Abstract
A taxonomic and faunistic review of the genus Omalium Gravenhorst, 1802 of Siberia and Far Eastern Russia is presented, including notes on some species from Mongolia and Japan. Four species groups are established for 16 species of the genus: Amplissimum group containing O. longicorne Luze, 1906 (Mongolia, eastern Siberia); Caesum group containing the Palaearctic O. caesum Gravenhorst, 1806 and O. golovatchi sp. nov. (Russia: Kemerovo Area); Oxyacanthae group containing the Holarctic O. oxyacanthae Gravenhorst, 1806 and Palaearctic O. septentrionis Thomson, 1857; Rivulare group containing the Palaearctic O. curtipenne Mäklin, 1878, Japanese O. diffine Sharp, 1889, O. hibernum Hayashi, 2006, as well as O. shibatai Watanabe, 1990 and O. kurilicum sp. nov. (Russia: Kuril islands: Iturup, Kunashir), O. japonicum Sharp, 1874 (Japan, South Korea), O. niponense Sharp, 1889 (Far Eastern Russia, Japan), Holarctic O. rivulare (Paykull, 1789) and O. strigicolle Wankowicz, 1869, Mongolian O. scabrum Smetana, 1975, and O. subsolanum Herman, 2001 (Siberia, Japan); and unspecified group containing the Japanese O. tenue Weise, 1877. All new species, and O. curtipenne, O. diffine, O. longicorne, O. niponense and O. subsolanum are (re-)described and illustrated. A new synonymy is established: O. curtipenne = O. lacki Bernhauer, 1940 syn. nov. Lectotypes are designated for O. clavatum Luze, 1906, O. japonicum and O. lacki. The habitus of O. oxyacanthae, O. scabrum and O. septentrionis, and, in addition, the aedeagus and female accessory sclerites for O. japonicum and O. strigicolle are illustrated. A checklist and key to all species of Omalium of Asian part of Russia eastwards of Ural Mts., Mongolia and Japan are provided. Three species, that were based on the misidentifications, are excluded from the fauna of Siberia: O. allardii Fairmaire & Brisout, 1859, O. excavatum Stephens, 1834 and O. foraminosum Mäklin, 1852. Distributional maps for the new species and O. curtipenne, O. diffine, O. longicorne, O. niponense and O. subsolanum are given. Several species of Omalium are recorded from a certain area for the first time: O. curtipenne from Tyumen Area, Evenkia, Chita Area, Yakutia, Magadan and Amur areas, O. longicorne and O. oxyacanthae from Amur Area, O. niponense from Russia (Maritime Province), O. rivulare from Khanti-Mansi Autonomous region, Tomsk and Novosibirsk areas, O. septentrionis from Khanty-Mansi and Taimyr autonomous regions, Evenkia, Tuva, Amur Area and Khabarovsk Territory, O. strigicolle from Khanty-Mansi Autonomous Region, Tyumen Area, Krasnoyarsk Territory, Evenkia, Tuva, Amur Area and Khabarovsk Territory, and O. subsolanum from Amur Area and Japan.
References
- Adachi, T. (1957) The staphylinid fauna of Japan. The Journal of the Tokyo University, 11, 166–200.
- Ananina, T.A. & Voincov, A.A. (2006) K poznaniyu fauny stafilinid (Coleoptera, Staphylinidae) Barguzinskogo zapovednika [To the knowledge of the fauna of rove beetles (Coleoptera, Staphylinidae) of Barguzinsky Nature Reserve]. Trudy gosudarstvennogo prirodnogo biosfernogo zapovednika “Barguzinskiy”, 9, 38–44. [in Russian]
- Anishchenko, A.V. & Shavrin, A.V. (1998) On the fauna of ground beetles (Coleoptera, Carabidae) and rove beetles (Coleoptera, Staphylinidae) of the Baikal-Lena Nature Reserve. Vestnik Irkutskoy Gosudarstvennoy Sel’skokhozyaystvennoy Akademii: IGSKhA, 13, 30–33. [in Russian]
- Averenskiy, A.I. (1999) Katalog zhukov Yakutii [The catalogue of beetles of Yakutia]. Yakutsk Scientific Centre SD RAS, Yakutsk, 76 pp. [in Russian]
- Averenskiy, A.I. (2003) Zhestkokrylye (Coleoptera) yakutskogo botanicheskogo sada [Beetles (Coleoptera) of Yakut botanical garden]. In: Isaev, A.P. (Ed.), Entomologicheskie issledovaniya v Yakutii [Entomological research in Yakutia]. IBPK SO RAN, Yakutsk, pp. 58–68. [in Russian]
- Babenko, A.S. (1982) Fauna i biotopicheskoe raspredelenie korotkonadkrylykh zhukov (Coleoptera, Staphylinidae) v yuzhnoy chasti lesnoy zony Zapadnoy Sibiri [The fauna and habitat distribution of rove beetles (Coleoptera, Staphylinidae) in southern part of the forest zone of West Siberia]. In: Zolotarenko, G.S. (Ed.), Poleznye i vrednye nasekomye Sibiri [Useful and harmful insects of Siberia]. Nauka, Novosibirsk, pp. 52–59. [in Russian]
- Belskaya, E.A. & Solodovnikov, A.Yu. (2003) Vliyanie piretroidnogo insekticida Detsisa na populyacii zhukov-stafilinid (Coleoptera, Staphylinidae) v agrocenoze yarovoy pshenitsi [The influence of the pyrethroid insecticide Detsis on populations of rove beetles (Coleoptera, Staphylinidae) in agrocenosis of spring wheat]. Agrokhimiya, 4, 56–62. [in Russian]
- Bernhauer, M. (1940) Neuheiten der paläarktischen Staphylinidenfauna (Col. Staph.). Mitteilungen der Münchner Entomologischen Gesellschaft, 30 (2), 622–642.
- Bernhauer, M. & Schubert, K. (1910) Staphylinidae I. In: Schenkling, S. (Ed.), Coleopterorum Catalogus. 5 (19). Junk, Berlin, pp. 1–86. https://doi.org/10.1007/978-94-011-9697-0_1
- Bukhkalo, S.P., Sergeeva, E.V. & Semenov, V.B. (2012) Rove-beetle fauna (Coleoptera, Staphylinidae) of the central part of the southern taiga in West Siberia, Russia. Euroasian Entomological Journal, 11 (4), 343–353. [in Russian]
- Coquerel, C. (1860) [New species]. In: Fairmaire, L. & Coquerel, C., Essai sur les Coléoptères de Barbarie. Annales de la Société Entomologique de France, Series 3, 8, 145–176, pl. 6.
- Dedyukhin, S.V., Nikitsky, N.B. & Semenov, V.B. (2005) Checklist of beetles (Insecta, Coleoptera) of Udmurtia. Euroasian Entomological Journal, 4 (4), 293–315.
- Eppelsheim, E. (1887) Neue Staphylinen vom Amur. Deutsche Entomologische Zeitschrift, 31, 419–430.
- Eppelsheim, E. (1893) Beitrag zur Staphylinen-Fauna des südwestlichen Baikal-Gebietes. Deutsche Entomologische Zeitschrift, 1893, 17–67. https://doi.org/10.1002/mmnd.48018930105
- Erichson, W.F. (1839) Die Käfer der Mark Brandenburg. 1 (2). F. H. Morin, Berlin, 385–740 pp.
- Ermakov, A.I., Belskaya, E.A., Nesterkov, A.V. & Semenov, V.B. (2017) K faune korotkonadkrylykh zhukov Srednego Urala [On the fauna of rove beetles of the Middle Ural]. Fauna Urala i Sibiri, 2, 9–18. [in Russian]
- Fairmaire, L. & Brisout, C. (1859) [New species]. In: Fairmaire, L. (Ed.), Miscellanea entomologica. Annales de la Société Entomologique de France, Series 3, 7, pp. 21–64.
- Fauvel, A. (1869) Remarques synonymiques sur les staphylinides du Catalogus Coleopterorum de MM v. Harold et Gemminger. L’Abeille. Mémoires d’Entomologie, 5, 479–494.
- Filatova, L.D. (1968) Ispolzovanie mer vklyucheniya dlya sravneniya stafilinid v raznykh mestoobitaniyakh [Using of measures of including for comparison of rove beetles in different habitats]. In: Teoretiko-graphovye metody v biogeographicheskikh issledovaniyakh. Isdatel`stvo DVO AN SSSR, Vladivostok, pp. 66–77. [in Russian]
- Gravenhorst, J.L.C. (1806) Monographia Coleopterorum Micropterorum. Henricus Dieterich, Gottingae, 236 + [12] pp. https://doi.org/10.5962/bhl.title.67769
- Hayashi, Y. (2006) A new species of Omalium from Japan (Coleoptera: Staphylinidae). The Entomological Review of Japan, 61 (1), 75–79.
- Hardy, J. (1851) Family 1. Staphylinidae, Leach. In: Hardy, J. & Bold, F. (Eds.), A catalogue of the insects of Northumberland and Durham. Part II. Transactions of the Tyneside Naturalists’ Field Club, 2, pp. 21–97.
- Heer, O. (1839) Fascicule II. Turici: Orelii, Fuesslini et Sociorum, pp. i–xii + 1–652 pp. [published in parts: 1838–1841]
- Herman, L.H. (2001a) Catalogue of the Staphylinidae (Insecta: Coleoptera). 1758 to the end of the second millennium. Bulletin of the American Museum of Natural History, 265 (1–7), i–vi + 1–4218. https://doi.org/10.1206/0003-0090.265.1.1
- Herman, L.H. (2001b) Nomenclatural changes in the Staphylinidae (Insecta: Coleoptera). Bulletin of the American Museum of Natural History, 264, 1–83. https://doi.org/10.1206/0003-0082(2003)416<0001:NCITPC>2.0.CO;2
- Ishihara, T., Miyatake, M., Hisamatsu, S., Sakai, M., Kinoshita, S. & Ito, K. (1974) The Coleoptera of the Ehime University Forest, I (Research of the insect-fauna of Komenono University Forest, 4). Bulletin of the Ehime University Forest, 11, 75–82. [in Japanese]
- Kashcheev, V.A. (1999) Novye Oxytelinae (Coleoptera, Staphylinidae) Srednei Azii i Kazakhstana. Tethys Entomological Research, 1, 141–156. [in Russian]
- Kim, T.-K. & Ahn, K.-J. (2014) Two Omaliine Species (Coleoptera: Staphylinidae) New to Korea. Animal Systematics, Evolution and Diversity, 30 (2), 151–157. https://doi.org/10.5635/ASED.2014.30.2.151
- Klimaszewski, J., Newton, A.F. & Thayer, M.K. (1996) A review of the New Zealand rove beetles (Coleoptera: Staphylinidae). New Zealand Journal of Zoology, 23, 143–160. https://doi.org/10.1080/03014223.1996.9518074
- Koch, C. (1940) Resultados científicos des los viajes entomolóticos en Espana patrocinados por Su Alteza el Príncipe Alessandro C. della Torre e Tasso. VI Congreso Internacional de Entomologia, Madrid, 1935, 369–390.
- Kolesnikova, A.A. (2000) Beetles. In: van Eerden, M.R. (Ed.), Pechora Delta. Structure and dynamics of the Pechora Delta ecosystems (1995–1999). University of Groningsen, Groningsen, pp. 127–131.
- Kryzhanovskiy, O.L., Tikhomirova, A.L. & Filatova, L.D. (1973) Stafilinidy (Coleoptera, Staphylinidae) Yuzhnogo Primoria [The rove beetles (Coleoptera, Staphylinidae) of South Primorye]. In: Ghilarov, M.S. (Ed.), Ekologiya pochvennykh bespozvonochnykh [Ecology of soil invertebrates]. Nauka, Moscow, pp. 144–173. [in Russian]
- Kurtcheva, G.F. (1977) Pochvennyie bespozvonochnyie sovetskogo Dal`nego Vostoka [Soil invertebrates of the soviet Far East]. Nauka, Moscow, 131 pp. [in Russian]
- Lobkova, L.E., Lobanova, V.I. & Semenov, V.B. (2017) Staphilinidy (Coleoptera, Staphylinidae) Kronotskogo zapovednika i sopredel`nykh territoriy Kamchatki. Dopolnenie 3 [Rove beetles (Coleoptera, Staphylinidae) of Kronotsky Nature Reserve and adjacent territories of Kamchatka. Supplement 3]. In: Tokranov, A.M. (Ed.), Trudy Kronotskogo gosudarstvennogo prirodnogo biosphernogo zapovednika, 5. Kamchatpress, Petropavlovsk-Kamchatskiy, pp. 54–82. [in Russian]
- Lobkova, L.E. & Semenov, V.B. (2014) Staphilinidy (Coleoptera, Staphylinidae) Kronotskogo zapovednika i sopredel`nykh territoriy Kamchatki. Dopolnenie 1 [Rove beetles (Coleoptera, Staphylinidae) of Kronotsky Nature Reserve and adjacent territories of Kamchatka. Supplement 1]. In: Nikonorov, A.P. (Ed.), Trudy Kronotskogo gosudarstvennogo prirodnogo biosphernogo zapovednika. Vol. 3. OOO “STP”, Voronezh, pp. 85–93. [in Russian]
- Lobkova, L.E. & Semenov, V.B. (2015) Staphilinidy (Coleoptera, Staphylinidae) Kronotskogo zapovednika i sopredel`nykh territoriy Kamchatki. Dopolnenie 2 [Rove beetles (Coleoptera, Staphylinidae) of Kronotsky Nature Reserve and adjacent territories of Kamchatka. Supplement 2]. In: Lobkov, E.G. (Ed.), Trudy Kronotskogo gosudarstvennogo prirodnogo biosphernogo zapovednika. Vol. 4. Kamchatpress, Petropavlovsk-Kamchatskiy, pp. 119–128. [in Russian]
- Luze, G. (1906) Revision der paläarktischen Arten der Staphyliniden-Genera: Xylodromus, Omalium, Phyllodrepa, Hypopycna, Dialycera, Pycnoglypta und Phloeonomus. Verhandlungen der Kaiserlich-Königlichen Zoologisch-Botanischen Gesellschaft in Wien, 56, 485–602.
- Mäklin, F.G. (1852) [new taxa]. In: Mannerheim, C.G. (Ed.), Zweiter Nachtrag zur Kaefer-Fauna der Nord-Amerikanischen Laender des Russischen Reiches. Bulletin de la Societe Imperiale des Naturalistes de Moscou, 25 (2), pp. 283–387.
- Mäklin, F.G. (1878) Diagnoser öfver nagra nya siberiska insektarter. Ofversigt af Finska Vetenskaps-Societetens Forhandlingar, 19, 15–32. [1876–1877]
- Mäklin, F.G. (1881) Coleoptera insamlade under den Nordenskiöld’ska expeditionen 1875 pa nagra öar vid Norges nordvestkust, pa Novaja Semlja och ön Waigastsch samt vid Jenissej i Sibirien. Köngliga Svenska Vetenskaps-Akademiens Handlingar, 18 (4), 1–48. [1880]
- Motschulsky, V. de. (1858) Énumération des nouvelles especes de coléopteres rapportés de ses voyages. Bulletin de la Société Imperiale des Naturalistes de Moscou, 30 (4), 490–517. [1857]
- Motschulsky, V. de. (1860) Énumération des nouvelles especes de coléopteres rapportées de ses voyages. 3-ieme article. Bulletin de la Société Imperiale des Naturalistes de Moscou, 33 (2), 539–588.
- Müller, O.F. (1776) Zoologia Danicae prodromus, seu animalium Daniae et Norvegiae indigenarum characteres, nomina, et synonyma imprimis popularium. Hallager, Hafniae, xxxii + 282 pp. https://doi.org/10.5962/bhl.title.13268
- Mulsant, E. & Rey, C. (1880) Tribu des brévipennes. Onzième famille: Omaliens. Douxième famille: Pholidiens. Annales de la Société Linnéenne de Lyon, Series 2, 27, 1–430. https://doi.org/10.3406/linly.1880.3986
- Nakane, T. (1963) Staphylinidae. In: Nakane, T., Ohbayashi, K., Nomura, S. & Kurosawa, Y. (Eds.), Iconographia Insectorum Japonicorum, Colore naturali edita. Vol. 2. Hokuryukan, Tokyo, pp. 81–100. [in Japanese]
- Nakane, T. (1971) Staphylinidae. In: Nakane, T., Ohbayashi, K., Nomura, S. & Kurosawa, Y. (Eds.), Iconographia insectorum Japonicorum, colore naturali edita, volumen II (Coleoptera). 4th Edition. Hokuryukan, Tokyo, pp. 81–100.
- Newton, A.F. (2022) StaphBase: Staphyliniformia world catalog database. Version August 2022. Staphylinoidea, Hydrophiloidea, Histeroidea (except Histeridae). In: Bánki, O., Roskov, Y. et al., Catalogue of Life Checklist. Digital resource. Available from: https://www.catalogueoflife.org/ (accessed 27 March 2025)
- Paykull, G. de (1789) Monographia Staphylinorum Sueciae. J. Edman, Upsaliae, 8 + 81 pp.
- Poppius, B. (1904a) Neue palaearktische Omaliiden. Meddelanden af Societas pro Fauna et Flora et Fennica, 29, 106–111.
- Poppius, B. (1904b) Neue paläarctische Coleopteren. Öfversigt af Finska Vetenskaps-Societetens Förhandlingar, 46 (16), 1–14. [1903–1904]
- Reitter, E. (1911) Vier neue Koleopteren-Arten aus der palaearctischen Fauna. Wiener Entomologische Zeitung, 30, 234–236.
- Roubal, J. (1930) Katalog coleopter (brouků) Slovenska a Podkarpatska, na zakladě bionomickem a zoogeografickem a spolu systematicky doplněk Ganglbauerovych “Die Kafer von Mitteleuropa” a Reitterovy “Fauna germanica”. Svazek 3. Učená Společnost Šarfaříkova v Bratislavě, Praha, 527 pp.
- Ryabukhin, A.S. (1999) A catalogue of rove beetles (Coleoptera: Staphylinidae exclusive of Aleocharinae) of the northeast of Asia. Pensoft, Sofia, 140 pp.
- Ryabukhin, A.S. (2002) Fauna and biotopic distribution of the rove beetles (Coleoptera: Staphylinidae) in forests of Magadan Oblast. Euroasian Entomological Journal, 1 (2), 197–200. [in Russian]
- Ryabukhin, A.S. (2010) Studying [sic!] the rove beetles fauna (Coleoptera, Staphylinidae) in the territory of Koryakia (Kamchatka). Vestnik SVNC DVO RAN, 3, 72–77. [in Russian]
- Ryabukhin, A.S. (2020) Materialy k faune stafilinid (Coleoptera: Staphylinidae) tundr Kamchatki (Podsemeystva Omaliinae, Proteininae, Pselaphinae, Tachyporinae i Oxytelinae) [Materials to the fauna of the rove beetles (Coleoptera: Staphylinidae) of the tundra of Kamchatka (Subfamilies Omaliinae, Proteininae, Pselaphinae, Tachyporinae and Oxytelinae)]. Vestnik Severo-Vostochnogo nauchnogo centra DVO RAN, 2, 59–68. [in Russian] https://doi.org/10.34078/1814-0998-2020-2-59-68
- Ryabukhin, A.S. (2024) Materials to the fauna of rove beetles (Coleoptera: Staphylinidae) of the north of Koryakia (Kamchatskiy Kray). Vestnik Severo-Vostochnogo nauchnogo centra DVO RAN, 1, 73–77 [in Russian, with Abstract in English]
- Sahlberg, J.R. (1871) Anteckningar till Lapplands coleopter-fauna. Notiser ur Sallskapets pro Fauna et Flora Fennica, 11, 385–440. [1870]
- Sahlberg, J.R. (1880) Bidrag till Nordvestra Sibiriens Insektfauna. Coleoptera. Insamlade under Expeditionerna till Obi och Jenissej 1876 och 1877. I. Cicindelidae, Carabidae, Dytiscidae, Hydrophilidae, Gyrinidae, Dryopidae, Georyssidae, Limnichidae, Heteroceridae, Staphylinidae och Micropepeplidae. Kungliga Svenska Vetenskaps-Akademiens Handlingar, 17 (4), 1–115.
- Saulcy, F.H.C. de (1864) Description de quatre nouvelles especes de coléopteres propres a la faune française et remarques sur quelques autres especes. Annales de la Société Entomologique de France, Series 4, 3, 653–658. [1863]
- Sazhnev, A.S. (2018) Materials to the rove beetles fauna (Coleoptera: Staphylinidae) of the Commander Islands (Kamchatka Region, Russia). Caucasian Entomological Bulletin, 14 (1), 19–23. [in Russian, with Abstract in English] https://doi.org/10.23885/1814-3326-2018-14-1-19-23
- Schawaller, W. (1990) Käfer aus Sibirien (Umgebung Novosibirsk) (Insecta: Coleoptera). Beträge zur Entomologie, 40 (1), 231–245.
- Schülke, M. & Smetana, A. (2015) Staphylinidae. In: Löbl, I. & Löbl, D. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 2. Hydrophiloidea—Staphylinoidea. Revised and Updated Edition. Brill, Leiden, pp. 304–1134.
- Scriba, W. (1867a) Beitrag zur Kenntniss der Staphylinen Unteritaliens. Berliner Entomologische Zeitschrift, 10, 376–378. [1866] https://doi.org/10.1002/mmnd.18660100408
- Scriba, W. (1867b) Fünf neue Leptusa-Arten. Coleopterologische Hefte, 1, 68–72.
- Sharp, D.S. (1874) The Staphylinidae of Japan. The Transactions of the Entomological Society of London, 1874, 1–103. https://doi.org/10.1111/j.1365-2311.1874.tb00159.x
- Sharp, D.S. (1889) The Staphylinidae of Japan. The Annals and Magazine of Natural History, Series 6, 3 (18), 463–476. https://doi.org/10.1080/00222938909460371
- Shavrin, A.V. (2001) K poznaniyu fauny zhukov-stafilinid (Coleoptera, Staphylinidae) Yuzhnogo Predbaykalya [To the knowledge of the staphylinid beetles fauna (Coleoptera, Staphylinidae) of South Cisbaikalia]. Proceedings of the Biology and Soil Department of the Irkutsk State University, 5, 80–96. [in Russian, with Abstract in English]
- Shavrin, A.V. (2007) Stafilinidy (Coleoptera, Staphylinidae) Pribaykal`skogo nacional`nogo parka [The rove beetles (Coleoptera, Staphylinidae) of Pribaikalsky National Park]. In: Trudy Pribaykal`skogo nacional`nogo parka. Vol. 2. Irkutsk State University, Irkutsk, pp. 132–148. [in Russian]
- Shavrin, A.V. (2009) Impact of Industrial Pollutions on Forest Communities of Rove Beetles (Coleoptera, Staphylinidae) in Shelekhov Raion of Irkutsk Oblast. Contemporary Problems of Ecology, 2 (1), 40–45. https://doi.org/10.1134/S199542550901007X
- Shavrin, A.V. (2010) Contribution to the knowledge of the fauna of the tribe Omaliini McLeay, 1825 (Coleoptera: Staphylinidae: Omaliinae) of the Baikal region and adjacent territories. Baltic Journal of Coleopterology, 10 (1), 27–43.
- Shavrin, A.V. (2020) New species and records of Paraphloeostiba Steel, 1960 from China and Laos, and descriptions of four new species of related genera (Coleoptera: Staphylinidae: Omaliinae: Omaliini). Zootaxa, 4890 (3), 301–329. https://doi.org/10.11646/zootaxa.4890.3.1
- Shavrin, A.V. (2023) A revision of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae) of the Himalayan region. Zootaxa, 5369 (2), 151–206. https://doi.org/10.11646/zootaxa.5369.2.1
- Shavrin, A.V. (2025) Review of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae: Omaliini) of China. Zootaxa, 5588 (4), 501–543. https://doi.org/10.11646/zootaxa.5588.4.1
- Shavrin, A.V. & Khachikov, E.A. (2023) Faunistic and diagnostic notes on some species of the genus Omalium Gravenhorst, 1802 of Caucasus, Turkey and Iran (Coleoptera: Staphylinidae: Omaliinae: Omaliini). Zootaxa, 5227 (4), 443–458. https://doi.org/10.11646/zootaxa.5227.4.3
- Shavrin, A.V. & Makarov, K.V. (2019) Contribution to the knowledge of the fauna of rove beetles of the subfamily Omaliinae MacLeay, 1825 (Coleoptera: Staphylinidae) of Kunashir Island, Kurile Islands. Russian Entomological Journal, 28 (1), 36–53. https://doi.org/10.15298/rusentj.28.1.06
- Shavrin, A.V., Shilenkov, V.G. & Veinberg, I.V. (1999) Fauna and ecology of Staphylinidae (Coleoptera) on the Norther [sic!] slope of Khamar-Daban in the Southern Cisbaikalia. Proceedings of the Biology and Soil Department of the Irkutsk State University, 1, 26–42. [in Russian, with English abstract]
- Shibata, Y. & Watanabe, Y. (1999) Staphylinidae. In: Uéno, S.-I., Kurosawa, Y. & Satô, M., The Coleoptera of Japan in color. Vol. II. 4th Edition. Hoikusha Publishing, Higashiosaka, pp. 261–321.
- Smetana, A. (1968) Ergebnisse der zoologischen Forschungen von Dr. Z. Kaszab in der Mongolei. 127. Staphylinidae III. Unterfamilien Oxytelinae und Oxyporinae (Coleoptera). Acta entomologica Bohemoslovaca, 65 (3), 226–238.
- Smetana, A. (1975) Ergebnisse der zoologischen Forschungen von Dr. Z. Kaszab in der Mongolie. 340. Staphylinidae IV. Unterfamilien Omaliinae bis Staphylininae (Coleoptera). Acta Zoologica Academiae Scientiarum Hungaricae, 21, 153–179.
- Stephens, J.F. (1834) Illustrations of British entomology; or, a synopsis of indigenous insects; containing their generic and specific distinctions; with an account of their metamorphoses, times of appearance, localities, food, and economy, as far as practicable. In: Mandibulata. Vol. 5. Baldwin and Cradock, London, pp. 305–368.
- Takahashi, K. (1988) Flight activity of insects sampled with a truck trap. Kontyȗ, Tokyo, 56 (2), 410–416.
- Thomson, C.G. (1857) Öfversigt af de arter inom Insektgruppen Stenini, som blifvit funna i Sverige. Öfversigt af Kongliga Vetenskaps-Akademiens Förhandlingar, 14 (6), 219–235.
- Thomson, C.G. (1884) Bidrag till Sveriges insectfauna. Opuscula Entomologica, 10, 1029–1040.
- Voincov, A.A. (2006) Zoogeograficheskiy analiz fauny stafilinid (Coleoptera, Staphylinidae) Selenginskogo srednegorya [Zoogeographical analysis of the fauna of staphylinid beetles (Coleoptera, Staphylinidae) of Selenga plateau]. In: Entomological studies in northern Asia. Proceedings of VII international meeting of entomologists of Siberia and Far East in framework of Siberian zoological conference, Novosibirsk, 20–24 September 2006, pp. 46–48. [in Russian]
- Voincov, A.A. (2010) To the knowledge of staphylinid fauna (Coleoptera, Staphylinidae) of the Vitim plateau. In: Entomological studies in northern Asia. Proceedings of VIII interregion meeting of entomologists of Siberia and Far East with participation of foreign scientists. Novosibirsk, 4–7 October 2010, Novosibirsk, pp. 46–48. [in Russian]
- Wankowicz, J. (1869) Notices entomologiques. Annales de la Société Entomologique de France, Series 4, 9, 411–422.
- Watanabe, Y. (1961) The staphylinid-fauna of the middle and southern Izu Islands. Journal of Agricultural Science, Tokyo, 6, 348–356.
- Watanabe, Y. (1990) A taxonomic study on the subfamily Omaliinae from Japan (Coleoptera, Staphylinidae). Memoirs of the Tokyo University of Agriculture, 31, 59–391.
- Watanabe, Y. (2013) Subfamily Omaliinae MacLeay, 1825. In: Shibata, Y. & Maruyama, M. (Eds.), Catalogue of Japanese Staphylinidae (Insecta: Coleoptera). Bulletin of the Kyushu University Museum, 11, pp. 73–80.
- Watanabe, Y. & Shibata, Y. (1972) The staphylinid-fauna of Yaku-shima Island, Japan, with descriptions of a new genus and new species. Journal of Agricultural Science of the Tokyo University of Agriculture, 17, 59–72.
- Weise, J. (1877) Japanische Staphilinidae [sic] und Pselaphidae. In: Beiträge zur Käferfauna von Japan, meist auf R. Hiller’s Sammlungen basirt. Deutsche Entomologische Zeitschrift, 21, 88–100.
- Yoshida, M., Kuroda, Y., Tanaka, K. & Sakuragi, D. (2008) Beetles of Mima Koyadaira region. Awa Society Bulletin, 54, 65–75. [in Japanese]
- Zanetti, A. (1987) Coleoptera. Staphylinidae. Omaliinae. In: Fauna d’Italia. Vol. 25. Calderini, Bologna, pp. i–xii + 1–472.
- Zanetti, A. (2002) Studies on Omalium Gravenhorst, 1802 from Turkey, Cyprus, and the Caucasus region, with notes on some European and Asian species (Coleoptera, Staphylinidae: Omaliinae). Bollettino del Museo Civivo di Storia Naturale di Verona. Botanica Zoologia, 26, 45–63.
- Zanetti, A. (2011) Unterfamilie Omaliinae. In: Assing, V. & Schülke, M. (Eds.), Freude-Harde-Lohse-Klausnitzer—Die Käfer Mitteleuropas. Band 4. Staphylinidae I. Zweite neubearbeitete Auflage. Spektrum Akademischer Verlag, Heidelberg, pp. 49–117.