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Published: 2023-02-08

The woodwasp genus Tremex (Hymenoptera, Siricidae) of Japan

Department of Zoology, National Museum of Nature and Science, 4–1–1 Amakubo, Tsukuba, Ibaraki 305–0005, Japan.
Symphyta new species new synonym new combination new distribution record new host record


The woodwasp genus Tremex Jurine, 1807, is revised for Japan and eight species are recognized, T. apicalis Matsumura, 1912, T. bicinctus Shinohara, n. sp., T. contractus Maa, 1949, T. fuscicornis (Fabricius, 1787), T. katayamai Shinohara & Kurihara, n. sp., T. longicollis Konow, 1896, T. nakanei Takeuchi, 1955, and T. okinawensis Togashi, 1997. Tremex bicinctus is described from Honshu and Kyushu and T. katayamai is described from Honshu, both based only on females. Tremex kaedei Togashi, 1997, is a new synonym of T. nakanei Takeuchi, 1955, and T. kurokivorus Togashi, 2006, is a new synonym of T. okinawensis Togashi, 1997. Tremex sudajii Togashi, 2006, is transferred to the genus Eriotremex Benson, 1943. Tremex okinawensis is newly recorded from Amami-ôshima Island and the male of this species is diagnosed for the first time. Pterocarya rhoifolia (Juglandaceae) is newly recorded as a host plant of T. nakanei. Keys are given for the females of eight Japanese species and for the males of six Japanese species. The stability and taxonomic use of some characters, mostly employed in previous works, are examined and discussed. Collection records are given for 443 specimens of eight species examined.


  1. Abe, M. & Togashi, I. (1989) Siricidae. In: Hirashima, Y. (Ed.), A Check List of Japanese Insects. Entomological Laboratory, Faculty of Agriculture, Kyushu University, Fukuoka, pp. 558–559. [in Japanese]

  2. Anonymous (2004) Tremex contractus Maa. In: [Endangered Wildlife in Hiroshima Prefecture, Revised Edition, Red Data Book Hiroshima 2003]. Hiroshima Prefectural Government, Hiroshima, pp. 169. [in Japanese]

  3. Anonymous (2013) [A Tremex species (Toyama-shi Einou Support Center/Toyama-shi, Tsukioka-cho)]. In: [Yamaboshi, A record book of nature]. Available from: (accessed 25 February 2022) [in Japanese]

  4. Anonymous (2014) 2014.05.16 Friday. [Ovipositing Tremex longicollis, (Erratum) Tremex contractus]. In: Insects of Tsushima Is. Available from: (accessed 25 February 2022) [in Japanese]

  5. Benson, R.B. (1943) Studies in Siricidae, especially of Europe and southern Asia (Hymenoptera; Symphyta). Bulletin of Entomological Research, 34 (1), 27–51.

  6. Benson, R.B. (1954) Some sawflies of the European Alps and the Mediterranean region (Hymenoptera: Symphyta). Bulletin of the British Museum (Natural History), Entomology Series, 3 (7), 267–295.

  7. Eichhorn, O. (1982) Familienreihe Siricoidea. In: Schwenke, W. (Ed.), Die Forstschädlinge Europas. Bd. 4. Hautflügler und Zweiflügler. Paul Parey, Hamburg und Berlin, pp. 196–231.

  8. Esaki, T., Hori, H. & Yasumatsu, K. (1939) Insectorum Japonicorum Illustratio Iconographica Coloribus Ad Naturam Depicta. Sanseido, Tokyo, 485 pp. [in Japanese]

  9. Fabricius, J.C. (1787) Mantissa Insectorum sistens eorum species nuper detectas adiectis characteribus genericis differentiis specificis emendationibus observationibus. Vol. 1. C.G. Proft, Hafniae, 348 pp.

  10. Funamoto, D. (2010) [Records of two Tremex woodwasps collected in Kanagawa Prefecture]. Kanagawa-Chûhô, 171, 10. [in Japanese]

  11. Gussakovskij, V.V. (1935) Insectes Hyménoptères, Chalastogastra 1, Fauna SSSR. 2 (1). Izdatel’stvo Akademii Nauk SSSR, Moskva, Leningrad, 453 pp. [in Russian and German]

  12. Haneda, Y., Tano, T., Okuno, H., Nozaka, C., Murota, T., Kurokawa, H. & Inoue, S. (1998) Hymenoptera. In: Fukui-ken Shizen-kankyo Hozen-chosa Kenkyu-kai Konchu Bukai (Ed.), [Catalogue of Insects of Fukui Prefecture, 2nd Ed]. Fukui-ken, Kenminseikatsu-bu, Fukui, pp. 314–404. [in Japanese]

  13. Hara, H., Ibuki, S. & Shinohara, A. (2021) Taxonomic notes and new distribution and host plant records for sawflies and woodwasps (Hymenoptera, Symphyta) of Japan VI. Bulletin of the National Museum of Nature and Science, Series A, 47, 163–188.

  14. Háva, J. (2021) The woodwasp Tremex incognitus sp. nov. (Hymenoptera: Siricidae) from Vietnam. Munis Entomology & Zoology, 16 (1), 111–114.

  15. Higurashi, T. (2019) [Collection of Tremex longicollis in Yachimata City]. Bôsô-no-konchû, 64, 93. [in Japanese]

  16. Jurine, L. (1807) Nouvelle Méthode de classer les Hyménoptères et les Diptères, Hyménoptères, Tome Premier. J.J. Paschoud, Genève, 319 pp., 14 pls.

  17. Kawamoto, Y. (2010) [A Tremex sp. new to Japan collected in Yamaguchi Prefecture]. Insects of Yamaguchi, 9, 168. [in Japanese]

  18. Kawano, T. (2014) The first record of Tremex okinawensis Togashi since the original description (Siricidae: Tremecinae). Tsunekibachi, Fukui, 24, 19–20. [in Japanese]

  19. Kim, C.-W. (1963) Hymenoptera of Korea. Humanities and Sciences, Natural Science, Korea University, 6, 243–374. [in Korean]

  20. Kim, C.-W. (1971) Tremex apicalis Matsumura as a host of an Ichneumon-fly, Megarhyssa jezoensis Matsumura. Korean Journal of Zoology, 14, 85–86. [in Korean with English summary]

  21. Kojima, K., Watanabe, H. & Nakamura, S. (1962) Hood[sic!]-plants of Siricidae and Xiphydriidae in Japan. Miscellaneous reports of Hiwa Museum for Natural History, 5, 8–15. [in Japanese with English summary]

  22. Kondo, T. (1961) [A record of Tremex longicollis with a fragmentary note on its biology]. Suzumushi, Kurashiki, 11 (2), 4–5. [in Japanese]

  23. Kondo, T. & Miyake, M. (1974) [A fragmentary note on the biology of Tremex longicollis]. Nature and Insects, Tokyo, 9 (14), 15–18. [in Japanese]

  24. Kondo, T. & Miyake, M. (1976) [Symphyta (Hymenoptera) of Okayama Prefecture, 2]. Suzumushi, Kurashiki, 113, 1–13. [in Japanese]

  25. Konishi, T. (2020) [Records of Tremex nakanei from Kyoto Prefecture]. Gekkan-Mushi, 594, 51. [in Japanese]

  26. Kono, H. & Sugihara, Y. (1939) A list of the food-plants of Siricidae and Xiphydriidae. Insecta Matsumurana, 13, 108–109.

  27. Konow, F.W. (1896) Verschiedenes aus der Hymenopteren-Gruppe der Tenthrediniden. Wiener Entomologische Zeitung, 15, 41–59.

  28. Konow, F.W. (1899) Neuer Beitrag zur Synonymie der Chalastogastra. Entomologische Nachrichten, 25 (6), 86–96.

  29. Kuramitsu, K., Ishihara, T., Sugita, A., Yooboon, T., Lustig, B., Matsumori, Y., Yamada, H. & Kinoshita, N. (2019) The attraction of Tremex apicalis (Hymenoptera, Siricidae, Tremecinae) and its parasitoid Ibalia japonica (Hymenoptera, Ibaliidae) to the fungus Cerrena unicolor. Journal of Hymenoptera Research, 68, 37–48.

  30. Kuramitsu, K., Kosaki, A., Ishihara, T., Yamada, H., Watanabe, K. (2016) Infestation of the woodwasp Tremex apicalis Matsumura (Hymenoptera, Siricidae) on the large-leaf dogwood Swida Macrophylla (Wall.) with biological notes on its parasitoid wasps. Journal of Hymenoptera Research, 52, 71–79.

  31. Kurosawa, Y. (1965) [Tremex nakanei collected in Ueno Park]. Natural Science and Museums, 32, 18. [in Japanese]

  32. Lee, J.-W., Choi, J.-K. & Park, B. (2019) Synoptic list of Symphyta (Hymenoptera) in Korea. Journal of Species Research, 8, 1–96.

  33. Lee, J.-W., Ryu, S.-M. & Chung, D.-K. (1998) Taxonomy of the genus Tremex Jurine (Hymenoptera: Siricidae) from Korea, with descriptions of one new species and one unrecorded species. Korean Journal of Biological Sciences, 2, 297–302.

  34. Maa, T.-C. (1949) A synopsis of Asiatic Siricoidea with notes on certain exotic and fossil forms. Notes d’Entomologie Chinoise, 13, 11–189.

  35. Marlatt, C.L. (1898) Japanese Hymenoptera of the family Tenthredinidae. Proceedings of the United States National Museum, 21 (1157), 493–506.

  36. Matsumoto, R., Yoshida, H. & Yashiro, M. (2009) [A Zukan (fully illustrated handbook) of the Hymenoptera not much treated in the previously published Zukans. 1. Tremex spp.]. Hachi-Kariudo, 1, 27. [in Japanese]

  37. Matsumura, S. (1906) Nihon Gaichu Mokuroku, Zen. Rokumeikan, Tokyo, 154 pp. [in Japanese]

  38. Matsumura, S. (1912) Thousand Insects of Japan. Supplement IV. Keiseisha, Tokyo, 1 + 247 pp., 14 pls., 2 +2 pp. [in Japanese and English]

  39. Matsumura, S. (1930) The Illustrated Thousand Insects of Japan, Vol. II (Hymenoptera). Tôkôshoin, Tokyo, 198 pp., 18 pls., 89 + 8 pp. [in Japanese and English]

  40. Matsumura, S. (1931) 6000 Illustrated Insects of Japan-Empire. Tôkôshoin, Tokyo, 1497 + 23 pp. [in Japanese]

  41. Matsumura, S. (1932a) Illustrated Common Insects of Japan. Hymenoptera, Vol. 4. Hymenoptera, Diptera, Rhynchota. Shûnyôdo, Tokyo, 23 pls., 99 + 154 pp. [in Japanese and English]

  42. Matsumura, S. (1932b) Conspectus of Japanese Injurious Insects. Meijitosho, Tokyo, 62 pls., 3 + 51 + 971 + 115 pp. [in Japanese]

  43. Miyatake, M. (1950) A contribution to the Siricoidea of Shikoku (Hymenoptera). Transactions of the Shikoku Entomological Society, Matsuyama, 1, 38–39.

  44. Miyoshi, K. (1988) Hymenoptera. In: The Insects of Yamaguchi Prefecture. Yamaguchi Museum, Yamaguchi, pp. 46 + 187–197. [in Japanese]

  45. Murota, T. & Kurokawa, H. (1985) Symphyta. In: [Catalogue of Insects of Fukui Prefecture]. Fukui Prefectural Government, Fukui, pp. 246–257. [in Japanese]

  46. Nagase, H. (2004) Hymenoptera (excl. Formicidae). In: Insect Fauna of Kanagawa. Kanagawa Konchû Danwakai, Odawara, pp. 1241–1326. [in Japanese]

  47. Nagase, H. (2007) Hymenoptera. In: Research Group of the Tanzawa Mountains (Ed.), Results of the Scientific Research on the Tanzawa Mountains, Appendix: A List of Plants and Animals. Hiraoka Environmental Science Laboratory, Sagamihara, pp. 286–310. [in Japanese]

  48. Nagase, H. (2008a) [Insect Fauna of Kanagawa (Hymenoptera), supplement 2]. Kanagawa-Chûhô, 161, 1–10. [in Japanese]

  49. Nagase, H. (2008b) [Insect Fauna of Kanagawa (Hymenoptera), supplement 3]. Kanagawa-Chûhô, 164, 41–54. [in Japanese]

  50. Nagase, H. & Watanabe, K. (2018) Hymenoptera. In: Insect Fauna of Kanagawa. III. Kanagawa Entomologists’ Association, Odawara, pp. 934–1038. [in Japanese]

  51. Naito, T. (2019) Family Siricidae. In: Editorial Committee of Catalogue of the Insects of Japan (Ed.), Catalogue of the Insects of Japan, Volume 9 Hymenoptera. Part 1. Symphyta. Entomological Society of Japan, Kyoto, pp. 18–20. [in Japanese]

  52. Naito, T. (2020) Family Siricidae. In: Naito, T., Shinohara, A., Hara, H. & Ito, F., Sawflies and Woodwasps of Japan. Hokkaido University Press, Sapporo, pp. 179–182 + 462–468. [in Japanese]

  53. Naito, T., Yoshida, H., Nakamine, H., Morita, T., Ikeda, T., Suzuki, H. & Nakanishi, A. (2004) Species diversity of sawflies in Hyogo Prefecture, central Japan. Museum of Nature and Human Activities, Hyogo, Monograph of Natural History and Environmental Science, 1, 1–2, pls. 1–10 + 1–85. [in Japanese]

  54. Nakahama, N., Tsuji, H., Morimoto, K. & Matsumoto, R. (2006) [Collection records and ecological notes on Tremex contractus in Osaka and Hyogo]. Nature Study, Osaka, 53, 5–6. [in Japanese]

  55. Nakamura, K. (2003) [Hymenoptera (excl. Formicidae)]. In: Tochigi-ken Shizen-kankyô Chôsa Kenkyû-Kai Konchû Bukai (Ed.), [Insects of Tochigi I, Basic Survey of Natural Environment in Tochigi Prefecture]. Tochigi-ken, Rinmu-bu, Utsunomiya, pp. 249–336. [in Japanese]

  56. Nakamura, S. & Fujishita, A. (1974) [Woodwasps of Hiroshima Prefecture]. Hiroshima Mushi-no-kai Kaiho, 13, 103–105. [in Japanese]

  57. Narita, K. (2018) [A record of a tremecine woodwasp Tremex sudajii Togashi from the mainland of Kyushu]. Gekkan Mushi, 568, 54–55. [in Japanese]

  58. Ogata, K. (2021) [Tremex longicollis ovipositing on Podocarpus macrophyllus f. angustifolius]. Satsuma, 167, 186. [in Japanese]

  59. Ogawa, K. & Shinohara, A. (2022) A new record of Tremex nakanei Takeuchi, 1955 (Hymenoptera, Siricidae) from Kyushu, Japan. Japanese Journal of Entomology, New Series, 25, 189–190.

  60. Okutani, T. (1959) [Symphyta]. In: Esaki, T., Ishii, T., Kawada, A., Shiraki T. & Yuasa, H. (Eds.), Illustrated Insect Larvae of Japan. Hokuryûkan, Tokyo, pp. 548–582. [in Japanese]

  61. Okutani, T. (1962) [On the larvae of Siricidae]. Transactions of the 72nd Meeting of the Japanese Forestry Society, 1962, 342–343. [in Japanese]

  62. Okutani, T. (1963) Description of the larvae of horntails. Science Reports of the Hyogo University of Agriculture, Series Agricultural Biology, 6 (1), 23–27.

  63. Okutani, T. (1965) [Symphyta]. In: Furukawa, H., Hasegawa, H. & Okutani, T. (Eds.), Genshoku Konchu Hyakka Zukan. Shûeisha, Tokyo, pp. 159–167 + 460–477. [in Japanese]

  64. Okutani, T. (1967) Food plants of Japanese Symphyta (I). Japanese Journal of Applied Entomology and Zoology, 11, 43–49. [in Japanese with English abstract]

  65. Okutani, T. (1970) Food-plants of Japanese Symphyta (III). Japanese Journal of Applied Entomology and Zoology, 14, 25–28. [in Japanese with English abstract]

  66. Okutani, T. (1971) [Symphyta of Japan II]. Nature and Insects, Tokyo, 6 (12), 17–19. [in Japanese]

  67. Okutani, T. (1973) Symphyta of Niigata Pref. with systematic notes on some species. Transactions of Essa Entomological Society, 42, 13–27. [in Japanese and English]

  68. Okutani, T. (1977) Siricidae. In: Ito, S., Okutani, T. & Hiura, I. (Eds.), Colored Illustrations of the Insects of Japan. Vol. II. Hoikusha, Osaka, pp. 285–286, pl. 54. [in Japanese]

  69. Palma, M.A., Valenzuela, E., Parra, P., Gutierrez, M. & Silva, L.T. (2005) Cerrena unicolor (Bull.) Murr (Basidiomycota) aislado de micangio de Tremex fuscicornis Fabr. (Hymenoptera Siricidae) asociado a decaimiento y pudrición del alamo (Populus sp.) en Chile. Boletín Micológico, 20, 57–61.

  70. Pažoutovaì, S. & Šrůtka, P. (2007) Symbiotic relationship between Cerrena unicolor and the horntail Tremex fuscicornis recorded in the Czech Republic. Czech Mycology, 59, 83–90.

  71. Saito, Y. (2013) [Tremex nakanei in Chiba Prefecture]. Bôsô-no-konchû, 51, 60. [in Japanese]

  72. Saito, Y. (2018) [Male of Tremex nakanei collected in Noda City]. Bôsô-no-konchû, 63, 61. [in Japanese]

  73. Saito, Y. & Higurashi, T. (2017) [Male of Tremex nakanei collected in Yachimata City]. Bôsô-no-konchû, (60), 53. [in Japanese]

  74. Saito, Y. & Shirota, C. (2016) [Tremex sp., still unrecorded in Chiba Prefecture, collected in Sakae Town, Inba District]. Bôsô-no-konchû, 58, 78. [in Japanese]

  75. Sakamoto, M. (1997) The siricid woodwasps (Hymenoptera, Symphyta, Siricidae) of Hiroshima Prefecture. Journal of the Hiba Society of Natural History, 177, 23–34. [in Japanese with English abstract]

  76. Schiff, N.M., Goulet, H., Smith, D.R., Boudreault, C., Wilson, A.D. & Scheffler, B.E. (2012) Siricidae (Hymenoptera: Symphyta: Siricoidea) of the Western Hemisphere. Canadian Journal of Arthropod Identification, Guelph, 21, 1–305.

  77. Semenov-Tian-Shanskij, A. (1921) Praecursoriae Siricidarum novorum diagnoses (Hymenoptera). Russkoe Entomologicheskoe Obozrenie, Petrograd, 17, 81–95.

  78. Shimoyama, K. (1986) Siricoidea (Hymenoptera) of Aomori Prefecture. Journal of Aomori-ken Biological Society, 23, 7–8. [in Japanese]

  79. Shinohara, A. (2000) Sawflies and woodwasps of the Imperial Palace, Tokyo. Memoirs of the National Science Museum, Tokyo, 36, 295–305. [in Japanese with English summary]

  80. Shinohara, A. (2001) Sawflies and woodwasps in the garden of the Institute for Nature Study. Report of the Institute for Nature Study, Tokyo, 33, 281–288. [in Japanese with English summary]

  81. Shinohara, A. (2005) Sawflies and woodwasps of the Akasaka Imperial Gardens and the Tokiwamatsu Imperial Villa, Tokyo. Memoirs of the National Science Museum, Tokyo, 39, 225–238. [in Japanese with English summary]

  82. Shinohara, A. (2014) Sawflies and woodwasps of the Imperial Palace, Tokyo. Memoirs of the National Museum of Nature and Science, Tokyo, 50, 461–475. [in Japanese with English abstract]

  83. Shinohara, A. & Hara, H. (2020) Taxonomic notes and new distribution and host plant records for sawflies and woodwasps (Hymenoptera, Symphyta) of Japan V. Bulletin of the National Museum of Nature and Science, Series A, 46, 183–202.

  84. Shinohara, A., Hara, H. & Smith, D.R. (2020) The Xiphydria annulitibia group in northeastern Asia (Hymenoptera, Xiphydriidae). Zootaxa, 4755 (2), 375–389.

  85. Shirota, C. (2017) [Tremex nakanei collected in Noda City]. Bôsô-no-konchû, 60, 52. [in Japanese]

  86. Smith, D.R. (1978) Suborder Symphyta (Xyelidae, Pararchexyelidae, Parapamphiliidae, Xyelydidae, Karatavitidae, Gigasiricidae, Sepulcidae, Pseudosiricidae, Anaxyelidae, Siricidae, Xiphydriidae, Paroryssidae, Xyelotomidae, Blasticotomidae, Pergidae). In: Vecht, J. & Shenefelt, R.D. (Eds.), Hymenopterorum Catalogues Part 14. Dr. W. Junk B.V. Publishers, Hague, pp. 43–128.

  87. Smith, D.R. (2010) The woodwasp genus Eriotremex (Hymenoptera: Siricidae), a review and a new species from Malaysia. Proceedings of the Entomological Society of Washington, 112 (3), 423–438.

  88. Sugimoto, H. & Sono, S. (2013) [Tremex sp. new to Japan collected again]. Insects of Yamaguchi, 12, 114. [in Japanese]

  89. Sundukov, Yu. N. (2015) To the sawfly fauna (Hymenoptera, Symphyta) of southern Kuriles. A.I. Kurentsov’s Annual Memorial Meetings, 26, 241–258. [in Russian with English abstract]

  90. Sundukov, Yu. N. (2017) Suborder Symphyta—Sawflies and wood wasps. In: Lelej, A.S., Proshchalykin, M. Yu. & Loktionov, V.M. (Eds.), Annotated catalogue of the Hymenoptera of Russia. Vol. I. Symphyta and Apocrita: Aculeata. Proceedings of the Zoological Institute of the Russian Academy of Sciences, Supplement 6, pp. 20–117.

  91. Sundukov, Yu. N. & Lelej, A.S. (2012) Podotryad Symphyta—Sidyachebryukhie. In: Lelej, A.S. (Ed.), Annotated Catalogue of the Insects of Russian Far East. Vol. I. Hymenoptera. Dal’nauka, Vladivostok, pp. 62–119. [in Russian]

  92. Tabata, M. & Abe, Y. (1995) Cerrena unicolor isolated from the mycangia of a horntail, Tremex longicollis, in Kochi Prefecture, Japan. Mycoscience, 36, 447–450.

  93. Taeger, A., Blank, S.M. & Liston, A.D. (2006) European Sawflies (Hymenoptera: Symphyta)—A Species Checklist for the Countries. In: Blank, S.M., Schmidt, S. & Taeger, A. (Eds.), Recent Sawfly Research — Synthesis and Prospects. Goecke &Evers, Keltern, pp. 399–504.

  94. Taeger, A., Blank, S.M. & Liston, A.D. (2010) World catalog of Symphyta (Hymenoptera). Zootaxa, 2580 (1), 1–1064.

  95. Takahashi, H. (2004) [Tremex nakanei in Higashiyamato City, Tokyo]. Gekkan-Mushi, 406, 16. [in Japanese]

  96. Takahashi, H. (2013) Male of Tremex nakanei Takeuchi, 1955 (Hymenoptera: Siricidae). Tsunekibachi, 23, 31–32. [in Japanese]

  97. Takakuwa, M. (2004) Kimadara-hiraashikibachi (Kibachi-ka) [Tremex fuscicornis (Siricidae)]. Newsletter of the Kanagawa Prefectural Museum of Natural History, 10, 31–32. [in Japanese]

  98. Takeuchi, K. (1938) A systematic study on the suborder Symphyta (Hymenoptera) of the Japanese Empire (1). Tenthredo, 2, 173–229.

  99. Takeuchi, K. (1949) A list of the food-plants of Japanese sawflies. Transactions of the Kansai Entomological Society, Osaka, 14, 47–50.

  100. Takeuchi, K. (1950) Siricidae. In: Ishii, T., Esaki, T., Kinoshita, S., Shiraki, T., Uchida, S., Kawamura, T., Kuwayama, S. & Yuasa, H. (Eds.), Iconographia Insectorum Japonicorum Editio Secunda Reformata. Hokuryûkan, Tokyo, pp. 1332–1334. [in Japanese]

  101. Takeuchi, K. (1955a) Siricidae of Japan (Hymenoptera). Akitu, Kyoto, 4, 1–9. [in Japanese with English abstract]

  102. Takeuchi, K. (1955b) Coloured Illustrations of the Insects of Japan. Vol. II. Hoikusha, Osaka, 190 pp., 68 pls. [in Japanese]

  103. Takeuchi, K. (1962) Insecta Japonica. Series 2. Part 4. Hymenoptera: Siricidae. Hokuryûkan, Tokyo, 2 + 12 pp., 1 pl. [in Japanese with English summary]

  104. Tanaka, C. (1971) A List of Insects of Toyama Prefecture. Privately published, Toyama, 242 pp. [in Japanese]

  105. Tanaka, C. (1979) Hymenoptera. In: Toyama-ken Konchû Kenkyû-kai (Ed.), [Insects of Toyama Prefecture]. Toyama Prefectural Government, Toyama, pp. 459–489. [in Japanese]

  106. Tanaka, S. (2015) [Catalogue of sawflies and woodwasps of Yamaguchi Prefecture]. Bulletin of the Firefly Museum of Toyata Town, (7), 27–44. [in Japanese]

  107. Terayama, M. (1982a) On the Siricidae (Hymenoptera) of Saitama Prefecture, Japan. Nature and Insects, Tokyo, 17 (9), 24–26. [in Japanese]

  108. Terayama, M. (1982b) [Hymenoptera of Kanagawa Prefecture, collection records]. Kanagawa-Chûho, 67, 25–31. [in Japanese]

  109. Togashi, I. (1965) Siricidae. In: Asahina, S., Ishihara, T. & Yasumatsu, K. (Eds.), Iconographia Insectorum Japonicorum Colore naturali edita. Vol. III. Hokuryûkan, Tokyo, pp. 244, pl. 122. [in Japanese]

  110. Togashi, I. (1973) Tenthredinoidea of Korea collected by Prof. K. Tsuneki in 1941–43 (Hym.). Life Study, Fukui, 17 (3–4), 103–112.

  111. Togashi, I. (1980) A new Tremex (Hymenoptera, Siricidae) from China. Kontyû, Tokyo, 48 (3), 321–323.

  112. Togashi, I. (1997) Two new species of Tremex Jurine (Hymenoptera: Siricidae) from Japan. Species Diversity, 2, 1–6.

  113. Togashi, I. (1998) Symphyta (Hymenoptera) of Hokkaido, Japan. Bulletin of the Biogeographical Society of Japan, Tokyo, 53, 39–47.

  114. Togashi, I. (2006) Two new species of Tremex (Hymenoptera: Siricidae) reared from Castanopsis and Symplocos in Japan. Proceedings of the Entomological Society of Washington, 108, 938–944.

  115. Ushirokita, M., Okuyama, S., Sakamoto, N., Nomoto, K., Kakumasa, M. & Nagashima, S. (2012) Catalogue of the Materials Deposited in the Itami City Museum of Insects, a Collection List of Insect Specimens of the Koyaike Park. Itami City Museum of Insects, Itami, 8 + 36 pp. [in Japanese]

  116. Verzhutskij, B.N. (1981) Rastitel'nojadnye Nasekomye v Ekosistemakh Vostochnoj Sibiri (Pilil'shchiki i Rogokhvosty). Nauka, Novosibirsk, 303 pp.

  117. Viitasaari, M. (2002) The suborder Symphyta of the Hymenoptera. In: Viitasaari, M. (Ed.), Sawflies (Hymenoptera, Symphyta) I. A review of the suborder, the Western Palaearctic taxa of Xyeloidea and Pamphilioidea. Tremex, Helsinki, pp. 11–174.

  118. Watanabe, A. (1986) [Woodwasps newly recorded from Okayama Prefecture]. Suzumushi, Kurashiki, 121, 31–32. [in Japanese]

  119. Xiao, G., Huang, X., Zhou, S., Wu, J. & Zhang, P. (1992) Economic Sawfly Fauna of China. Tianze Eldonejo, Beijing, 4 + 8 + 221 pp. [in Chinese]

  120. Xiao, G. & Wu, J. (1983) The Siricid woodwasps of China (Hymenoptera, Symphyta). Scientia Silvae Sinicae, 19, 1–29. [in Chinese with English abstract]

  121. Yafuso, M. (2002) Siricidae. In: Azuma, S., Yafuso, M., Kinjo, M., Hayashi, M., Kohama, T., Sasaki, T., Kimura, M. & Kawamura, M. (Eds.), Check List of the Insects of the Ryukyu Islands, Second Edition. The Biological Society of Okinawa, Nishihara, pp. 328. [in Japanese]

  122. Yamazaki, K. & Matsumoto, R. (2009) Predation on the woodwasp Tremex longicollis Konow (Hymenoptera: Siricidae) and its parasitoid Megarhyssa jezoensis (Matsukura) (Hymenoptera: Ichneu­monidae) adults during oviposition. Journal of Asia-Pacific Entomology, 12, 313–315.

  123. Yano, M. (1917) A catalogue of recorded Japanese Siricidae, with description of a new species. In: Nagano, K. (Ed.), A Collection of Essays for Mr. Yasushi Nawa: Written in Commemoration of his Sixtieth Birthday, October 8, 1917. Nawa Entomological Laboratory, Gifu, pp. 115–121. [in Japanese with English abstract]

  124. Yano, M. (1932) Siricidae. In: Issiki, S., Ishii, T., Niijima, Y., Hori, H., Hozawa, S., Okamoto, H., Oguma, M., Kawada, A., Kawamura, T., Yokoyama, K., Takahashi, R., Uchida, S., Ueno, M., Kuwana, I., Kuwayama, S., Yano, M., Yamada, S., Maruge, N., Koizumi, M., Kojima, T., Esaki, T., Kinoshita, S., Kishida, K., Yuasa, H., Shiraki, T. & Shinji, O., Iconographia Insectorum Japonicorum Editio Prima. Hokuryûkan, Tokyo, pp. 471–475. [in Japanese]

  125. Yonekura, K. & Kajita, T. (2022) Shokubutsu Wamei–Gakumei Index, YList. Available from: (accessed 18 November 2022) [in Japanese]

  126. Yoshida, H. (2006) Symphyta (Hymenoptera) of Osaka Prefecture, Japan. West Japan Hymenopterists’ Club, Kakogawa, 24 pls., 128 pp. [in Japanese]