Skip to main content Skip to main navigation menu Skip to site footer
Responsive image
Issue: Vol. 5383 No. 2: 11 Dec. 2023
Type: Article
Published: 2023-12-11
DOI: 10.11646/zootaxa.5383.2.5
Page range: 179-215
Abstract views: 250
PDF downloaded: 21

Alpheid shrimps of the genera Athanas Leach, 1814, Athanopsis Coutière, 1897 and Pseudathanas Bruce, 1983 of the coasts of the Arabian Peninsula (Malacostraca: Decapoda: Caridea)

Universidade Federal de Pelotas (UFPEL); Departamento de Ecologia; Zoologia e Genética; Instituto de Biologia; Campus Universitário Capão do Leão; RS; 96010-610; Brazil; Red Sea Research Center; King Abdullah University of Science and Technology (KAUST); Thuwal; Saudi Arabia
Crustacea Alpheidae Athanas Athanopsis Pseudathanas infauna commensal shrimp suction pump echiurans new records new species western Indian Ocean Saudi Arabia Oman

Abstract

Several species of alpheid shrimps belonging the genera Athanas Leach, 1814, Athanopsis Coutière, 1897 and Pseudathanas Bruce, 1983 (Decapoda: Caridea) are reported based on material collected on the Red Sea coast of Saudi Arabia and in Oman. Athanas claereboudti sp. nov. is described based on specimens from Masirah Island and Bandar Khayran east of Muscat, Oman, whereas A. iranicus Anker, Naderloo & Marin, 2010, A. minikoensis Coutière, 1903, A. parvus De Man, 1910, and a species tentatively identified as A. cf. dentirostris Anker, Jeng & Chan, 2001, are reported from various Omani localities. In the genus Athanopsis, four species are reported, including one species new to science: A. brevirostris Banner & Banner, 1981 from Saudi Arabia, A. dawa sp. nov. and A. rubricinctuta Berggren, 1991 from Masirah Island, Oman, and A. tarahomii Marin, Sheibani & Sari, 2014 from Saudi Arabia. The records of A. rubricinctuta and A. tarahomii represent second findings and significant extensions of their previously known ranges for these species. An important error in the description and illustration of the colour pattern of A. tarahomii is corrected, whereas the colour pattern of A. brevirostris is compared again with that of A. saurus Anker, 2011; the latter species is newly recorded from New Caledonia. The taxonomic identity of A. platyrhynchus Coutière, 1897, the type species of the genus from Djibouti, remains somewhat problematic. Finally, the monotypic genus Alberta Kazmi & Kazmi, 2010 established for Alberta banneri Kazmi & Kazmi, 2010 is placed in the synonymy of Pseudathanas Bruce, 1983 and Pseudathanas banneri (Kazmi & Kazmi, 2010) comb. nov. is reported from Muscat, Oman, for the first time since its original description. The generic diagnoses of Athanas, Athanopsis and Pseudathanas are emended to accommodate new or transferred species. All species reported are illustrated in colour.

 

References

  1. Ahyong, S.T. (2015) Decapod Crustacea of the Kermadec Biodiversity Expedition 2011. Bulletin of the Auckland Museum, 20, 405–442.
  2. Al-Kandari, M., Anker, A., Al-Hasheem, A., Al-Yassen, S., Sattari, Z. & De Grave, S. (2020) New records of decapod crustaceans from Kuwait (Malacostraca: Decapoda). Zootaxa, 4803 (2), 251–280. https://doi.org/10.11646/zootaxa.4803.2.2
  3. Anker, A. (2001) Évolution et taxonomie des Alpheidae (Crustacea, Decapoda). Thèse de Doctorat, Muséum National d’Histoire Naturelle, Paris, 547 + 331 pp.
  4. Anker, A. (2011a) Description of a third species in the alpheid shrimp genus Richalpheus Anker & Jeng, 2006 (Crustacea, Decapoda). Zootaxa, 3059 (1), 59–68. https://doi.org/10.11646/zootaxa.3059.1.4
  5. Anker, A. (2011b) Four new infaunal decapod crustaceans (Caridea: Alpheidae and Gebiidea: Axianassidae) from Lizard Island, Australia, one of them also occurring in Moorea, French Polynesia. Zootaxa, 2734 (1), 1–22. https://doi.org/10.11646/zootaxa.2734.1.1
  6. Anker, A. (2012) Notes on the Indo-West Pacific shrimp genus Athanopsis Coutière, 1897 (Crustacea, Decapoda, Alpheidae), with the description of a new species associated with echiurans (Annelida, Thalassematidae). Zootaxa, 3307 (1), 48–61. https://doi.org/10.11646/zootaxa.3307.1.3
  7. Anker, A. (2019) On three symbiotic species of the alpheid shrimp genus Salmoneus Holthuis, 1955 from the Indo-West Pacific, including one new to science (Malacostraca: Decapoda: Caridea). Zootaxa, 4651 (1), 51–63. https://doi.org/10.11646/zootaxa.4651.1.3
  8. Anker, A. (2022a) On two goby-associated snapping shrimps from the Red Sea, one of them new to science (Malacostraca: Decapoda: Alpheidae: Alpheus). Zootaxa, 5105 (3), 421–438. https://doi.org/10.11646/zootaxa.5105.3.5
  9. Anker, A. (2022b) Three new species of the shrimp genus Salmoneus Holthuis, 1955 from the Red Sea, with a new record of S. poupini Anker, 2010 (Malacostraca: Decapoda: Alpheidae). Zootaxa, 5138 (3), 261–282. https://doi.org/10.11646/zootaxa.5138.3.3
  10. Anker, A. & Ahyong, S.T. (2007) A rediagnosis of Athanopsis australis Banner & Banner, 1982, a rare alpheid shrimp from southern Australia, with a phylogeny of Athanopsis Coutière, 1897 and remarks on antitropical distributions in the Alpheidae (Decapoda, Caridea). Crustaceana, 80, 685–697. https://doi.org/10.1163/156854007781360612
  11. Anker, A., Ahyong, S.T., Palmer, A.R. & Noël, P.Y. (2006a) Morphological phylogeny of alpheid shrimps: parallel preadaptation and the origin of a key morphological innovation, the snapping claw. Evolution, 60, 2507–2528. https://doi.org/10.1111/j.0014-3820.2006.tb01886.x
  12. Anker, A., Al-Kandari, M. & De Grave, S. (2020) On two species of the alpheid shrimp genus Salmoneus Holthuis, 1955 from Kuwait, one of them new to science (Malacostraca: Decapoda: Caridea). Zootaxa, 4780 (1), 77–92. https://doi.org/10.11646/zootaxa.4780.1.3
  13. Anker, A. & Ashrafi, H. (2019) Salmoneus durisi sp. nov., an infaunal alpheid shrimp probably associated with callianassid ghost shrimps in the tropical Indo-West Pacific (Malacostraca: Decapoda: Caridea). Zootaxa, 4651 (1), 64–74. https://doi.org/10.11646/zootaxa.4651.1.4
  14. Anker, A. & De Grave, S. (2016) An updated and annotated checklist of marine and brackish caridean shrimps of Singapore (Crustacea, Decapoda). The Raffles Bulletin of Zoology, Supplement 34, 343–454.
  15. Anker, A. & De Grave, S. (2009) A new snapping shrimp (Crustacea, Decapoda, Alpheidae, Alpheus) from the estuarine mudflats of Kuwait. Zoologische Mededelingen, 83, 811–817.
  16. Anker, A. & Ďuriš, Z. (2022) Athanas philippei n. sp., a new alpheid shrimp associated with stomatopods in New Caledonia (Decapoda: Caridea). Zootaxa, 5094 (2), 288–300. https://doi.org/10.11646/zootaxa.5094.2.4
  17. Anker, A. & Dworschak, P.C. (2007) Description of a new species of Richalpheus Anker and Jeng, 2006 (Crustacea: Decapoda: Alpheidae) from the Red Sea. Journal of Natural History, 41, 2331–2340. https://doi.org/10.1080/10503300701579943
  18. Anker, A. & Jeng, M.S. (2006) Richalpheus palmeri, n. gen., n. sp., an infaunal alpheid shrimp from the Philippines, with redescription of Amphibetaeus jousseaumei Coutière, 1896 (Decapoda: Caridea). Journal of Crustacean Biology, 26, 379–391. https://doi.org/10.1651/S-2647.1
  19. Anker, A. & Jeng, M.S. (2007) Establishment of a new genus for Arete borradailei Coutière, 1903 and Athanas verrucosus Banner and Banner, 1960, with redefinitions of Arete Stimpson, 1860 and Athanas Leach, 1814 (Crustacea: Decapoda: Alpheidae). Zoological Studies, 46, 454–472.
  20. Anker, A., Jeng, M.S. & Chan, T.Y. (2001) Two unusual species of Alpheidae (Decapoda: Caridea), associated with upogebiid mudshrimps in the mudflats of Taiwan and Vietnam. Journal of Crustacean Biology, 21, 1049–1061. https://doi.org/10.1163/20021975-99990197
  21. Anker, A. & Komai, T. (2010) Description of a new species of Athanas Leach, 1814 (Crustacea, Decapoda, Alpheidae) from Madagascar and Japan. Zootaxa, 2680 (1), 45–54. https://doi.org/10.11646/zootaxa.2680.1.4
  22. Anker, A., Naderloo, R. & Marin, I. (2010) On a new species of the shrimp genus Athanas Leach, 1814 (Crustacea, Decapoda, Alpheidae) from Iran. Zootaxa, 2372 (1), 53–60. https://doi.org/10.11646/zootaxa.2372.1.6
  23. Anker, A., Poddoubtchenko, D. & Jeng, M.S. (2006b) Acanthanas pusillus, new genus, new species, a miniature alpheid shrimp with spiny eyes from the Philippines (Crustacea: Decapoda). Raffles Bulletin of Zoology, 54, 341–348.
  24. Ashrafi, H., Dehghani, A., Sari, A. & Naderloo, R. (2020a) An updated checklist of caridean shrimps of the Persian Gulf and Gulf of Oman. Zootaxa, 4747 (3), 521–534. https://doi.org/10.11646/zootaxa.4747.3.6
  25. Ashrafi, H., Ďuriš, Z. & Naderloo, R. (2020b) A new species and a new record of alpheid shrimps of the genus Salmoneus Holthuis, 1955 (Crustacea: Decapoda: Alpheidae) from Iran. Zootaxa, 4838 (4), 503–514. https://doi.org/10.11646/zootaxa.4838.4.4
  26. Ashrafi, H., Sari, A. & Naderloo, R. (2020c) A new sponge-dwelling species of Synalpheus Spence Bate, 1888 (Decapoda: Caridea: Alpheidae) from the Persian Gulf. Zootaxa, 4861 (3), 338–348. https://doi.org/10.11646/zootaxa.4861.3.2
  27. Ashrafi, H., Anker, A. & Ďuriš, Z. (2022) Salmoneus shojaei, a new species of symbiotic alpheid shrimp (Decapoda: Caridea) from the mangrove forests of Iran. Zootaxa, 5165 (1), 121–132. https://doi.org/10.11646/zootaxa.5165.1.7
  28. Banner, A.H. & Banner, D.M. (1983) An annotated checklist of the alpheid shrimp from the western Indian Ocean. Travaux et Documents de l’ORSTOM, 158, 1–164.
  29. Banner, A.H. & Banner, D.M. (1985) The alpheid shrimp of Indonesia, based upon J.G. de Man’s “The Decapoda of the Siboga Expedition, Part II: Family Alpheidae” (1911). Marine Research in Indonesia, 25, 1–79. https://doi.org/10.14203/mri.v25i0.431
  30. Banner, D.M. & Banner, A.H. (1973) The alpheid shrimp of Australia, part I: the lower genera. Records of the Australian Museum, 28, 291–382. https://doi.org/10.3853/j.0067-1975.28.1973.407
  31. Banner, D.M. & Banner, A.H. (1981) Annotated checklist of the alpheid shrimp of the Red Sea and Gulf of Aden. Zoologische Verhandelingen, Leiden, 190, 1–99.
  32. Banner, D.M. & Banner, A.H. (1982) The alpheid shrimp of Australia, Part III: The remaining alpheids, principally the genus Alpheus, and the family Ogyrididae. Records of the Australian Museum, 34, 1–357. https://doi.org/10.3853/j.0067-1975.34.1982.434
  33. Barnard, K.H. (1947) Descriptions of new species of South African Decapoda Crustacea, with notes on synonymy and new records. Annals and Magazine of Natural History, Series 11, 13 (102), 361–392. https://doi.org/10.1080/00222934608654562
  34. Barnard, K.H. (1950) Descriptive catalogue of South African decapod Crustacea. South African Museum Annals, 38, 1–837.
  35. Barnard, K.H. (1962) New records of marine Crustacea from the East African region. Crustaceana, 3, 239–245. https://doi.org/10.1163/156854062X00463
  36. Berggren, M. (1991) Athanopsis rubricinctuta, new species (Decapoda: Natantia: Alpheidae), a shrimp associated with an echiuroid at Inhaca Island, Moçambique. Journal of Crustacean Biology, 11, 166–178. https://doi.org/10.2307/1548555
  37. Bruce, A.J. (1983) Pseudathanas darwiniensis, new genus, new species, an alpheid shrimp from the Northern Territory, Australia. Journal of Crustacean Biology, 3 (3), 463–471. https://doi.org/10.2307/1548146
  38. Chace, F.A. Jr. (1988) The caridean shrimps (Crustacea: Decapoda) of the Albatross Philippine Expedition, 1907-1910, part 5: Family Alpheidae. Smithsonian Contributions to Zoology, 466, i–v +1–99. https://doi.org/10.5479/si.19436696.391.1
  39. Chow, L.H., De Grave, S., Anker, A., Poon, K.K.Y., Ma, K.Y, Chu, K.H, Chan, T.Y. & Tsang, L.M. (2021) Distinct suites of pre- and post-adaptations indicate independent evolutionary pathways of snapping claws in the shrimp family Alpheidae (Decapoda: Caridea). Evolution, 75 (11), 2898–2910. https://doi.org/10.1111/evo.14351
  40. Coutière, H. (1896) Note sur un nouvel Alphéidé, Betæus Jousseaumei [Crust.]. Bulletin de la Société Entomologique de France, 13, 313–317. https://doi.org/10.3406/bsef.1896.21892
  41. Coutière, H. (1897) Note sur un nouveau genre d’Alphéidés. Bulletin du Muséum d’Histoire Naturelle, 1 (3), 301–303. https://doi.org/10.5962/bhl.part.19260
  42. Coutiere, H. (1899) Les “Alpheidae”, morphologie externe et interne, formes larvaires, bionomie. Annales des Sciences Naturelles, Zoologie, Série 8, Vol. 9. Masson, Paris, 559 pp., pls. 1–6. https://doi.org/10.5962/bhl.title.13143
  43. Coutière, H. (1903) Note sur quelques Alpheidae des Maldives et Laquedives. Bulletin de la Société Philomatique de Paris, Série 9, 5, 72–90.
  44. Coutière, H. (1905) Les Alpheidae. In: Stanley Gardiner, J. (Ed.), The fauna and geography of the Maldive and Laccadive Archipelagoes, 2, pp. 852–921, pls. 70–87.
  45. De Grave, S. & Anker, A. (2000) A new genus and new species of alpheid shrimp (Crustacea: Decapoda) from Papua New Guinea. Raffles Bulletin of Zoology, 48, 249–256.
  46. De Grave, S. & Anker, A. (2008) Leptathanas powelli gen. nov., sp. nov., a new infaunal alpheid shrimp associated with upogebiid mudshrimps in Nigeria (Crustacea, Decapoda). Zootaxa, 1750 (1), 43–52. https://doi.org/10.11646/zootaxa.1750.1.4
  47. De Haan, W. (1849) [1833–1850]. Crustacea. In: von Siebold, P.F. (Ed.), Fauna Japonica sive descriptio animalium, quae in itinere per Japoniam, jussu et auspicilis superiorum, qui summum in India Batava Imperium tenent, suspecto, annis 1823–1830 collegit, notis, observationibus et adumbrationibus illustravit. Apud Auctorem, Lugduni Batavorum, i–xxxi + ix–xvi + 1–243, pls. A–J + L–Q + 1–55.
  48. De Grave, S. & Fransen, C.H.J.M. (2011) Carideorum catalogus: the recent species of the dendrobranchiate, stenopodidean, procarididean and caridean shrimps (Crustacea: Decapoda). Zoologische Mededelingen, Leiden, 85 (9), 195–589.
  49. Dehghani, A., Sari, A. & Naderloo, R. (2019) Three new species of narrowly endemic snapping shrimp, genus Alpheus (Decapoda: Caridea: Alpheidae) from the Persian Gulf. Journal of the Marine Biological Association of the United Kingdom, 99, 911–920. https://doi.org/10.1017/S0025315418000693
  50. De Man, J.G. (1910) Diagnoses of new species of macrurous decapod Crustacea from the “Siboga-Expedition”, V. Tijdschrift der Nederlandsche Dierkundige Vereeniging, 2 (11), 287–319.
  51. De Man, J.G. (1911) The Decapoda of the Siboga-Expedition, II: Family Alpheidae. Siboga-Expeditie, 39 a1 (2). E.J. Brill, Leiden, pp. 133–465, Supplement (1915), Explanations of plates of Alpheidae, pls. 1–23.
  52. Ďuriš, Z. & Anker, A. (2014) Athanas manticolus sp. nov., a new stomatopod-associated alpheid shrimp from Vietnam (Crustacea, Decapoda). Zootaxa, 3784 (5), 550–558. https://doi.org/10.11646/zootaxa.3784.5.4
  53. Fabricius, J.C. (1798) Supplementum Entomologiae Systematicae. Proft et Storch, Hafniae, 572 pp.
  54. Fourmanoir, P. (1958) Crevettes Alphéides de Nosy-Bé. Naturaliste Malgache, 10, 115–127.
  55. Goto, R. & Kato, M. (2012) Geographic mosaic of mutually exclusive dominance of obligate commensals in symbiotic communities associated with a burrowing echiuran worm. Marine Biology, 159, 319–330. https://doi.org/10.1007/s00227-011-1810-8
  56. Hayashi, K.I. (2002) A new species of the genus Athanas (Decapoda, Caridea, Alpheidae) living in the burrows of mantis shrimp. Crustaceana, 75, 395–403. https://doi.org/10.1163/156854002760095462
  57. Heller, C. (1861a) Synopsis der im Rothen Meere vorkommenden Crustaceen. Verhandlungen der Koeniglich-Kaiserlichen Zoologisch-Botanischen Gesellschaft in Wien, 11, 3–32.
  58. Heller, C. (1861b) Beiträge zur Crustaceen-Fauna des Rothen Meeres. Erster Theil. Sitzungsberichte der Mathematisch-Naturwissenschaftlichen Classe der Kaiserlichen Akademie der Wissenschaften, Wien, 44, 297–374, pls. 1–4. https://doi.org/10.5962/bhl.title.120013
  59. Heller, C. (1862) Beiträge zur Crustaceen-Fauna des Rothen Meeres. Sitzungsberichte der Mathematisch-Naturwissenschaftlichen Classe der Kaiserlichen Akademie der Wissenschaften, Wien, 44, 241–295, pls. 1–3.
  60. Holthuis, L.B. (1955) The recent genera of the caridean and stenopodidean shrimps (Class Crustacea: Order Вecapoda: Supersection Natantia) with keys to their determination. Zoologische Verhandelingen, Leiden, 26, 1–157.
  61. Holthuis, L.B. (1958) Contributions to the knowledge of the Red Sea. N°. 8. Crustacea Decapoda from the northern Red Sea (Gulf of Aqaba and Sinai Peninsula). I. Macrura. Bulletin of the Sea Fisheries Research Station Haifa, 17, 1–40.
  62. Kazmi, Q.B. & Kazmi, M.A. (2010) Biodiversity and biogeography of caridean shrimps of Pakistan. Higher Education Commission, Islamabad, xvii + 516 pp.
  63. Kensley, B.F. (1970) Some decapod Crustacea from northern South-West Africa, including a new species of Hippolyte. Cimbebasia, SWA Research, Series A, 1, 180–187.
  64. Kensley, B.F. (1972) Shrimps and prawns of southern Africa. South African Museum, Cape Town, 65 pp.
  65. Kensley, B.F. (1981) On the zoogeography of southern African decapod Crustacea, with a distributional checklist of the species. Smithsonian Contributions to Zoology, 338, 1–64. https://doi.org/10.5479/si.00810282.338
  66. Komai, T. & Henmi, Y. (2023) A new species of the alpheid shrimp genus Athanas Leach, 1814 (Decapoda: Caridea) from the Sea of Japan. Zootaxa, 5277 (2), 363–373. https://doi.org/10.11646/zootaxa.5277.2.6
  67. Leach, W.E. (1814) Crustaceology. In: Brewster, D. (Ed.), The Edinburgh Encyclopaedia, 7, pp. 383–437.
  68. Ledoyer, M. (1970) Étude systématique et remarques écologiques sur les Caridea recueillis principalement dans les biotopes de substrat meuble. Régions de Tuléar et de Nosy-Bé. Annales de l’Université de Madagascar (Sciences Naturelles et Mathématiques), 7, 121–157, pls. 1–25.
  69. Macnae, W. & Kalk, M. (1962) The fauna and flora of sand flats at Inhaca Island, Mozambique. Journal of Animal Ecology, 31, 93–128. https://doi.org/10.2307/2334
  70. Marin, I. (2017) Athanas alpheusophilus sp. nov. (Decapoda: Alpheidae)—a new Alpheus-associated shrimp from the Russian coast of the Sea of Japan. Zootaxa, 4324 (1), 50–62. https://doi.org/10.11646/zootaxa.4324.1.3
  71. Marin, I., Sheibani, R. & Sari, A. (2014) A re-description of rare alpheid shrimp Amphibetaeus jousseaumei (Coutière, 1896) and description of a new species of the genus Athanopsis Coutière, 1897 (Crustacea: Decapoda: Alpheidae) from Iranian coast of the Persian Gulf. Zootaxa, 3846 (3), 398–410. https://doi.org/10.11646/zootaxa.3846.3.4
  72. Miya, Y. (1980) Two new records of the genera, Athanopsis and Prionalpheus, from Japan, with description of a new species (Crustacea, Decapoda Alpheidae). Publications from the Amakusa Marine Biological Laboratory, Kyushu University, 5, 117–131.
  73. Miya, Y. & S. Miyake. 1968a. Revision of the genus Athanas of Japan and the Ryukyu Islands, with description of a new species (Crustacea, Decapoda, Alpheidae). Publications from the Amakusa Marine Biological Laboratory, Kyushu University, 1, 129–162.
  74. Naderloo, R. & Türkay, M. (2012) Decapod crustaceans of the littoral and shallow sublittoral Iranian coast of the Persian Gulf: faunistics, biodiversity and zoogeography. Zootaxa, 3374 (1), 1–67. https://doi.org/10.11646/zootaxa.3374.1.1
  75. Ortmann, A. (1890) Die Unterordnung Natantia Boas. Theil I. Die Decapoden Krebse des Strassburger Museums, mit besonderer Berücksichtigung der von Herrn Dr. Döderlein bei Japan und bei den Liu-Kiu-Inseln gesammelten und z. Z. im Strassburger Museum aufbewahrten Formen. Zoologische Jahrbücher, Systematik, 5, 437–542, pls. 36, 37.
  76. Paulson, O. (1875) Podophthalmata i Edriophthalmata. Issledovaniya Rakoobraznykh Krasnogo morya s zametkami otnositel’no rakoobraznykh drugikh morei, chast’ I. S.V. Kul’zhenko, Kiev, xiv + 144 pp., 21 pls.
  77. Rafinesque, C.S. (1815) Analyse de la Nature ou Tableau de l’Univers et des corps organisés, Palermo, 224 pp. https://doi.org/10.5962/bhl.title.106607
  78. Ramadan, M.M. (1936) Report on a collection of Stomatopoda and Decapoda from Ghardaga, Red Sea. Bulletin of the Faculty of Sciences, Egyptian University, 6, 1–43, pls. 1, 2.
  79. Sakai, K. & Türkay, M. (1995) Two upogebiid species from the Persian-Arabian Gulf, with a description of a related new species from Taiwan (Crustacea: Decapoda: Upogebiidae). Senckenbergiana Maritima, 25, 197–208.
  80. Spence Bate, C. (1888) Report on the Crustacea Macrura dredged by H.M.S. Challenger during the years 1873–76. The voyage of H.M.S. Challenger. Zoology, Vol. 24. Eyre & Spottiswoode, London, xi + 942 pp., 157 pls. (separate volume).
  81. Stimpson, W. (1860) Prodromus descriptionis animalium evertebratorum, quae in Expeditione ad Oceanum Pacificum Septentrionalem, a Republica Federata missa, Cadwaladaro Ringold et Johanne Rodgers Ducibus, observavit et descripsit. Pars VIII. Crustacea Macrura. Proceedings of the Academy of Natural Sciences of Philadelphia, 1860, 22–47.
  82. Tattersall, W.M. (1921) Report on the Stomatopoda and macrurous Decapoda collected by Mr. Cyril Crossland in the Sudanese Red Sea. Journal of Linnean Society of London, Zoology, 34, 345–398, pls. 27 + 28. https://doi.org/10.1111/j.1096-3642.1921.tb01838.x
  83. Titgen, R.H. (1982) The systematics and ecology of the decapods of Dubai, and their zoogeographic relationships to the Arabian Gulf and the western Indian Ocean. PhD thesis, Texas A&M University, College Station, Texas, 269 pp.