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Type: Article
Published: 2023-04-19
Page range: 67-91
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The genus Armadillo Latreille, 1802 (Oniscidea, Armadillidae) from Cyprus, with descriptions of two new species

Department of Biological Sciences; University of Cyprus; Lefkosia (Nicosia); Cyprus.
Department of Biological Sciences; University of Cyprus; Lefkosia (Nicosia); Cyprus.
Istituto di Ricerca sugli Ecosistemi Terrestri; Consiglio Nazionale delle Ricerche; Sesto Fiorentino (Florence); Italy.; Museo di Storia Naturale; Sezione di Zoologia “La Specola”; Florence; Italy.,Department of Biological Sciences; University of Cyprus; Lefkosia (Nicosia); Cyprus.
Department of Biological Sciences; University of Cyprus
Crustacea terrestrial isopods cryptic species endogean species oceanic island Mediterranean region

Abstract

The present work aims to morphologically characterize the species of Armadillo from Cyprus. Moreover, two new species of the genus are described, A. konstantinoui sp. nov. from several places along the island, and A. karametae sp. nov. from Pafos region. The circum-Mediterranean species A. officinalis is also redescribed.

 

References

  1. Agodi, A., Conti, G.O., Barchitta, M., Quattrocchi, A., Lombardo, B.M., Montesanto, G., Messina, G., Fiore, M. & Ferrante, M. (2015) Validation of Armadillo officinalis Duméril, 1816 (Crustacea, Isopoda, Oniscidea) as a bioindicator: In vivo study of air benzene exposure. Ecotoxicology and Environmental Safety, 114, 171–178. https://doi.org/10.1016/j.ecoenv.2015.01.011
  2. Anderson, L.E. (1954) Hoyer’s Solution as a Rapid Permanent Mounting Medium for Bryophytes. The Bryologist, 57, e242. https://doi.org/10.2307/3240091
  3. Arcangeli, A. (1957) Il genere Armadillo Br. emend. Arc. (Crostacei Isopodi terrestri). Bollettino dell’Istituto e Museo di Zoologia dell’Università di Torino, 5, 165–191, pls. 1–5.
  4. Barnard, K.H. (1958) Terrestrial isopods and amphipods from Madagascar. Mémoires de l‘Institute scientifique de Madagascar, Série A, 12, 67–111.
  5. Barnard, K.H. (1960a) Terrestrial Isopoda from the Transvaal. Annals of the Natal Museum, 15, 45–55. [https://hdl.handle.net/10520/AJA03040798_794]
  6. Barnard, K.H. (1960b) A new species of Calmanesia [Isopoda Terrestrial] from Madagascar. Mémoires de l’Institute scientifique de Madagascar, Série A, 14, 59–61.
  7. Barth, J.M.I., Gubili, C., Matschinaer, M., Tørresen, O.K., Watanabe, S., Egger, B., Han, Y-S., Feunteun, E., Sommaruga R., Jehle, R. & Schabetsberger, R. (2020) Stable species boundaries despite ten million years of hybridization in tropical eels. Nature Communications, 11, 1433. https://doi.org/10.1038/s41467-020-15099-x
  8. Bedek, H.E., Zimmermann, N.E., McVicar, T.R., Vergopolan, N., Berg, A. & Wood, E.F. (2018) Present and future Köppen-Geiger climate classification maps at 1-km resolution. Scientific Data, 5, e180214. https://doi.org/10.1038/sdata.2018.214
  9. Bedek, H.E., Zimmermann, N.E., McVicar, T.R., Vergopolan, N., Berg, A. & Wood, E.F. (2020) Publisher Correction: Present and future Köppen-Geiger climate classification maps at 1-km resolution. Scientific Data, 7, e274. https://doi.org/10.1038/s41597-020-00616-w
  10. Boyko, C.B., Bruce, N.L., Hadfield, K.A., Merrin, K.L., Ota, Y., Poore, G.C.B. & Taiti, S. (2008 onwards). World Marine, Freshwater and Terrestrial Isopod Crustaceans database. Armadillo Latreille, 1802. Available from: https://www.marinespecies.org/aphia.php?p=taxdetails&id=248455 (accessed 17 August 2022)
  11. Brandt, J.F. (1831) Isopoda. Gleichfüßler. In: Brandt, J.F. & Ratzeburg, J.C.T. (Eds.), Medizinische Zoologie oder getreue Darstellung und Beschreibung der Tiere die in der Arzneimittellehre in Betracht kommen, in systematischer Folge herausgegeben. Vol. 2. Isopoda. Hirschwald, Berlin, pp. 70–84, pls. 12–13. https://doi.org/10.5962/bhl.title.120464
  12. Brian, A. (1932) Spedizione scientifica all’Oasi di Cufra (Marzo-Luglio 1931). Gli isopodi terrestri. Annali del Museo civico di Storia naturale Giacomo Doria, 55, 329–352, pls. 8–10.
  13. Budde-Lund, G. (1885) Crustacea Isopoda Terrestria per Familias et Genera et Species Descripta. Copenhagen, Nielsen & Lydiche, 319 pp.
  14. Budde-Lund, G. (1904) A revision of Crustacea Isopoda terrestria, with additions and illustrations. Pt. 1 Eubelum. Pt. 2 Spherilloninae. Pt. 3 Armadillo. H. Hagerup, Copenhagen, pp. 33–144, pls. 6–10. https://doi.org/10.5962/bhl.title.9883
  15. Budde-Lund, G. (1908) Isopoda von Madagaskar und Ostafrika mit Diagnosen verwandter Arten. In: Voeltzkow, A. (Ed.) Reise in Ostafrika in den Jahren 1903–1905. Vol. 2. Wissenschaftliche Ergebnisse, Stuttgart, pp. 265–308, pls. 12–18. https://doi.org/10.5962/bhl.title.12989
  16. Budde-Lund, G. (1912) Die Fauna von Südwest-Australien. Oniscoidea (nachgelassenes Fragment, hrsg. von W. Michaelsen). In: Michaelsen, W. & Hartmeyer, R. (Ed.), Ergebnisse der Südwestaustralischen Forschungsreise 1905. Vol. 4. G. Fischer, Jena, pp. 1–4 + 17–44, 1 pl.
  17. Campos-Filho, I.S., Fernandes, C.S., Cardoso, G.M., Bichuette, M.E., Aguiar, J.O. & Taiti, S. (2019) Two new species and new records of terrestrial isopods (Crustacea, Isopoda, Oniscidea) from Brazilian caves. Zootaxa, 4564 (2), 422–448. https://doi.org/10.11646/zootaxa.4564.2.6
  18. Campos-Filho, I.S. & Taiti, S. (2021) Oniscidea taxonomy: present and future. Abstract book of the 11th International Symposium on Terrestrial Isopod Biology, “Facets of terrestrial isopod biology”. Spinicornis, Ghent, 9 pp. [https://spinicornis.be/istib2021/presentations/]
  19. Cividini, S. & Montesanto, G. (2018a) Changes in turn alternation pattern in response to substrate-borne vibrations in terrestrial isopods. Behavioural Processes, 146, 27–33. https://doi.org/10.1016/j.beproc.2017.11.005
  20. Cividini, S. & Montesanto, G. (2018b) Aggregative behavior and intraspecific communication mediated by substrate-borne vibrations in terrestrial arthropods: An exploratory study in two species of woodlice. Behavioural Processes, 157, 422–430. https://doi.org/10.1016/j.beproc.2018.07.006
  21. Collinge, W.E. (1922) Two new terrestrial isopods from Madagascar. Journal of the Linnean Society, Zoology, 35, 107–113, pl. 9. https://doi.org/10.1111/j.1096-3642.1922.tb00466.x
  22. Constantinou, G. & Panayides, I. (2013) Cyprus and Geology, Science—Environment—Culture. Bank of Cyprus Cultural Foundation, Nicosia. [unknown pagination, in Greek]
  23. Cruz, A. (1994) Redescripción de Agabiformius obtusus (Budde-Lund, 1909) y de Armadillo hirsutus (Koch 1856) (Isopoda: Oniscidea) de la Peninsula Ibérica. Butlletí de la Institució catalana d’Historia natural, 62, 65–76.
  24. Cyprus Department of Metereology (2022) Climatological Information. Available from: http://www.moa.gov.cy/moa/dm/dm.nsf/climateinfo_en/climateinfo_en?OpenDocument (accessed 27 July 2022)
  25. Dalens, H. (1990) Les Australiodillinae Vandel, 1973 (Isopoda, Oniscidea): révision des espèces connues et description d’un nouveau genre et d’une nouvelle espéce de Papouasie-Nouvelle Guinée. Bulletin de la Société d’Histoire naturelle de Toulouse, 126, 25–34.
  26. Dalens, H. (1993) Two new genera of terrestrial isopods (Crustacea: Isopoda: Oniscidea) from north-western Western Australia. Records of the Western Australian Museum, 16 (2), 257–267.
  27. Darriba, D., Taboada, G.L., Doallo, R. & Posada, D. (2012) JModelTest 2: More models, new heuristics and parallel computing. Nature Methods, 9, 772. https://doi.org/10.1038/nmeth.2109
  28. Delipetrou, P., Makhzoumi, J., Dimipoulos, P. & Georghiou, K. (2008) Cyprus. In: Vogiatzakis, I.N., Pungetti, G. & Mannion, A.M. (Ed.) Mediterranean Island Landscapes. Natural and Cultural Approaches. Springer, Dordrecht, pp. 170–203. https://doi.org/10.1007/978-1-4020-5064-0
  29. Dimitriou, A.C., Antoniou, A., Alexiou, I., Poulakakis, N., Parmakelis, A. & Sfenthourakis, S. (2022) Diversification within an oceanic Mediterranean island: Insights from a terrestrial isopod. Molecular Phylogenetics and Evolution, 175, 107585. https://doi.org/10.1016/j.ympev.2022.107585
  30. Dimitriou, A.C., Taiti S. & Sfenthourakis, S. (2019) Genetic evidence against monophyly of Oniscidea implies a need to revise scenarios for the origin of terrestrial isopods. Nature Scientific Reports, 9, e18508. https://doi.org/10.1038/s41598-019-55071-4
  31. Dollfus, A. (1892) Voyage de M. Ch. Alluaud dans le territoire d’Assinie (Afrique occidentale) en juillet et août 1886. Crustacés isopodes terrestres. Annales de la Société entomologique de France, 1892, 385–390, pl. 7.
  32. Dollfus, A. (1905) Liste des isopodes terrestres recueillis par M. le Dr. Cecconi dans l’île de Chypre. Feuille des jeunes Naturalistes, 4me Série, 35 (163), 172–177.
  33. Duméril, A. (1819) Insectes. ARM. In: Jardin du Roi and Écoles de Paris (Ed.), Dictionnaire des Sciences Naturelles, Tome Troisième. Le Normant, Paris, pp. 115–117.
  34. Fabricius, J.C. (1798) Supplementum entomologiae systematicae. CG Proft & Storch, Hafniae, pp. 296–306. https://doi.org/10.5962/bhl.title.122153
  35. Felsenstein, J. (1985) Confidence limits on phylogenies: an approach using the bootstrap. Evolution, 39 (4), 783–791. https://doi.org/10.1111/j.1558-5646.1985.tb00420.x
  36. Gao, Y.-D., Gao, X.-F. & Harris, A.J. (2019) Species boundaries and parapatric speciation in the complex of alpine shrubs, Rosa sericea (Rosaceae), based on population genetics and ecological tolerances. Frontiers in Plant Science, 10, 321. https://doi.org/10.3389/fpls.2019.00321
  37. Gruner, H. (1953) Der Rollmechanismus bei kugelnden Landisopoden und Diplopoden. Mitteilungen des zoologischen Museums Berlin, 29, 148–179.
  38. Hadjikyriakou, G. & Hadjisterkotis, E. (2002) The adventive plants of Cyprus with new records of invasive species. Zeitschrift für Jagdwissenschaft, 48 (Supplement 1), 59–71. https://doi.org/10.1007/BF02192393
  39. Hadjisterkotis, E., Masala, B. & Reese, D. (2000) The origin and extinction of the large endemic Pleistocene mammals of Cyprus. Biogeographia Anatolia, 21, 593–606. https://doi.org/10.21426/B6110069
  40. Hurtado, L.A., Lee, E.J., Mateos, M. & Taiti, S. (2014) Global diversification at the harsh sea-land interface: mitochondrial phylogeny of the supralittoral isopod genus Tylos (Tylidae, Oniscidea). PLoS ONE, 9(4), e94081. https://doi.org/10.1371/journal.pone.0094081
  41. Iwamoto, K. (1943) On Japanese terrestrial isopods. Shokobutsu oyobi dobutsu, 11, 959–974 [in Japanese].
  42. Javidkar, M., Cooper, S.J.B., King, R.A., Humphreys, W.F. & Austin, A. (2015) Molecular phylogenetic analyses reveal a new southern hemisphere oniscidean family (Crustacea: Isopoda) with a unique water transport system. Invertebrate Systematics, 29, 554–577. https://doi.org/10.1071/IS15010
  43. Jorna, J., Linde, J.B., Searle, P.C., Jackson, A.C., Nielsen, M-E., Nate, M.S., Saxton, N.A., Grewe, F., Herrera-Campos, M.A., Spjut, R.W., Wu, H., Ho, B., Lumbsch, H.T. & Leavitt, S.D. (2021) Species boundaries in the messy middle—A genome-scale validation of species delimitation in a recently diverged lineage of coastal fog desert lichen fungi. Ecology and Evolution, 11, 18615–18632. https://doi.org/10.1002/ece3.8467
  44. Khila, M., Zaabar, W., Bouslama, M.F. & Achouri, M.S. (2018) Comparison of terrestrial isopod (Crustacea: Oniscidea) assemblages from two preserved areas (Bouhedma and Chambi) in arid regions. The European Zoological Journal, 85 (1), 159–169. https://doi.org/10.1080/24750263.2018.1455902
  45. Koch, C. (1856) Crustacea. In: Rosenhauer, W.G. (Ed.), Die Thiere Andalusiens nach dem Resultate einer Reise zusammengestellt, nebst den Beschreibungen von 249 neuen oder bis jetzt noch unbeschriebenen Gattungen und Arten. T. Blaesing, Erlangen, pp. 418–423. https://doi.org/10.5962/bhl.title.66016
  46. Kozlov, A.M., Darriba, D., Flouri, T., Morel, B. & Stamatakis, A. (2019) RAxML-NG: a fast, scalable and user-friendly tool for maximum likelihood phylogenetic inference. Bioinformatics, 35 (21), 4453–4455. https://doi.org/10.1093/bioinformatics/btz305
  47. Latreille, A. (1802) Histoire naturelle, générale et particulière des crustacés et des insects. Tome III. Famille Seconde. Cloportides. L’Imprimeria de F. Dufart, Paris, pp. 42–43.
  48. Lewis, F. (1998) Oniscidea (Isopoda) from Lord Howe Island. Crustaceana, 71 (7), 743–777. [http://www.jstor.org/stable/20106052?origin=JSTOR-pdf]
  49. Lins, L.S.F., Ho, S.Y.W. & Lo, N. (2017) An evolutionary timescale for terrestrial isopods and a lack of molecular support for the monophyly of Oniscidea (Crustacea: Isopoda). Organisms, Diversity and Evolution, 17, 813–820. https://doi.org/10.1007/s13127-017-0346-2
  50. Mallet, J. (2006) Concepts of species. In: Levin, S.A. (Ed.), Encyclopedia of Biodiversity. 2nd Edition. Academic Press (Elsevier), Oxford, pp. 679–691. https://doi.org/10.1016/B978-0-12-384719-5.00131-3
  51. Marchese, C. (2015) Biodiversity hotspots: A shortcut for a more complicated concept. Global Ecology and Conservation, 3, 297–309. https://doi.org/10.1016/j.gecco.2014.12.008
  52. Mattern, D. (1999) Die Gattung Armadillo im westlichen Mittelmeergebiet (Isopoda: Oniscidea: Armadillidae). Faunistische Abhandlungen, 21, 169–185.
  53. Montesanto, G. (2015) A fast GNU method to draw accurate scientific illustrations for taxonomy. ZooKeys, 515, 191–206. https://doi.org/10.3897/zookeys.515.9459
  54. Montesanto, G. (2016) Drawing setae: a GNU way for digital scientific illustrations. Nauplius, 24, e2016017. https://doi.org/10.1590/2358-2936e2016017
  55. Montesanto, G. & Cividini, S. (2017) The moult cycle of the terrestrial isopod Armadillo officinalis Duméril, 1816 (Crustacea: Isopoda: Oniscidea). Acta Zoologica, 99 (3), 263–273. https://doi.org/10.1111/azo.12210
  56. Myers, N., Mittermeier, R.A., Mittermeier, C.G., Da Fonseca, G.A. & Kent, J. (2000) Biodiversity hotspots for conservation priorities. Nature, 403, 853–858. https://doi.org/10.1038/35002501
  57. Omer-Cooper, J. (1923) The terrestrial Isopoda of Mesopotamia and the surrounding districts. Journal of the Bombay natural Science Society, 29, 93–106. [https://biostor.org/reference/148278]
  58. Paoli, P., Ferrara, F. & Taiti, S. (2002) Morphology and evolution of the respiratory apparatus in the family Eubelidae (Crustacea, Isopoda, Oniscidea). Journal of Morphology, 253, 272–289. https://doi.org/10.1002/jmor.10008
  59. Plötner, J., Uzzell, T., Beerli, P., Cigdem, A.C., Bilgin, C., Haefeli, C., Ohst, T., Köhler, F., Schreiber, R., Guex, G.D., Litvinchuk, S.N., Westaway, R., Reyer, H.U., Pruvost, N. & Hotz, H. (2010) Genetic divergence and evolution of reproductive isolation in eastern Mediterranean water frogs. In: Glaubrecht, M. (Ed.) Evolution in action. Springer, Berlin, Heidelberg, pp. 373–403.
  60. Poulakakis, N. & Sfenthourakis, S. (2008) Molecular phylogeny and phylogeography of the Greek populations of the genus Orthometopon (Isopoda, Oniscidea) based on mitochondrial DNA sequences. Zoological Journal of the Linnean Society, 152, 707–715. https://doi.org/10.1111/j.1096-3642.2007.00378.x
  61. Queiroz, K. (2007) Species concepts and species delimitation. Systematic Biology, 56 (6), 879–886. https://doi.org/10.1080/10635150701701083
  62. Rambaut, A. & Drummond, A. J. (2007) Tracer. Version 1.5. 0. University of Edinburgh, Edinburgh. [program]
  63. Renner, S.S. (2016) A return to Linnaeus’s focus on diagnosis, not description: the use of DNA characters in the formal naming of species. Systematic Biology, 65 (6), 1085–1095. https://doi.org/10.1093/sysbio/syw032
  64. Ronquist, F., Teslenko, M., Van Der Mark, P., Ayres, D. L., Darling, A., Höhna, S., Larget, B., Liu, L., Suchard, M.A. & Huelsenbeck, J.P. (2012) MrBayes 3.2: efficient Bayesian phylogenetic inference and model choice across a large model space. Systematic biology, 61 (3), 539–542. https://doi.org/10.1093/sysbio/sys029
  65. Said, A.B., Bohli-Abderrazek, Jelassi, R. & Nasri-Ammar, K. (2020) Locomotor behaviour of three sympatric species Ligia italica, Armadillo officinalis and Chaetophiloscia elongata (Crustacea, Oniscidea) from Ghar El Melh lagoon (Northeast Tunisia). Biological Rhythm Research, 51 (3), 338–348. https://doi.org/10.1080/09291016.2018.1528696
  66. Said, A.B., Jelassi, R., Ouni, A., Habassi, A. & Nasri-Ammar, K. (2022) Behavioral responses of two sympatric species Armadillo officinalis and Chaetophiloscia elongata (Crustacea, Oniscidae) towards zinc contaminated litter. Biological Rhythm, 53 (2), 329–340. https://doi.org/10.1080/09291016.2020.1746032
  67. Santamaria, C.A. (2019) Molecular taxonomy of endemic coastal Ligia isopods from the Hawaiian Islands: re-description of L. hawaiensis and description of seven novel cryptic species. PeerJ, 7, e7531. http://doi.org/10.7717/peerj.7531
  68. Schmalfuss, H. (1978) Morphology and function of cuticular micro-scales and corresponding structures in terrestrial isopods (Crust., Isop., Oniscoidea). Zoomorphologie, 91, 263–274. https://doi.org/10.1007/BF00999815
  69. Schmalfuss, H. (1986) Die Land-Isopoden (Oniscidea) Syriens und des Libanon. Teil I. Stuttgarter Beiträge zur Naturkunde, Serie A, 391, 1–21.
  70. Schmalfuss, H. (1990) Land-Isopoden aus dem Kaukasus-Gebiet. 3. Porcellionidae, Armadillidiidae, Armadillidae. Stuttgarter Beiträge zur Naturkunde, Serie A, 444, 1–11.
  71. Schmalfuss, H. (1996) The terrestrial isopod genus Armadillo in Western Asis (Oniscidea: Armadillidae), with descriptions of five new species. Stuttgarter Beiträge zur Naturkunde, Serie A, 544, 43 S., 1–43.
  72. Schmalfuss, H. (2003) World catalog of terrestrial isopods (Isopoda: Oniscidea). Stuttgarter Beiträge zur Naturkunde, Serie A, 654, 1–341.
  73. Schmalfuss, H. & Ferrara, F. (1983) Terrestrial isopods from West Africa, Part 3: Family Armadillidae Verhoeff, 1917. Monitore zoologico italiano, Nuova Serie, Supplemento, 18, 111–157. https://doi.org/10.1080/00269786.1983.11758568
  74. Schmalfuss, H., Paragamian, K. & Sfenthourakis, S. (2004) The terrestrial isopods (Isopoda: Oniscidea) of Crete and the surrounding islands. Stuttgarter Beiträge zur Naturkunde, Serie A, 662, 1–74.
  75. Schmidt, C. (2003) Contribution to the phylogenetic system of the Crinocheta (Crustacea, Isopoda). Part 2. (Oniscoidea to Armadillidiidae). Mitteilungen aus dem Museum für Naturkunde in Berlin, Zoologische Reihe, 79, 3–179. https://doi.org/10.1002/mmnz.20030790102
  76. Schmidt, C. (2008) Phylogeny of the Terrestrial Isopoda (Oniscidea): a Review. Arthropod Systematics & Phylogeny, 66, 191–226.
  77. Schmidt, C. & Leistikow, A. (2004) Catalogue of genera of the terrestrial Isopoda (Crustacea: Isopoda: Oniscidea). Steenstrupia, 28, 1–118.
  78. Schnitzler, H.J. (1853) De Oniscineis Agri Bonnensis. Dissertatio Zoologica. Cologne, Clouthius, 28 pp.
  79. Sfenthourakis, S. & Taiti, S. (2015) Patterns of taxonomic diversity among terrestrial isopods. ZooKeys, 515, 13–25. https://doi.org/10.3897/zookeys.515.9332
  80. Simon, E. (1885) Étude sur les Crustacés terrestres et fluviatiles recueillis en Tunisie en 1883, 1884 et 1885 par M.M.A. Letourneux, M. Sedillot et Valery-Mayet. In: Exploration scientifique de la Tunisie. Crustaces. Sciences Naturelles, Paris, pp. 5–21
  81. Strouhal, H. (1968) Die Landisopoden der Insel Zypern. Annalen des naturhistorischen Museums in Wien, 72, 299–387. [http://www.jstor.org/stable/41781786]
  82. Taiti, S. (2014) New subterranean Armadillidae (Crustacea, Isopoda, Oniscidea) from Western Australia. Tropical Zoology, 27 (4), 153–165. https://doi.org/10.1080/03946975.2014.984510
  83. Taiti, S. & Cardoso, G.M. (2020) New species and records of Exalloniscus Stebbing, 1911 from southern Asia (Malacostraca, Isopoda, Oniscidea). Tropical Zoology, 33 (4), 125–158. https://doi.org/10.4081/tz.2020.83
  84. Taiti, S., Ferrara, F. & Kwon, D.H. (1992) Terrestrial Isopoda (Crustacea) from the Togian Islands, Sulawesi, Indonesia. Invertebrate Taxonomy, 6 (3), 787–842. https://doi.org/10.1071/IT9920787
  85. Taiti, S. & Montesanto, G. (2018) New species of subterranean and endogean terrestrial isopods (Crustacea, Oniscidea) from Tuscany (central Italy). Zoosystema, 40 (11), 197–226. https://doi.org/10.5252/zoosystema2018v40a11
  86. Taiti, S. Paoli, P. & Ferrara, F. (1998) Morphology, biogeography, and ecology of the family Armadillidae (Crustacea, Oniscidea) Israel Journal of Zoology, 44, 291–301. https://doi.org/10.1080/00212210.1998.10688952
  87. Vandel, A. (1955) Mission Henri Coiffait au Liban (1951). 8. Isopodes terrestres. Archives de Zoologie expérimentale et générale, Notes et Revue, 91, 455–531.
  88. Vandel, A. (1962) Isopodes terrestres (Deuxième Partie). In: Fédération Française des Sociétes de Sciences Naturelles (ed) Faune de France 66. P. Lechevalier, Paris, pp. 416–931.
  89. Vandel, A. (1965) La faune isopodique de l’île de Chypre. Bulletin du Muséum national d’Histoire naturelle, 2e Série, 36, 818–830.
  90. Verhoeff, K.W. (1919) Über augenlose Armadillidien und kritische Prüfung der Familie Armadillidiidae. 25. Isopoden-Aufsatz. Archiv für Naturgeschichte, 83A (2), 160–170.
  91. Verhoeff, K.W. (1926) Isopoda terrestria von Neu-Caledonien und den Loyalty-Inseln. In: Sarasin, F. & Roux, J. (Ed.), Nova Caledonia. Forschungen in Neu-Caledonien und auf den Loyalty-Inseln. Recherches scientifiques en Nouvelle-Calédonie et aux iles Loyalty. A. Zoologie. Vol. 4. C.W. Kreidel, Wiesbaden, pp. 243–366. [https://biostor.org/reference/98771]
  92. Vogl, C.V. (1876) Beiträge zur Kenntnis der Land-Isopoden. Verhandlungen der zoologisch-botanischen Gesellschaft Wien, 25, 501–522.
  93. Wang, J., Yang, J., Zeng, X. & Li, W. (2022) Integrative taxonomy on the rare sky‑island Ligidium species from southwest China (Isopoda, Oniscidea, Ligiidae). BMC Zoology, 7, 26. https://doi.org/10.1186/s40850-022-00120-1
  94. Warburg, M.R. (2013) Post-parturial reproduction in terrestrial isopods: a partial review. Invertebrate Reproduction & Development, 57 (1), 10–26. https://doi.org/10.1080/07924259.2011.633620
  95. WoRMS [World Register of Marine Species] (2022) Oniscidea. Available from: https://www.marinespecies.org/aphia.php?p=taxdetails&id=146505 (accessed 26 May 2022)
  96. Zachos, F.E. (2016) Species concepts in Biology. Historical development, theoretical foundations and practice relevance. Springer Nature, Cham, 227 pp.
  97. Zimmermann, B.L., Campos-Filho, I.S. & Araujo, P.B. (2018b) Integrative taxonomy reveals a new genus and new species of Philosciidae (Crustacea: Isopoda: Oniscidea) from Neotropical region. Canadian Journal of Zoology, 96, 473–485. https://doi.org/10.1139/cjz-2017-0289
  98. Zimmermann, B.L., Campos-Filho, I.S., Cardoso, G.M., Santos, S., Aguiar, J.O. & Araujo, P.B. (2018a) Two new species of Atlantoscia Ferrara & Taiti, 1981 (Isopoda: Oniscidea: Philosciidae) from southern Brazil described in the light of integrative taxonomy. Zootaxa, 4482 (3), 551–565. https://doi.org/10.11646/zootaxa.4482.3.7
  99. Zimmermann, B.L., Campos-Filho, I.S., Deprá, M. & Araujo, P.B. (2015) Taxonomy and molecular phylogeny of the Neotropical genus Atlantoscia (Oniscidea, Philosciidae): DNA barcoding and description of two new species. Zoological Journal of the Linnean Society, 174, 702–717. https://doi.org/10.1111/zoj.12256