Abstract
Currently, all Australian and New Zealand “weak-veined” Lasioglossum Curtis 1883 species are placed in the subgenus Austrevylaeus Michener 1965 which has L. sordidum (Smith 1853), a New Zealand species, as the type species. A comparative analysis was conducted for Austrevylaeus against three Northern Hemisphere “weak-veined” subgenera Evylaeus Robertson 1902b, Hemihalictus Cockerell 1897 and Sphecodogastra Ashmead 1899. This resulted in a reclassification of the Australian “weak-veined” Lasioglossum fauna with three new subgenera erected: Cnephalictus new subgenus, Enigmalictus new subgenus and Evyloides new subgenus and Austrevylaeus is now restricted to the New Zealand species. Diagnoses are provided for the Australian and New Zealand “weak-veined” subgenera. All Australian species are described, in some cases with one sex for the first time. In addition, citation lists, type designations, type depositories, synonymy lists, diagnoses, distribution maps, SEMs of female inner metatibial spurs, montage images (specimen and male genitalia), examined material, etymologies of species names and comments on taxonomy and floral visitations are presented. Keys to both sexes for Australian and New Zealand species are provided.
The New Zealand “weak-veined” Lasioglossum fauna contains three species: subgenus Austrevylaeus—L. mataroa Donovan 2007, L. maunga Donovan 2007 and L. sordidum.
The Australian “weak-veined” Lasioglossum fauna contains 11 species. Valid, new taxa and new synonyms include: subgenus Cnephalictus—L. amber new species, L. bernhardti new species, L. contaminatum (Cockerell 1910a) and L. lorienae new species; subgenus Enigmalictus—L. blighi (Cockerell 1915) (junior subjective synonym is Halictus pertasmaniae Rayment 1953 new synonymy), L. disclusum (Cockerell 1914) (junior subjective synonym is H. pertribuarius Rayment 1935 new synonymy), L. excelsum new species, L. exoneuroides (Rayment 1953) (junior subjective synonym is H. portlandensis Rayment 1953 new synonymy), L. melanurus (Cockerell 1919) and L. rufibasis (Cockerell 1923); subgenus Evyloides—L. ewarti (Cockerell 1910a).
A re-evaluation of the number of subgenera and species included in the world-wide Lasioglossum “strong-veined” and “weak-veined” series is presented.
Examination of specimens in the Bishop Museum revealed the first known specimen and new species of a “weak-veined” Lasioglossum from West Papua (Indonesia) with an elongated head length—L. oblongulum new species. The head elongation and subgeneric placement of this species are discussed.
References
- ABRS (2024) Australian Faunal Directory. Australian Biological Resources Study, Canberra. Available from: https://biodiversity.org.au/afd/taxa/04a3cd49-5a82-4e12-a949-14bec6ca3cf (accessed 1 May 2024)
- Aguiar, A.P. & Gibson, G.A.P. (2010) The spatial complexity in describing leg surfaces of Hymenoptera (Insecta), the problem and a proposed solution. Zootaxa, 2415 (1), 54–62. https://doi.org/10.11646/zootaxa.2415.1.5
- Aliev, K.A., Guseinzade, G.A. & Aliev, A.V. (2005) Towards understanding the bee fauna of the family Halictidae (Hymenoptera: Apoidea) of Apsheron Peninsula. Caucasian Entomological Bulletin, 1 (2), 153–157. https://doi.org/10.23885/1814-3326-2005-1-2-153-157
- APNI (2024) Australian Plant Name Index. Available from: https://www.anbg.gov.au/apni/ (accessed 26 February 2024)
- Ascher, J.S. & Pickering, J. (2024) Discover Life bee species guide and world checklist (Hymenoptera: Apoidea: Anthophila). Available from: https://www.discoverlife.org/mp/20q?guide=Apoidea_species (accessed 30 September 2024)
- Ashmead, W.H. (1899) Classification of the bees, or the superfamily Apoidea. Transactions of the American Entomological Society, 26, 49–100.
- AVH (2024) The Australasian Virtual Herbarium: Map view for Lophostemon grandifloras. Available from: https://avh.ala.org.au/occurrences/search?q=lsid%3Ahttps%3A%2F%2Fid.biodiversity.org.au%2Fnode%2Fapni%2F2888397&disableAllQualityFilters=true&qc=data_hub_uid%3Adh9&fq=taxon_name%3A%22Lophostemon%20grandiflorus%22#tab_mapView (accessed 27 March 2024)
- Bernhardt, P., Sage, T., Weston, P., Azuma, H., Lam, M., Thien, L.B. & Bruhl, J. (2003) The Pollination of Trimenia morrei (Trimeniaceae): Floral Volatiles, Insect/Wind Pollen Vectors and Stigmatic Self-incompatibility in a Basal Angiosperm. Annals of Botany, 92, 445–458. https://doi.org/10.1093/aob/mcg157
- Blüthgen, P. (1923) Beiträge zur Kenntnis der Bienengattung Halictus Latr. Archiv für Naturgeschichte, 89 (5), 232–332.
- Brady, S.G., Sipes, S., Pearson, A. & Danforth, B.N. (2006) Recent and simultaneous origins of eusociality in halictid bees. Proceedings of the Royal Society B, 273, 1643–1649. https://doi.org/10.1098/rspb.2006.3496
- Brothers, D.J. (1976) Modifications of the metapostnotum and origin of the “propodeal triangle” in Hymenoptera Aculeata. Systematic Entomology, 1, 177–182. https://doi.org/10.1111/j.1365-3113.1976.tb00036.x
- Cane, J.H. (1987) Estimation of bee size using intertegular span (Apoidea). Journal of the Kansas Entomological Society, 60, 145–147.
- Cockerell, T.D.A. (1897a) New and little-known bees. Transactions of the American Entomological Society, 24, 144–162.
- Cockerell, T.D.A. (1897b) On the generic position of some bees hitherto referred to Panurgus and Calliopsis. The Canadian Entomologist, 29, 287–290. https://doi.org/10.4039/Ent29287-12
- Cockerell, T.D.A. (1898) Tables for the determination of New Mexico bees. Bulletin of the University of New Mexico, 1, 41–73.
- Cockerell, T.D.A. (1905) Descriptions and records of bees. II. Annals and Magazine of Natural History, Series 7, 16 (93), 292–301. https://doi.org/10.1080/03745480509442865
- Cockerell, T.D.A. (1910a) New and little—known bees. Transactions of the American Entomological Society, 36, 199–249.
- Cockerell, T.D.A. (1910b) Descriptions and records of bees. XXXIII. Annals and Magazine of Natural History, Series 8, 6 (34), 356–366. https://doi.org/10.1080/00222931008692862
- Cockerell, T.D.A. (1914) Australian halictine bees. The Entomologist, 47, 242–244. https://doi.org/10.5962/bhl.part.15127
- Cockerell, T.D.A. (1915) Descriptions and records of bees. LXIX. Annals and Magazine of Natural History, Series 8, 16 (92), 96–104. https://doi.org/10.1080/00222931508693693
- Cockerell, T.D.A. (1919) Descriptions and records of bees. LXXXIII. Annals and Magazine of Natural History, Series 9, 3 (13), 118–125. https://doi.org/10.1080/00222931908673805
- Cockerell, T.D.A. (1923) Descriptions and records of bees. XCVIII. Annals and Magazine of Natural History, Series 9, 12 (68), 238–247. https://doi.org/10.1080/00222932308632941
- Cockerell, T.D.A. (1924) Descriptions and records of bees. CI. Annals and Magazine of Natural History, Series 9, 14 (79), 179–185. https://doi.org/10.1080/00222932408633105
- Cockerell, T.D.A. (1929) Descriptions and records of bees—CXX. Annals and Magazine of Natural History, Series 10, 4 (24), 584–594. https://doi.org/10.1080/00222932908673101
- Cockerell, T.D.A. (1932) The bees of Australia. The Australian Zoologist, 7, 206–218.
- Cockerell, T.D.A. (1933) The bees of Australia. The Australian Zoologist, 7, 291–324.
- Crawford, J.C. (1906) Some new species of Halictus. The Canadian Entomologist, 38, 4–6. https://doi.org/10.4039/Ent384-1
- Crawford, J.C. (1910) New Hymenoptera from the Philippine Islands. Proceedings of the United States National Museum, 38, 119–133. https://doi.org/10.5479/si.00963801.38-1733.119
- Cresson, E.T. (1872) Hymenoptera Texana. Transactions of the American Entomological Society, 4, 153–292. https://doi.org/10.2307/25076272
- Curtis, J. (1833) British Entomology. Vol. 10. Privately published, London, 48 pls. [pls. 434–481]
- Dalla Torre, C.G. (1896) Catalogus Hymenopterorum, Hucusque Descriptorum Systematicus et Synonymicus. Vol. X. Apidae (Anthophila). Engelmann, Lipsiae, 643 pp.
- Danforth, B.N. (1989) The evolution of hymenopteran wings: the importance of size. Journal of Zoology, 218, 247–276. https://doi.org/10.1111/j.1469-7998.1989.tb02536.x
- Danforth, B.N. (1999) Phylogeny of the bee genus Lasioglossum (Hymenoptera: Halictidae) based on mitochondrial COI sequence data. Systematic Entomology, 24, 377–393. https://doi.org/10.1046/j.1365-3113.1999.00087.x
- Danforth, B.N. & Ji, S. (2001) Australian Lasioglossum + Homalictus Form a Monophyletic Group: Resolving the “Australian Enigma”. Systematic Biology, 50 (2), 268–283. https://doi.org/10.1093/sysbio/50.2.268
- Danforth, B.N. (2002) Evolution of sociality in a primitively eusocial lineage of bees. Proceedings of the National Academy of Science, 99 (1), 286–290. https://doi.org/10.1073/pnas.012387999
- Danforth, B.N., Conway, L. & Ji, S. (2003) Phylogeny of eusocial Lasioglossum reveals multiple losses of eusociality within a primitively eusocial clade of bees (Hymenoptera: Halictidae). Systematic Biology, 52, 23–36. https://doi.org/10.1080/10635150390132687
- Danforth, B.N., Brady, S.G., Sipes, S.D. & Pearson, A. (2004) Single-copy nuclear genes recover Cretaceous-Age divergences in bees. Systematic Biology, 53 (2), 309–326. https://doi.org/10.1080/10635150490423737
- Danforth, B.N., Eardly, C., Packer, L., Walker, K., Pauly, A. & Randrianambinintsoa, F.J. (2008) Phylogeny of Halictidae with an emphasis on endemic African Halictinae. Apidologie, 39, 86–101. https://doi.org/10.1051/apido:2008002
- Danforth, B.N., Minckley, R.L., Neff, J.L. & Fawcett, F. (2019) The solitary bees: biology, evolution, conservation. Princeton University Press, Princeton, New Jersey, 488 pp. https://doi.org/10.1515/9780691189321
- Davison, P.J. & Field, J. (2016) Social polymorphism in the sweat bee Lasioglossum (Evylaeus) calceatum. Insectes Sociaux, 63, 327–338. https://doi.org/10.1007/s00040-016-0473-3
- Donovan, B.J. (2007) Apoidea (Insecta: Hymenoptera). Fauna of New Zealand, 57, 295. https://doi.org/10.7931/J2/FNZ.57
- Do-pham, T.T., Plateaux-quénu, C. & Plateaux, L. (1984) Étude Comparative des Genitalia Males de Quelques Halictinae (Hymenoptera) Répercussion Éventuelle sur la Classification. Annales de la Société entomologique de France, 20 (1), 3–46. https://doi.org/10.1080/21686351.1984.12278743
- Dorey, J.B. (2018) Bees of Australia: A Photographic Exploration. CSIRO Publishing, Clayton, Victoria, 224 pp. https://doi.org/10.1071/9781486308507
- Dorey, J.B., Schwarz, M.P. & Stevens, M.I. (2019) Review of the bee genus Homalictus Cockerell (Hymenoptera: Halictidae) from Fiji with description of nine new species. Zootaxa, 4674 (1), 1–46. https://doi.org/10.11646/zootaxa.4674.1.1
- Dorey, J.B., Fagan-Jeffries, E.P., Stevens, M.I. & Schwarz, M.P. (2020) Morphometric comparisons and novel observations of diurnal and low-light foraging bees. Journal of Hymenoptera Research, 79, 117–144. https://doi.org/10.3897/jhr.79.57308
- Ebmer, A.W. (1971) Die Bienen des Genus Halictus Latr. s.l. im Grossraum von Linz (Hymenoptera, Apidae) Teil III: Lasioglossum (Evylaeus Rob.). Naturkundliches Jahrbuch der Stadt Linz, 1971, 63–156.
- Ebmer, A.W. (1974) Beiträge zur Kenntnis der Fauna Afghanistans. Halictus Latr. et Lasioglossum Curt., Halictidae, Apoidea, Hymenoptera. Časopis Moravského Musea, 59, 183–210.
- Ebmer, A.W. (1976) Halictus und Lasioglossum aus Marokko. Linzer Biologische Beiträge, 8, 205–266.
- Ebmer, A.W. (1995) Asiatische Halictidae, 3. Die Artengruppe der Lasioglossum carinate-Evylaeus (Insecta: Hymenoptera: Apoidea: Halictidae: Halictinae). Linzer Biologische Beiträge, 27, 525–652.
- Ebmer, A.W. (1997) Asiatische Halictidae, 6. Lasioglossum carinaless–Evylaeus: Ergänzungen zu den Artengruppe von L. nitidiusculum und L. punctatissimum s.l., sowie die Artengruppe des L. marginellum (Insecta: Hymenoptera: Apoidea: Halictidae: Halictinae). Linzer Biologische Beiträge, 29 (2), 921–982.
- Ebmer, A.W. (2002) Asiatische Halictidae—10. Neue Halictidae aus China sowie diagnostische Neubeschreibungen der von Fan and Ebmer 1992 beschriebenen Lasioglossum-Arten (Insecta: Hymenoptera: Apoidea: Halictidae: Halictinae). Linzer Biologische Beiträge, 34 (2), 819–934.
- Ebmer, A.W. (2003) Hymenopterologische Notizen aus Österreich—16 (Insecta: Hymenoptera: Apoidea). Linzer Biologische Beitraege, 35 (1), 313–403.
- Ebmer, A.W., Maeta, Y. & Sakagami, S.F. (1994) Six new halictine bee species from southwest archipelago, Japan (Hymenoptera, Halictidae). Bulletin of the Faculty of Agriculture, Shimane University, 28, 23–36.
- Engel, M.S. (2001) A monograph of the Baltic amber bees and evolution of the Apoidea (Hymenoptera). Bulletin of the American Museum of Natural History, 259, 1–192.
- http://dx.doi.org/10.1206/0003-0090
- Engel, M.S. (2009) Revision of the bee genus Chlerogella (Hymenoptera, Halictidae), Part I: Central American species. Zookeys, 23, 47–75. https://doi.org/10.3897/zookeys.23.248
- Engel, M.S., Hinjosa-Díaz, I.A. & Yáñez-Ordóñez, Y. (2007) The Augochlora–like Dialictus from Guatemala and southern México (Hymenoptera: Halictidae). Acta Zoológica Mexicana, 23, 125–134. https://doi.org/10.21829/azm.2007.233596
- Engler, J.D., Gorman, M., Cappaert, D., Stanton, M., Best, L.R. & Jackson, A.S. (2024) Bees of the Pacific Northwest: Key to Lasioglossum Species for Females of the Subgenera Hemihalictus, Sphecodogastra, and Evylaeus in Oregon (Hymenoptera: Halictidae). Version 1. February 2024. Department of Horticulture, Oregon State University, Corvallis, Oregon, 39 pp. https://doi.org/10.5399/osu/1171
- Gardner, J. & Gibbs, J. (2020) The ‘red-tailed’ Lasioglossum (Dialictus) (Hymenoptera: Halictidae) of the western Nearctic. European Journal of Taxonomy, 725, 1–242. https://doi.org/10.5852/ejt.2020.725.1167
- Gardner, J.D. & Gibbs, J. (2022) New and little-known Canadian Lasioglossum (Dialictus) (Hymenoptera: Halictidae) and an emended key to species. The Canadian Entomologist, 154, 1–37. https://doi.org/10.4039/tce.2021.47
- Gardner, J.D. (2023) Bees of the subgenus Lasioglossum (Dialictus) of the western Nearctic region (Hymenoptera: Halictidae: Halictini). A PhD thesis submitted to the Faculty of Graduate Studies of The University of Manitoba, 939 pp. Available from: https://mspace.lib.umanitoba.ca/items/f7617f07-c18d-4065-9f23-aceff5dc0754 (accessed 30 September 2024)
- GBIF (2025) Secretariat. Lasioglossum buccale (Pérez, 1903) in GBIF Backbone Taxonomy. Checklist dataset. Available from: https://www.gbif.org/species/1354640 (accessed 02 February 2025)
- Genaro, J.A. (2021) Especies nuevas de Lasioglossum Curtis de las altas montañas de las Antillas Mayores y estado actual del conocimiento del taxon Habralictellus Moure y Hurd (Hymenoptera: Apoidea: Halictidae). Insecta Mundi, 853, 1–17. https://doi.org/10.33800/nc.v0i10.28
- Gibbs, J. (2010) Revision of the metallic species of Lasioglossum (Dialictus) in Canada (Hymenoptera, Halictidae, Halictini). Zootaxa, 2591 (1), 1–382. https://doi.org/10.11646/zootaxa.2591.1.1
- Gibbs, J. (2011) Revision of the metallic Lasioglossum (Dialictus) of eastern North America (Hymenoptera: Halictidae: Halictini). Zootaxa, 3073 (1), 1–216. https://doi.org/10.11646/zootaxa.3073.1.1
- Gibbs, J. (2016) Bees of the family Halictidae Thomson, 1869 from Dominica, Lesser Antilles (Hymenoptera: Apoidea). European Journal of Taxonomy, 180, 1–50. https://doi.org/10.5852/ejt.2016.180
- Gibbs, J. (2018) DNA barcoding a nightmare taxon: assessing barcode index numbers and barcode gaps for sweat bees. Genome, 61 (1), 21–31. https://doi.org/10.1139/gen-2017-0096
- Gibbs, J., Brady, S.G., Kanda, K. & Danforth, B.N. (2012) Phylogeny of halictine bees supports origin of eusociality for Halictus and Lasioglossum (Apoidea: Anthophila: Halictidae). Molecular Phylogenetics and Evolution, 65, 926–939. https://doi.org/10.1016/j.ympev.2012.08.013
- Gibbs, J., Packer, L., Dumesh, S. & Danforth, B.N. (2013) Revision and reclassification of Lasioglossum (Evylaeus), Lasioglossum (Hemihalictus) and Lasioglossum (Sphecodogastra) in eastern North America (Hymenoptera: Apoidea: Halictidae). Zootaxa, 3672 (1), 1–117. https://doi.org/10.11646/zootaxa.3672.1.1
- Gibbs, J. & Dumesh, S. (2013) A new species, Lasioglossum (Eickwortia) hienae, from Mexico (Apoidea: Halictidae). Journal of Melittology, 13, 1–11. https://doi.org/10.17161/jom.v0i13.4518
- Graham, L. & Packer, L. (2024) Three New Species of Microsphecodes s.str. (Hymenoptera: Halictidae) with an updated key to species of the subgenus. Zootaxa, 5404 (1), 189–205. https://doi.org/10.11646/zootaxa.5404.1.12
- Hacker, H. (1921) Catalogue of Australian bees. Memoirs of Queensland Museum, 7, 99–163.
- Harris, R.A. (1979) A glossary of surface sculpturing. Occasional Papers in Entomology, 28, 1–31.
- https://zenodo.org/records/26215
- Harvey, M.S., Rix, M.G., Framenau, V.W., Hamilton, Z.R., Johnson, M.S., Teale, R.J., Humphreys, G. & Humphreys, W.F. (2011) Protecting the innocent: studying short-range endemic taxa enhances conservation outcomes. Invertebrate Systematics, 25, 1–10. https://doi.org/10.1071/IS11011
- Hatten, T.D., Griswold, T. & Gibbs, J. (2024) Spatiotemporal variability and foraging behavior of bee visitors to a rare long-lived iteroparous forb, Silene spaldingii (Caryophyllaceae). Scientific Reports, 14 (24667), 1–14. https://doi.org/10.1038/s41598-024-75836-w
- Hettiarachchi, T. (2024) Taxonomic studies of western North American Lasioglossum (Hemihalictus) and a problematic L. (Sphecodogastra) species complex. A Masters thesis submitted to the Faculty of Graduate Studies of The University of Manitoba, Winnipeg, Manitoba, 370 pp. Available from: https://mspace.lib.umanitoba.ca/items/66e2251c-b09c-4997-b13f-265cffc61000 (accessed 27 September 2024)
- Hettiarachchi, T. & Gibbs, J. (2024) A problematic species complex. for Lasioglossum subgeneric diagnostics in North America (Hymenoptera: Halictidae). Zootaxa, 5404 (1), 206–235. https://doi.org/10.11646/zootaxa.5404.1.13
- Hirashima, Y. (1953) The insect fauna of Mt. Ishizuchi and Omogo valley, Iyo, Japan. Four new species of Apoidea (Hymenoptera). Transactions of the Shikoku Entomological Society, 3, 132–138.
- Hopkins, M. (2024) Photographed Lasioglossum bees on iNaturalist. Available from: https://www.inaturalist.org/observations?place_id=any&taxon_id=57678&user_id=maramahopkins&verifiable=any (accessed 19 April 2024)
- Hurd, P.D. Jr. (1979) Catalog of Hymenoptera in America North of Mexico. In: Krombein, K.V., Hurd, P.D. Jr., Smith, D.R. & Burks, B.D. (Eds.), Superfamily Apoidea. Smithsonian Institution Press, Washington, D.C., pp. 1741–2209. https://doi.org/10.1007/BF02223710
- ICZN (2025) International Commission of Zoological Nomenclature. International Code of Zoological Nomenclature. 4th Edition, International Trust for Zoological Nomenclature, 2000. Recommendation 73F. Avoidance of assumption of holotype. Available from: https://code.iczn.org/types-in-the-species-group/article-73-name-bearing-types-fixed-in-the-original-publication-holotypes-and-syntypes/?frame=1 (accessed 30 January 2025)
- Kirby, W. (1802) Monographia Apum Angliae. Vol. 2. Privately published, Ipswich, United Kingdom, 388 pp., 18 pls.
- Knerer, G. & Atwood, C.E. (1964) Further Notes on the Genus Evylaeus Robertson (Hymenoptera: Halictidae). The Canadian Entomologist, 96 (7), 957–962. https://doi.org/10.4039/Ent96957-7
- Latreille, P.A. (1804) Tableau méthodique des insects. In: Nouveau Dictionnaire d’Histoire Naturelle, 24, Détereville, Paris, pp. 129–200.
- Lhomme, P., Michez, D., Christmann, S., Scheuch, E., Abdouni, I.E., Hamroud, L., Ihsane, O., Sentil, A., Smaili, M.C., Schwarz, M., Dathe, H.H., Straka, J., Pauly, A., Schmid-Egger, C., Patiny, S., Terzo, M., Müller, A., Praz, C., Risch, S., Kasparek, M., Kuhlmann, M., Wood, T., Bogusch, P., Ascher, J. & Rasmont, P. (2020) The wild bees (Hymenoptera: Apoidea) of Morocco. Zootaxa, 4892 (1), 1–159. https://doi.org/10.11646/zootaxa.4892.1.1
- Lucas, R. (1910) Hymenoptera für 1910. Archiv für Naturgeschichte, 7, 1–316.
- McGinley, R.J. (1999) Eickwortia (Apoidea, Halictidae), a new genus of bees from Mesoamerica. University of Kansas Natural History Museum Special Publication, 24, 111–120. https://doi.org/10.17161/jom.v0i13.4518
- Meyer, R. (1920) Apidae—Halictinae. I. Gatt. Archiv für Naturgeschichte, 85 (A) (11), 112–137.
- Michener, C.D. (1944) Comparative external morphology, phylogeny, and a classification of the bees (Hymenoptera). Bulletin of the American Museum of Natural History, 82 (6), 151–326.
- Michener, C.D. (1951) Superfamily Apoidea. In: Muesebeck, C.F., Krombein, K.V. & Townes, H.K. (Eds.), Superfamily Apoidea. United States Government Printing Office, Washington, D.C., pp. 1043–1255.
- Michener, C.D. (1965) A classification of the bees of the Australian and South Pacific regions. Bulletin of the American Museum of Natural History, 130, 1–362.
- Michener, C.D. (1974) The Social Behavior of the Bees. Belknap Press, Cambridge, Massachusetts, 404 pp.
- Michener, C.D. (1980) The large species of Homalictus and related Halictinae from the New Guinea area. American Museum Novitates, 2693, 1–21.
- Michener, C.D. (2002) The bee genus Chilicola in the tropical Andes, with observations on nesting biology and a phylogenetic analysis of the subgenera (Hymenoptera: Colletidae, Xeromelissinae). Scientific Papers Natural History Museum University of Kansas, 26, 1–47. https://doi.org/10.5962/bhl.title.8449
- Michener, C.D. (2007) The bees of the world. 2nd Edition. The Johns Hopkins University Press, Baltimore, Maryland, xvi + [i] + 953 pp.
- Morawitz, F. (1876) Bees (Mellifera). II. Andrenidae. In: Fedchenko, A.P. (Ed.), Travel to Turkestan ... Izvestiya Imperatorskogo Obshχhestva Ljubitelei Estestvoznaniya, Antropologii i Etnografii Moscovskogo Universiteta. Proceedings of the Society of Naturalists, Anthropologists, and Ethnographists of the Moscow University, 21 (3) (2), 161–303, pls. 1–3.
- Moure, J.S. & Hurd, P.D. (1982) On two new groups of Neotropical halictine bees (Hymenoptera, Apoidea). Dusenia, 23, 46.
- Murao, R. & Tadauchi, O. (2007) A revision of the subgenus Evylaeus of the Genus Lasioglossum in Japan (Hymenoptera, Halictidae) Part 1. Easkia, 47, 169–254. https://doi.org/10.5109/8325
- Murao, R., Tadauchi, O. & Yamauchi, T. (2009) Taxonomic Revision of the Subgenus Ctenonomia of the Genus Lasioglossum (Hymenoptera, Halictidae) in Japan. Esakia, 49, 75–94. https://doi.org/10.5109/16149
- Murao, R., Lee, H.S. & Tadauchi, O. (2014) Lasioglossum (Acanthalictus) dybowskii (Hymenoptera, Halictidae) newly recorded from South Korea, with a checklist of the genus Lasioglossum in Korean Peninsula. Journal of Hymenoptera Research, 38, 141–153. https://doi.org/10.3897/JHR.38.7572
- Murao, R. (2017a) First Description of the Males of both Lasioglossum bicornutum Murao, 2010 and L. latilabrum Murao & Tadauchi, 2006 (Hymenoptera: Halictidae) Occurring in the Ryukyu Islands, Japan. Japanese Journal of Systematic Entomology, 23 (1), 67–73. https://doi.org/10.11646/zootaxa.2669.1.1
- Murao, R. (2017b) Bees of the Hemihalictus series of Lasioglossum Curtis (Hymenoptera, Halictidae) in South Korea. Zootaxa, 4268 (4), 451–488. https://doi.org/10.11646/zootaxa.4268.4.1
- Murao, R., Tadauchi, O. & Miyanaga, R. (2017) The bee family Halictidae (Hymenoptera, Apoidea) from Central Asia collected by the Kyushu and Shimane Universities Expeditions. Biodiversity Data Journal, 5, 1–43. https://doi.org/10.3897/BDJ.5.e15050
- Murao, R. (2021) Redefinition of the sexstrigatus group of Lasioglossum (Hemihalictus) Cockerell, 1897 (Hymenoptera, Apoidea, Halictidae), with a revision of Japanese species. European Journal of Taxonomy, 763, 1–74. https://doi.org/10.5852/ejt.2021.763.1463
- Murao, R., Yeh, W.C. & Lu, S.S. (2023) A review of Lasioglossum (Leuchalictus) in Taiwan (Hymenoptera: Apoidea: Halictidae). Acta Entomologica, 63 (2), 363–382. https://doi.org/10.37520/aemnp.2023.023
- Niu, Z.-Q., Zhang, D. & Zhu, C.-D. (2020) Extraordinary bees of the genus Lasioglossum Curtis, 1833 (Hymenoptera: Apoidea: Halictidae) from China. Zoological Systematics, 45 (1), 50–58. https://doi.org/10.11865/zs.202005
- Packer, L. (1991) The evolution of social behaviour and nest architecture in sweat bees of the subgenus Evylaeus (Hymenoptera: Halictidae). Behaviorial Ecology and Sociobiology, 29 (3), 153–160. [https://www.jstor.org/stable/4600600] https://doi.org/10.1007/BF00166396
- Packer, L. (1998) A phylogenetic analysis of western European species of the Lasioglossum leucozonium species-group (Hymenoptera: Halictidae): Sociobiological and taxonomic implications. Canadian Journal of Zoology, 76 (9), 1611–1621. https://doi.org/10.1139/z98-102
- Packer, L. (2000) The biology of Thrincohalictus prognathus (Perez) (Hymenoptera: Halictidae: Halictini). Journal of Hymenoptera Research, 9, 53–61.
- Packer, L. (2023) Bees of the world. A guide to every family. Princeton University Press, Princeton, New Jersey, 240 pp. https://doi.org/10.2307/j.ctv2v6pd2p
- Pauly, A. (1984) Paradialictus, un nouveau genre cleptoparasite récolté au Parc National des Virungas (Zaïre) (Hymenoptera Apoidea Halictidae). Revue de Zoologie Africaine, 98 (4), 689–692.
- Pauly, A. (1986) Les abeilles de la sous-famille des Halictinae en Nouvelle-Guinee et dans L’Archipel Bismarck (Hymenoptera: Apoidea: Halictidae). Zoologische Verhandelingen, 227, 1–58.
- Pauly, A. (1999) Classification of the Halictini of the Afrotropical zone (Hymenoptera Apoidea Halictidae). Bulletin de l’Institut Royal des Sciences Naturelles de Belgique, Entomologie, 69, 137–196.
- Pauly, A. (2001) Ipomalictus Pauly, 1999, subgenus of Lasioglossum, new for the Oriental region, with notes on some Afro-tropical species (Hymenoptera: Halictidae). Bulletin de l’Institut Royal des Sciences Naturelles de Belgique, Entomologie, 71, 145–154.
- Pauly, A., Gibbs, J. & Kuhlmann, M. (2012) Capalictus, a new subgenus of Lasioglossum Curtis, 1833 from South Africa, with description of three new species (Hymenoptera, Apoidea, Halictidae). European Journal of Taxonomy, 28, 1–28. https://doi.org/10.5852/ejt.2012.28
- Pauly, A., Walker, K., Munzinger, J. & Donovan, B. (2013) Endémisme insulaire et cleptoparasitisme chez les Lasioglossum Curtis 1833 (Hymenoptera: Apoidea: Halictidae) de Nouvelle-Calédonie. Annales de la Société entomologique de France, New Series, 49, 127–153. https://doi.org/10.1080/00379271.2013.815036
- Pauly, A. (2016a) Le genre Lasioglossum, sous-genre Evylaeus Robertson, 1902, de la Region Paléarctique. Atlas Hymenoptera. Available from: http://www.atlashymenoptera.net/search.aspx?search=longirostre (accessed 24 September 2024)
- Pauly, A. (2016b) Le genre Lasioglossum, sous-genre Evylaeus Robertson, 1902, de la Region Paléarctique. Atlas Hymenoptera. Available from: http://www.atlashymenoptera.net/page.aspx?id=45 (accessed 24 September 2024)
- Pauly, A., Levy, K., Noel, G., Sonet, G., Boeve, J.L. & Yael Mandelik, Y. (2020) Lasioglossum dorchini (Hymenoptera: Apoidea: Halictidae) a new species of bee from Israel. Belgian Journal of Entomology, 105, 1–24.
- Perkins, R.C.L. & Cheesman, L.E. (1928) Insects of Samoa and Other Samoan Terrestrial Arthropoda. Part V. Hymenoptera. Fasc.1. Apoidea, Sphecoidea, and Vespoidea. British Museum (Natural History), London, 32 pp.
- Pesenko, Y.A. (2007a) A taxonomic study of the bee genus Evylaeus Robertson of Eastern Siberia and the Far East of Russia (Hymenoptera: Halictidae). Zoosystematica Rossica, 16 (1), 79–123. https://doi.org/10.31610/zsr/2007.16.1.79
- Pesenko, Y.A. (2007b) Subgeneric classification of the Palaearctic bees of the genus Evylaeus Robertson (Hymenoptera: Halictidae). Zootaxa, 1500 (1), 1–54. https://doi.org/10.11646/zootaxa.1500.1.1
- PlantNET NSW (2024a) “Pollia crispata” Available from: https://plantnet.rbgsyd.nsw.gov.au/cgi-bin/NSWfl.pl?page=nswfl&lvl=sp&name=Pollia~crispata (accessed 18 April 2024)
- PlantNET NSW (2024b) “Psychotria loniceroides” Available from: https://plantnet.rbgsyd.nsw.gov.au/cgi-bin/NSWfl.pl?page=nswfl&lvl=sp&name=Psychotria~loniceroides (accessed 18 April 2024)
- PlantNET NSW (2024c) “Cordyline stricta” Available from: https://plantnet.rbgsyd.nsw.gov.au/cgi-bin/NSWfl.pl?page=nswfl&lvl=sp&name=Cordyline~stricta (accessed 18 April 2024)
- PlantNET NSW (2024d) “Senecio amygdalifolius” Available from: https://plantnet.rbgsyd.nsw.gov.au/cgi-bin/NSWfl.pl?page=nswfl&lvl=sp&name=Senecio~amygdalifolius (accessed 22 April 2024)
- PlantNET NSW (2024e) “Trimenia moorei” Available from: https://plantnet.rbgsyd.nsw.gov.au/cgi-bin/NSWfl.pl?page=nswfl&lvl=sp&name=Trimenia~moorei (accessed 22 April 2024)
- PlantNET NSW (2024f) “Leucopogon affinis” Available from: https://plantnet.rbgsyd.nsw.gov.au/cgi-bin/NSWfl.pl?page=nswfl&lvl=sp&name=Leucopogon~lanceolatus (accessed 22 April 2024)
- Provancher, L. (1888) 1885–1889 Additions et Corrections au Vol. II. de la Faune Entomologique du Canada Traitant des Hyménoptères. Darveau, Quebec, 475 pp. https://doi.org/10.5962/bhl.title.46411
- Rayment, T. (1935) A Cluster of Bees. Sixty essays on the life-histories of Australian bees, with specific descriptions of over 100 new species, and an introduction by Professor E.F. Phillips, D.Ph. Cornell University, Ithaca, New York and Endeavour Press, Sydney, 752 pp.
- Rayment, T. (1953) Bees of the Portland District. Victoria. Portland Field Naturalist’s Club, Portland, Oregon, 39 pp.
- Richards, M.H., Lucas, Z., Proulx, Corbin, L., Jacks, F. & Kehler, D. (2023) Phenology and social status of the endemic Sable Island Sweat Bee, Lasioglossum sablense. Journal of Melittology, 124, 1–156. https://doi.org/10.17161/jom.vi124.22513
- Robertson, C. (1893) Notes on bees, with descriptions of new species. Transactions of the American Entomological Society, 20 (2), 145–149.
- Robertson, C. (1901) Some new or little-known bees. The Canadian Entomologist, 33, 229–231. https://doi.org/10.4039/Ent33229-8
- Robertson, C. (1902a) Some new or little-known bees—II. The Canadian Entomologist, 34, 48–49. https://doi.org/10.4039/Ent3448-2
- Robertson, C. (1902b) Synopsis of Halictinae. The Canadian Entomologist, 34, 243–250. https://doi.org/10.4039/Ent34243-9
- Schenck, A. (1853) Nachtrag zu der Beschreibung nassauischer Bienen. Jahresbücher des Vereins für Naturkunde im Herzogthum Nassau, 9 (1), 88–306.
- Scopoli, I.A. (1763) Entomologia Carniolica; Exhibens Insecta Carnioliae Indigena et Distributa in Ordines, Genera, Species, Varietates. Methodo Linnaeana. J.T. Trattner, Wien, xxxvi + 420 pp. https://doi.org/10.5962/bhl.title.119976
- Sheffield, C.S. & Heron, L.M. (2018) The bees of British Columbia (Hymenoptera: Apoidea, Apiformes). Journal of the Entomological Society of British Columbia, 115, 44–85.
- Shorthouse, D.P. (2010) SimpleMappr, an online tool to produce publication-quality point maps. Available from: https://www.simplemappr.net (accessed 21 August 2024)
- Smith, F. (1853) Catalogue of Hymenopterous Insects in the Collection of the British Museum Part 1. British Museum, London, i + 198 pp., vi pls.
- Strand, E. (1915) Apidae von Tsingtau (Hym.). Mitteilungen der Münchner Entomologischen Gesellschaft, 4 (1–3), 62–78.
- Su, Y.N. (2016) A simple and quick method of displaying liquid-preserved morphological structures for microphotography. Zootaxa, 4208 (6), 592–593. https://doi.org/10.11646/zootaxa.4208.6.6
- Timmermann, K. & Kuhlmann, M. (2008) The biology of a Patellapis (s. str.) species (Hymenoptera: Apoidea: Halictidae): sociality described for the first time in this bee genus. Apidologie, 39, 189–197. https://doi.org/10.1051/apido:2008003
- USNM (2024) United States National Museum. Images of Halictus melanurus type specimen. Available from: https://collections.si.edu/search/results.htm?q=record_ID=nmnhentomology_9172796&repo=DPLA (accessed 7 October 2024)
- Vachal, J. (1903) Hyménoptères rapportés du Japon par M. Harmand., Mellifères. Bulletin du Muséum national d’histoire naturelle, 9, 129–132.
- Walker, K.L. (1986) Revision of the Australian species of the genus Homalictus Cockerell (Hymenoptera: Halictidae). Memoirs of Museum Victoria, 47, 105–200. https://doi.org/10.24199/j.mmv.1986.47.05
- Walker, K.L. (1995) Revision of the Australian native bee subgenus Lasioglossum (Chilalictus) (Hymenoptera: Halictidae). Memoirs of the Museum of Victoria, 55 (Parts 1 & 2), 1–214 & 215–423. https://doi.org/10.24199/j.mmv.1995.55.01 https://doi.org/10.24199/j.mmv.1995.55.02
- Walker, K.L. (2022a) Taxonomic revision of the Australian native bee subgenus Callalictus (Hymenoptera: Halictidae: Halictini: genus Lasioglossum). Australian Journal of Taxonomy, 7, 1–19. https://doi.org/10.54102/ajt.mk64x
- Walker, K.L. (2022b) Taxonomic revision of the Australian native bee subgenus Australictus (Hymenoptera: Halictidae: Halictini: genus Lasioglossum)—“Wood-Splitting Axe Bees”. Memoirs of Museum Victoria, 81, 135–162. https://doi.org/10.24199/j.mmv.2022.81.06
- Walker, K.L. & Sparks, K.S. (2024) Taxonomic revision of the native bee subgenus Parasphecodes Smith 1853 in Australia (Hymenoptera: Halictidae: Halictini: Lasioglossum Curtis 1833). Zootaxa, 5408 (1), 001–184. https://doi.org/10.11646/zootaxa.5408.1.1
- Warncke, K. (1975) Beitrag zur Systematik und Verbreitung der Furchenbienen in der Türkei (Hymenoptera, Apoidea, Halictus). Polskie Pismo Entomologiczne, 45 (1), 81–128.
- Wcislo, W.T. & Tierney, S.M. (2009) Behavioural environments and niche construction: the evolution of dim-light foraging in bees. Biological Reviews, 84, 19–37. https://doi.org/10.1111/j.1469-185X.2008.00059.x
- Wheeler, W.M. (1928) The Social Insects: Their Origin and Evolution. Kegan Paul, Trench, Trubner and Co., Ltd. London, 24 pp. https://doi.org/10.5962/bhl.title.140774
- Zhang, D., Niu, Z.-Q., Luo, A.R., Orr, M.C., Ferrari, R.R., Jin, J.-F., Wu, Q.-T., Zhang, F. & Zhu, C.-D. (2022) Testing the systematic status of Homalictus and Rostrohalictus with weakened cross-vein groups within Halictini (Hymenoptera: Halictidae) using low-coverage whole-genome sequencing. Insect Science, 29 (6), 1819–1833. https://doi.org/10.1111/1744-7917.13034