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Published: 2023-03-10

New or little-known Neotropical Dolichopodidae (Diptera) (II): genus Hercostomus Loew excluded from the Neotropics

Graduate Program in Entomology (PPG-Ent), Instituto Nacional de Pesquisas da Amazônia (INPA).
Instituto Federal do Triângulo Mineiro, Campus Uberaba, Rua João Batista Ribeiro 4000, Distrito Industrial II, Uberaba, Minas Gerais 38064-790, Brazil.
3Coordenação de Biodiversidade (COBIO, INPA), Av. André Araújo, 2936, Petrópolis, CEP 69067-375, Manaus, Amazonas, Brazil.
Diptera Dolichopodinae Paraclius Tachytrechus Gymnopternus new combinations new species new synonym long-legged flies


The Neotropical species of Hercostomus Loew are revised and the following new combinations are proposed: Paraclius panamensis (Van Duzee, 1931) comb. nov. (Panama), P. plumitarsis (Parent, 1931) comb. nov. (Bolivia), P. problematicus (Parent, 1930) comb. nov. (Brazil and Guyana) and Tachytrechus flavimanus (Van Duzee, 1934) comb. nov. (Brazil, Ecuador and Guyana). The type of Hercostomus petulans Parent, 1939 is lost, the species cannot be confidently identified and is here treated as a nomen dubium. As a result, the genus Hercostomus is formally excluded from the Neotropical Region. Paraclius problematicus comb. nov., P. latipes (Aldrich) and Tachytrechus flavimanus comb. nov. are redescribed based on additional material and photographs of type specimens. In addition, Paraclius maculipennis (Van Duzee, 1934) is regarded as a junior synonym of P. problematicus comb. nov. A new species of Paraclius from Costa Rica, P. norrbomi sp. nov. is described in the newly proposed latipes-group, along with P. aberrans Robinson, P. latipes and P. panamensis comb. nov.



  1. Aldrich, J.M. (1896) Dolichopodidae; Phoridae. In: Williston, S.W. (Ed.), On the Diptera of St. Vincent (West Indies). Transactions of the Entomological Society of London, 1896, pp. 309–345, pl. 12, figs. 108–119, pp. 435–438.
    Bevilaqua, M. (2020) Guide to image editing and production of figures for scientific publications with an emphasis on taxonomy. Zoosystematics and Evolution, 96 (1), 139–158.
    Brooks, S.E. (2005) Systematics and phylogeny of Dolichopodinae (Diptera: Dolichopodidae). Zootaxa, 857 (1), 1–158.
    Brooks, S.E., Cumming, J.M. & Pollet, M.A.A. (2010) Revision of the Neotropical genus Cheiromyia Dyte (Diptera: Dolichopodidae). Zootaxa, 2333 (1), 41–58.
    Brooks, S.E., Cumming, J.M., Limeira-de-Oliveira, F. & Pollet, M. (2018) New species and records of Cheiromyia Dyte from Brazil and French Guiana (Diptera: Dolichopodidae). Zootaxa, 4444 (3), 267–282.
    Capellari, R.S. & Amorim, D.S. (2009) Four new species of Paraclius Loew (Diptera: Dolichopodidae) from Brazilian Atlantic Forest, and notes on the systematic position of Cheiromyia Dyte. Zootaxa, 2274 (1), 51–61.
    Capellari, R.S. (2013) A remarkable new species of Paraclius Loew from Brazil (Diptera: Dolichopodidae). Zootaxa, 3681 (3), 293–296.
    Capellari, R.S. & Bickel, D.J. (2020) New or little-known Neotropical Dolichopodidae (Diptera) (I): the enigmatic genus Notobothrus Parent. Journal of Insect Biodiversity, 15 (1), 16–21.
    Chandler, P.J. (Ed.) (1998) Checklists of Insects of the British Isles (New Series). Part 1. Diptera. Handbooks for the Identification of British Insects, 12 (1), 1–234.
    Coquillett, D.W. (1910) The type-species of the North American genera of Diptera. Proceedings of the United States National Museum, 37, 499–647.
    Cumming, J.M. & Wood, D.M. (2017) Adult morphology and terminology. In: Kirk-Spriggs, A.H. & Sinclair, B.J. (Eds.), Manual of Afrotropical Diptera. Vol. 1. Introductory chapters and keys to Diptera families. Suricata 4. South African National Biodiversity Institute, Pretoria, pp. 89–113.
    Haliday, A.H. (1851) Family XXI. Dolichopidae. In: Walker, F., Insecta Britannica, Diptera. Vol. I. Reeve & Benham, London, pp. 144–221.
    Latreille, P.A. (1809) Genera crustaceorum et insectorum secundum ordinem naturalem in famílias disposita, iconibus exemplisque plurimis explicata. Vol. 4. Parisiis et Argentorat [Paris and Strasbourg], 399 pp.
    Loew, H. (1861) Neue Beiträge zur Kenntniss der Dipteren. Achter Beitrag. Mittler & Sohn, Berlin, 100 pp.
    Loew, H. (1864) Monographs of the Diptera of North America. Part II. Smithsonian Miscellaneous Collections, 6 (2 [= pub. 171]), 1–360, 5 pls.
    Parent, O. (1928) Etude sur les Diptères Dolichopodides exotiques conservés au Zoologisches Staatsinstitut und Zoologisches Museum de Hambourg. Mitteilungen aus dem Zoologischen Staatsinstitut und Zoologischen Museum in Hamburg, 43, 155–198.
    Parent, O. (1930) Ergebnisse einer zoologischen Sammelreise nach Brasilien, insbesondere in das Amazonasgebiet, ausgerführt von Dr. H. Zerny III. Teil. Diptera: Dolichopodidae. Annalen des Naturhistorischen Museums in Wien, 44, 5–26.
    Parent, O. (1931) Diptères dolichopodides de l’Amérique du Sud. Espèces nouvelles figurant dans la collection Schnuse conservées aux Staatliche Museen für Tierkunde und Völkerkunde zu Dresden. Abhandlungen und Berichte Dresdener Staatliches Museen für Tierkunde und Völkerkunde, 18, 1–21, 3 pls.
    Parent, O. (1939) Quelques Diptères Dolichopodides de la region neotropicale. Arbeiten über morphologische und taxonomische Entomologie, 6, 140–151.
    Pollet, M. (1990) Phenetic and ecological relationships between species of the subgenus Hercostomus (Gymnopternus) in western Europe with the description of two new species (Diptera: Dolichopodidae). Systematic Entomology, 15, 359–382.
    Pollet, M. (2004) A critical note on the systematic position of Gymnopternus (Diptera: Dolichopodidae). Studia Dipterologica, 10, 537–548.
    Robinson, H. (1964) A synopsis of the Dolichopodidae (Diptera) of the southeastern United States and adjacent regions. Miscellaneous Publications of the Entomological Society of America, 4, 103–192.
    Robinson, H. (1970) 40. Family Dolichopodidae (Dolichopidae). In: Papavero, N. (Ed.), A Catalogue of the Diptera of the Americas, south of the United States. Vol. 40. Museu de Zoologia, Universidade de São Paulo, São Paulo, pp. 1–92.
    Robinson, H. (1975) Bredin-Archbold-Smithsonian biological survey of Dominica. The family Dolichopodidae with some related Antillean and Panamanian species. Smithsonian Contributions to Zoology, 185, 1–141.
    Rohlfein, K. & Ewald, B. (1979) Katalog der in den Sammlungen der Abteilung Taxonomie der Insekten des Instituts für Pflanzenschutzforschung, Bereich Eberswalde (ehemals Deutsches Entomologisches Institut), aufbewahrten Typen-XVIII. (Diptera: Brachycera). Beiträge zur Entomologie, 29, 201–247.
    Rondani, C. (1856) Dipterologiae Italicae prodromus. Vol: I. Genera Italica ordinis dipterorum ordinatim disposita et distincta et in familias et stirpes aggregata. A. Stocchi, Parmae [Parma], 226 + [2] pp.
    Shorthouse, D.P. (2010) SimpleMappr, an online tool to produce publication-quality point maps. Available from: (accessed 20 September 2022)
    Soares, M.M.M., Capellari, R.S. & Ale-Rocha, R. (2023) Species of Paraclius Loew (Diptera: Dolichopodidae) with bi-lamellate antennal stylus: new combination, new synonym, and two new species. Zootaxa, 5231 (1), 37–51.
    Stannius, F.H. (1831) Die europäischen Arten der Zweyflüglergattung Dolichopus. Isis, Oken’s, 1831, 28–68.
    Van Duzee, M.C. (1931) Dolichopidae of the Canal Zone. Bulletin of the American Museum of Natural History, 61, 161–205.
    Van Duzee, M.C. (1934) Dolichopidae. In: Curran, C.H. (Ed.), The Diptera of Kartabo, Bartica District, British Guiana. Bulletin of the American Museum of Natural History, 66 (III), pp. 365–374 + 524.
    Yang, D., Zhu, Y., Wang, M. & Zhang, L. (2006) World Catalog of Dolichopodidae (Insecta: Diptera). China Agricultural University Press, Beijing, 704 pp.