10.3897/jor.32.94918
  74. Suhaimi, M.A.N. (2007) Comparison of Butterfly (Insecta: Lepidoptera) Diversity Between Two Different Land Use Areas at Sekayu, Terengganu. Undergraduate thesis, Kolej Universiti Sains dan Teknologi Malaysia, Terengganu. [unknown pagination, Universiti Malaysia Terengganu Digital Archive Collection]
  75. Storozhenko, S.Y. & Dawwrueng, P. (2015) New and little-known pygmy grasshoppers (Orthoptera: Tetrigidae) from Thailand. Zootaxa, 4052 (5), 527–554. https://doi.org/10.11646/zootaxa.4052.5.2
  76. Storozhenko, S.Y., Tan, M.K., Vu, T.T., Le, C.V.C. & Pham, H.-T. (2023) Pygmy grasshoppers (Orthoptera: Tetrigidae) from Bach Mã National Park, central Vietnam. Zootaxa, 5319 (2), 199–223. https://doi.org/10.11646/zootaxa.5319.2.3
  77. Tan, M.K., Choi, J. & Shankar, N. (2017) Trends in new species discovery of Orthoptera (Insecta) from Southeast Asia. Zootaxa, 4238 (1), 127–134. https://doi.org/10.11646/zootaxa.4238.1.10
  78. Tan, M.K., Storozhenko, S.Y., Hwang, W.S. & Meier, R. (2017) Integrative taxonomy reveals two sympatric species of the genus Eucriotettix Hebard, 1930 (Orthoptera: Tetrigidae). Zootaxa, 4268 (3), 377–39. https://doi.org/10.11646/zootaxa.4268.3.4
  79. Tan, M.K. & Wahab, R.A. (2018) Preliminary study on the diversity of Orthoptera from Kuala Belalong Field Studies Centre, Brunei Darussalam, Borneo. Journal of Orthoptera Research, 27 (2), 119–142. https://doi.org/10.3897/jor.27.24152
  80. Tinkham, E.R. (1937) Notes on the identity of Formosan Acrydiinae with descriptions of a new genus and two new species (Orth.: Acrid.). Transactions of the Natural History Society of Formosa, 27, 229–243, 1 pl.
  81. Tsai, P. (1933) A note on some Chinese grouse locusts (Subfam. Tetriginae) in the British Museum and author’s collection. Journal of the Agricultural Association of China, Shanghai, 118.
  82. Tumbrinck, J. (2014) Taxonomic revision of the Cladonotinae (Orthoptera: Tetrigidae) from the islands of South-East Asia and from Australia, with general remarks to the classification and morphology of the Tetrigidae and descriptions of new genera and species from New Guinea and New Caledonia. In: Telnov, D. (Ed.), Biodiversity, Biogeography and Nature Conservation in Wallacea and New Guinea, II, pp. 350–396.
  83. Tumbrinck, J. (2015) Weitere Dornschrecken (Insecta: Orthoptera: Tetrigidae) aus Nepal in der Sammlung des Naturkundemuseums Erfurt. In: Hartmann, M. & Weipert, J. (Eds.), Biodiversität und Naturausstattung im Himalaya V. Erfurt. Naturkundemuseum, Erfurt, pp. 275–286.
  84. Tumbrinck, J. (2019) Taxonomic and biogeographic revision of the genus Lamellitettigodes (Orthoptera: Tetrigidae) with description of two new species and additional notes on Lamellitettix, Probolotettix, and Scelimena. Journal of Orthoptera Research, 28 (2), 167–180. https://doi.org/10.3897/jor.28.34605
  85. Vanschuytbroeck, P. (1980) Catalogue et liste du matériel typique des Tetrigidae conservés dans les collections entomologiques de l’Institut royal des Sciences naturelles de Belgique. Bulletin de l’Institut Royal des Sciences Naturelles de Belgique, 52 (22), 1–25.
  86. Wahizatul-Afzan, A., Amirrudin, A. & Raja Noor Balqhis, R.S. (2006a) Diversity of aquatic insects in relation to water quality in stream of Sekayu recreational forest, Terengganu. Proceedings of The National Seminar On Science, Technology And Social Sciences, 22006, 79–286.
  87. Wahizatul-Afzan, A., Julia, J. & Amirrudin, A. (2006b) Diversity and distribution of dragonflies (Insecta: Odonata) in Sekayu recreational forest, Terengganu. Journal of Sustainability Science and Management, 1 (2), 97–106.
  88. Walker, F. (1871) n.k. In: Catalogue of the Specimens of Dermaptera Saltatoria in the Collection of the British Museum. London, pp. 811–850.
  89. Wei, S.-Z., Deng, W.-A. & Lu, X.-Y. (2019) Pygmy grasshoppers of the genus Formosatettix Tinkham, 1937 (Orthoptera: Tetrigidae: Tetriginae). Journal of Natural History, 53 (17–18), 1001–1022. https://doi.org/10.1080/00222933.2019.1609113
  90. Wei, S.-Z., Zheng, Z.-M. & Deng, W.-A. (2007) A review of the genus Criotettix Bolívar (Orthoptera: Scelimenidae) from China. Zootaxa, 1423 (1), 39–49. https://doi.org/10.11646/zootaxa.1423.1.3
  91. Wei, T., Nong, C.-G. & Huang, J. (2014) A preliminary study on grasshoppers in Nonggang Nature Reserve of Guangxi, China. Natural Enemies of Insects, 36 (5), 843–852.
  92. Willemse, C.J.M. (1930) Fauna Sumatrensis (Bijdrage Nr. 62) Preliminary revision of the Acrididae (Orthoptera). Tijdschrift voor Entomologie, 73, 1–210.
  93. Willemse, C.J.M. (1930) List of the species of Acridiidae occuring in Sumatra and adjacent islands. Tijdschrift voor Entomologie, 73, 207–210.
  94. Wu, J. (2023) Malayic Varieties of Kelantan and Terengganu. Netherlands Graduate School of Linguistics.
  95. Xin, L. & Deng, W.-A. (2019) A new species and a new record of Xistrella from China (Orthoptera: Tetrigidae). Acta Entomologica Musei Nationalis Pragae, 59 (2), 453–462. https://doi.org/10.2478/aemnp-2019-0034
  96. Yin, X.-C., Shi, J. & Yin, Z. (1996) Synonymic Catalogue of Grasshoppers and their Allies of the World (Orthoptera: Caelifera). China Forestry Publishing House, Beijing, 1266 pp.
  97. Zheng, Z.-M. & Jiang, G. (1994) Five new species of Tetrigoidea from Guangxi (Orthoptera). Guangxi Sciences, 1 (2), 32–36.
  98. Zheng, Z.-M. & Jiang, G. (2000) Three new species of Tetrigoidea (Orthoptera) from Longzhou’s region of Guangxi. Entomotaxonomia, 22 (2), 85–90.
  99. Zheng, Z.-M. & Li, T. (2001) Two new species of Tetrigoidea (Orthoptera) from Guangxi. Entomotaxonomia, 23 (3), 243–224.
  100. Zheng, Z.-M. & Xie, L.-D. (2000) New species of Tetrigoidea from Xishuangbanna (Orthoptera). Journal of Shaanxi Normal University (Natural Science Edition), 28 (3), 90–95.
  101. Zheng, Z.-M. & Xie, L.-D. (2002) Two new species of the genus Criotettix Bolívar from Guangxi (Tetrigoidea: Scelimenidae). Acta Entomologica Sinica Supplement, 45, 6–8.
  102. Zheng, Z.-M. (1992) Orthoptera: Tetrigidae. In: Insects of the Hengduan Mountains region. Vol. 1. Science Press, Beijing, pp. 82–94.
  103. Zheng, Z.-M. (2004) Orthoptera: Tetrigoidea. In: Insects from Mt. Shiwandashan Area of Guangxi. China Forestry Publishing House, Beijing, pp. 69–89.
  104. Zheng, Z.-M. (2005) Fauna of the Tetrigoidea from Western China. Science Press, Beijing, 501 pp.
  105. Zheng, Z.-M. (2012) Key to the species of Criotettix, Coptotettix and Formosatettix from China, with descriptions of seven new species (Orthoptera: Tetrigoidea). Journal of Shangqiu Normal University, 28 (6), 1–18.
  106. Zheng, Z.-M. (2014) A survey of Tetrigoidea from Hunan province (Orthoptera). Journal of Shaanxi Normal University (Natural Science Edition), 42 (6), 54–59.

How to Cite

Muhammad, A.A., Tan, M.K., Azizan, I.A. & Abdullah, N.A. (2025) Pygmy grasshoppers (Orthoptera: Tetrigidae) from Sekayu, Terengganu; with the description of a new species. Zootaxa, 5666 (2), 151–174. https://doi.org/10.11646/zootaxa.5666.2.1