Skip to main content Skip to main navigation menu Skip to site footer
Type: Monograph
Published: 2022-11-22
Page range: 1-96
Abstract views: 304
PDF downloaded: 37

Checklist of Oribatid mites (Acari: Oribatida) of Ecuador

Colegio de Postgraduados, Campus Montecillo, Texcoco, Mexico state, Mexico.
Acari biodiversity Latin America soil taxonomy

Abstract

The present checklist of the oribatid mites compiles all known published records of Ecuador with 479 species belonging to 232 genera and 89 families; 180 of these species were described from Ecuador. Oribatid mites were found in 15 of the 24 provinces that conform the Ecuadorian territory. In the provinces of Zamora Chinchipe, Galápagos, Loja, Cotopaxi, Pichincha and Napo, 95% of the species were recorded. 23.6% of these species are known only from Ecuador, 47.8% are distributed in Neotropical region, the remaining are Pantropical, semicosmopolitan and cosmopolitan. This high number of species makes Ecuador the country with the second largest number of species records in Latin America after Brazil.

 

References

  1. Acharya, S. & Datta, T. (2019) Diversity of soil cryptostigmatid mites (Acari: Oribatida) of Himachal Pradesh, India, from an altitudinal perspective. Journal of Asia-Pacific Biodiversity, 12 (3), 357–362. https://doi.org/10.1016/j.japb.2019.03.014
    Balogh, J. & Balogh, P. (1985) Fifteen new species of the genus Xenillus Robineau-Desvoidy, 1839 (Acari: Oribatei) from South America. Acta Zoologica Hungarica, 31 (1–3), 53–79.
    Balogh, J. & Balogh, P. (1988) Oribatid mites of the Neotropical Region I. 3rd Edition. Elsevier Science Publishing Company, Inc., Amsterdam. 355 pp.
    Balogh, J. & Balogh, P. (1990) The soil mites of the world. Oribatid mites on Neotropical Region II. Akadémiai Kiado, Budapest, 333 pp.
    Balogh, J. & Balogh, P. (2002a) Identification Keys to the Oribatid Mites of the Extra-Holarctic Regions II. Well-PRess Publishing Limited, Budapest, 372 pp.
    Balogh, J. & Balogh, P. (2002b) Identification Keys to the Oribatid Mites of the Extra-Holartic Regions I. Well-PRess Publishing Limited, Budapest, 453 pp.
    Balogh, P. (1985) Galapagacarus schatzi gen. et sp. n. (Acari: Oribatei). Opuscula Zoologica Budapest, 21, 31–34.
    Balogh, P. (1986) Four new Xenillus species from the Neotropical Region (Acari: Oribatei). Opuscula Zoologica Budapest, 22, 43–50.
    Balogh, P. (1988) Oribatid mites from Ecuador (Acari). Acta Zoologica Hungarica, 34 (4), 321–338.
    Balogh, P. (1989) Oribatid mites from Ecuador II. Acta Zoologica Hungarica, 35 (1–2), 17–28.
    Bayartogtokh, B. & Ermilov, S. (2019) A new species of Hydrozetes (Acari: Oribatida, Hydrozetidae) from Ecuador and a key to Neotropical species. International Journal of Acarology, 45 (5), 328–334. https://doi.org/10.1080/01647954.2019.1630483
    Beck, L. (1963) Zur Ökologie und Taxionomie der neotropischen Bodentiere I. Zur Oribatiden-Fauna Perus. Zoologische Jahrbücher, 90, 299–392.
    Behan-Pelletier, V. & Ermilov, S. (2019) Trichoribates sidorchukae sp. nov. (Acari, Oribatida, Ceratozetidae) from tropical montane Ecuador, with revised generic diagnosis. Zootaxa, 4647 (1), 348–361. https://doi.org/10.11646/zootaxa.4647.1.21
    Behan-Pelletier, V. & Walter, D.E. (2009) Biodiversity of oribatid mites (Acari: Oribatida) in tree canopies and litter. In: Coleman, D. & Hendrix, P.F. (Eds.), Invertebrates as webmasters in ecosystems. CABI Publishing,Wallingford, pp. 187–202. https://doi.org/10.1079/9780851993942.0187
    Bernal, A., Castaño-Meneses, G., Palacios-Vargas, J.G. & García, N. (2009) Oribatid mites and springtails from a coffee plantation in Sierra Sur, Oaxaca, Mexico. Pesquisa Agropecuária Brasileira, 44 (8), 988–995. https://doi.org/10.1590/s0100-204x2009000800030
    Colloff, M.J. (1993) A taxonomic revision of the oribatid mite genus Camisia (Acari: Oribatida). Journal of Natural History, 1993, 1325–1428. https://doi.org/10.1080/00222939300770761
    Dhooria, M.S. (2016) Fundamentals of Applied Acarology. Springer, Singapore, 473 pp. https://doi.org/10.1007/978-981-10-1594-6
    Dindal, D. (1977) Biology of Oribatid Mites. State University of New York. 130 pp.
    Ermilov, S. (2016a) A new species of the genus Malaconothrus (Acari, Oribatida, Malaconothridae) from Ecuador. Ecologica Montenegrina, 9, 31–37. https://doi.org/10.37828/em.2016.9.5
    Ermilov, S. (2016b) Contribution to the knowledge of the oribatid mite genus Unguizetes (Acari, Oribatida, Mochlozetidae). Systematic and Applied Acarology, 21 (10), 1281–1299. https://doi.org/10.11158/saa.21.10.1
    Ermilov, S. (2019a) New findings of oribatid mites of the family Mochlozetidae (Acari: Oribatida) from Ecuador. Biologia, 74 (10), 1313–1324. https://doi.org/10.2478/s11756-019-00248-5
    Ermilov, S. (2019b) New Galumnidae (Acari, Oribatida) from Ecuador. Systematic and Applied Acarology, 24 (3), 348–361. https://doi.org/10.11158/saa.24.3.2
    Ermilov, S. (2020) Two new species of oribatid mites (Acari, Oribatida) of the genus Xenillus Robineau-Desvoidy (Liacaridae) from Ecuador. Zoologicheskii Zhurnal, 99 (2), 157–164. https://doi.org/10.31857/S0044513420020105
    Ermilov, S. Bayartogtokh, B., Sandmann, D., Marian, F. & Maraun, M. (2013) New and little known species of oribatid mites of the family Haplozetidae (Acari, Oribatida) from Ecuador. ZooKeys, 346, 43–57. https://doi.org/10.3897/zookeys.346.6436
    Ermilov, S. & Behan-Pelletier, V.M. (2014) Revision of Fenestrobates (Acari, Oribatellidae) with description of F. marauni sp. nov., from South America, and new diagnosis for Oribatellidae. Zootaxa, 3827 (2), 258–272. https://doi.org/10.11646/zootaxa.3827.2.7
    Ermilov, S., Friedrich, S. & Kalúz, S. (2016) A new species of Neoribates (Neoribates) (Acari: Oribatida: Parakalummidae) with key to the Neotropical species of the subgenus. Biologia, 71 (6), 673–677. https://doi.org/10.1515/biolog-2016-0079
    Ermilov, S. & Mary, N.J. (2019) To the knowledge of oribatid mites (Acari, Oribatida) of the Comoro lslands. Systematic & Applied Acarology, 24 (4), 624–637. https://doi.org/10.11158/saa.24.4.8
    Ermilov, S., Sandmann, D. & Maraun, M. (2013a) A new species of the genus Orthozetes from Ecuador (Acari: Oribatida: Microzetidae). Acarina, 21 (2), 100–103.
    Ermilov, S., Sandmann, D. & Maraun, M. (2013b) Two new species of Schalleria (Acari: Oribatida: Microzetidae) from Ecuador with a key to all species of the genus. International Journal of Acarology, 39 (3), 200–208. https://doi.org/10.1080/01647954.2012.762040
    Ermilov, S., Sandmann, D., Marian, F. & Maraun, M. (2013a) New oribatid mites of the genera Machadobelba and Campachipteria (Acari: Oribatida) from Ecuador. Systematic and Applied Acarology, 18 (2), 145–152. https://doi.org/10.11158/saa.18.2.7
    Ermilov, S., Sandmann, D., Marian, F. & Maraun, M. (2013b) Oribatid mites of the superfamily Oppioidea from Ecuador (Acari: Oribatida). Systematic and Applied Acarology, 18 (3), 218–224. https://doi.org/10.11158/saa.18.3.3
    Ermilov, S., Sandmann, D., Marian, F. & Maraun, M. (2013c) Perscheloribates paratzitzikamaensis n. sp., with supplementary descriptions of Scheloribates elegans and Monoschelobates parvus (Acari, Oribatida, Scheloribatidae) from Ecuador. Acarologia, 53 (4), 429–437. https://doi.org/10.1051/acarologia/20132104
    Ermilov, S., Sandmann, D., Marian, F. & Maraun, M. (2013d) Three new species of oribatid mites (Acari, Oribatida) from Ecuador. Acarologia, 53 (1), 111–123. https://doi.org/10.1051/acarologia/20132075
    Ermilov, S., Sandmann, D., Marian, F. & Maraun, M. (2013e) Three new species of the genus Sternoppia (Acari: Oribatida: Sternoppiidae) from Ecuador. Zootaxa, 3641 (5), 565–576. https://doi.org/10.11646/zootaxa.3641.5.5
    Ermilov, S., Sandmann, D., Marian, F. & Maraun, M. (2013f) Two new oribatid mite species of the genus Gittella from Ecuador (Acari, Oribatida, Oppiidae). Spixiana, 36 (1), 1–8.
    Ermilov, S., Sandmann, D., Marian, F. & Maraun, M. (2013g) Two new species of oribatid mites of the genus Truncozetes (Acari, Oribatida, Epactozetidae) from Ecuador. ZooKeys, 303, 23–31. https://doi.org/10.3897/zookeys.303.5309
    Ermilov, S., Sandmann, D., Marian, F. & Maraun, M. (2014) New species of oribatid mites of the genera Hermannobates and Rhynchoribates (Acari: Oribatida: Hermanniellidae, Rhynchoribatidae) from Ecuador. Systematic and Applied Acarology, 19 (3), 313–321. https://doi.org/10.11158/saa.19.3.5
    Ermilov, S., Starý, J., Sandmann, D., Marian, F. & Maraun, M. (2013) New taxa and new records of oribatid mites of the family Galumnidae (Acari: Oribatida) from Ecuador. Zootaxa, 3700 (2), 259–270. https://doi.org/10.11646/zootaxa.3700.2.4
    Ermilov, S. & Kalúz, S. (2012a) A new species of the genus Unguizetes (Acari: Oribatida: Mochlozetidae) from Ecuador. Acarina, 20 (2), 180–184.
    Ermilov, S. & Kalúz, S. (2012b) A new subgenus and three new species of oribatid mites of the family Scheloribatidae (Acari: Oribatida) from Ecuador. Annales Zoologici, 62 (4), 773–787. https://doi.org/10.3161/000345412X659795
    Ermilov, S. & Kalúz, S. (2012c) Four new species of the superfamily Galumnoidea (Acari: Oribatida) from Ecuador. Zootaxa, 3481 (1), 27–38. https://doi.org/10.11646/zootaxa.3481.1.2
    Ermilov, S. & Kalúz, S. (2012d) The oribatid mite genus Ceratorchestes (Acari: Oribatida: Peloppiidae). Acarologia, 52 (2), 165–172. https://doi.org/10.1051/acarologia/20122046
    Ermilov, S. & Kalúz, S. (2012e) Two new species of Oppiidae (Acari: Oribatida) from Ecuador. International Journal of Acarology, 38 (6), 521–527. https://doi.org/10.1080/01647954.2012.687499
    Ermilov, S. & Kalúz, S. (2012f) Two new species of oribatid mites (Acari, Oribatida) from Ecuador. Systematic and Applied Acarology, 17 (3), 269–280. https://doi.org/10.1134/S0013873815010169
    Ermilov, S. & Kalúz, S. (2014) A new oribatid mite of the genus Rhynchoribates (Acari, Rhynchoribatidae) from Ecuador. Entomological Review, 94 (5), 777–780. https://doi.org/10.1134/S0013873814050108
    Franklin, E., Santos, E. & Albuquerque, M. (2006) Diversity and distribution of oribatid mites (Acari: Oribatida) in a lowland rain forest in Peru and in several environments of the Brazilians states of Amazonas, Rondonia, Roraima an Pará. Brazilian Journal of Biology, 66 (4), 999–1020. https://doi.org/10.1590/S1519-69842006000600007
    Fredes, N. (2019) Catalogue of oribatid mites (Acari: Oribatida) from Argentina. Zootaxa, 4406 (1), 1–190. https://doi.org/10.11646/zootaxa.4716.1.1
    Halliday, R. & Norton, R.A. (2019) Case 3785—Proposal to remove the homonymy of Cepheidae Berlese, 1896 (Acariformes) with Cepheidae Agassiz, 1862 (Cnidaria), by emending the former to Cepheusidae. Bulletin of Zoological Nomenclature, 76 (1), 34–42. https://doi.org/10.21805/bzn.v76.a040
    Haq, M. (1994) Role of oribatid mites in soil ecosystem. In: Bhandari, S. & Somani, L. (Eds.), Ecology and Biology of Soil Organism. Agrotech Publishing Academy, Udaipur, pp. 143–178.
    Illig, J., Langel, R., Norton, R., Scheu, S. & Maraun, M. (2005) Where are the decomposers? Uncovering the soil food web of a tropical montane rain forest in southern Ecuador using stable isotopes (15N). Journal of Tropical Ecology, 21 (5), 589–593. https://doi.org/10.1017/S0266467405002646
    Illig, J., Norton, R.A., Scheu, S. & Maraun, M. (2010) Density and community structure of soil and bark-dwelling microarthropods along an altitudinal gradient in a tropical montane rainforest. Experimental and Applied Acarology, 52, 49–62. https://doi.org/10.1007/s10493-010-9348-x
    Illig, J., Sandmann, S., Schatz, H., Scheu, S. & Maraun, N. (2007) Oribatida (Mites) Checklist Reserva Biológica San Francisco (Prov. Zamora Chinchipe, S. Ecuador). Ecotropical Monographs, 4, 221–230.
    Illig, J., Schatz, H., Scheu, S. & Maraun, M. (2008) Decomposition and colonization by micro-arthropods of two litter types in a tropical montane rain forest in southern Ecuador. Journal of Tropical Ecology, 24, 157–167. https://doi.org/10.1017/S0266467407004750
    Kuty, M. (2005) The morphology of juvenile stages of Crotonia pulcher (Beck, 1962) (Acari: Oribatida: Crotonioidea) with a redescription of the adult. Zoologischer Anzeiger, 244, 125–136. https://doi.org/10.1016/j.jcz.2005.06.001
    Kuty, M. (2006a) A new species of the genus Malaconothrus (Acari, Oribatida, Malaconothridae) from Ecuador. Genus, 17 (2), 307–310. https://doi.org/10.37828/em.2016.9.5
    Kuty, M. (2006b) Nothrus olszanowskii and Nothrus pallidulus (Acari: Oribatida: Nothridae) new Crotonioid mites from Ecuador. Annales Zoologici, 56 (4), 799–804.
    Kuty, M. (2006c) Nothrus piriformis a new crotonioid mite from Ecuador (Acari: Oribatida: Nothridae). Genus, 17 (4), 621–626.
    Kuty, M. (2007) Description of juvenile stages and adults of two new nothroid mites from Ecuador (Acari: Oribatida: Nothridae). Journal of Natural History, 41 (9–12), 597–618. https://doi.org/10.1080/00222930701265126
    Kuty, M. & Olszanowski, Z. (2006) A new species of Nothrus (Acari: Oribatida: Nothridae) from Ecuador and remarks on N. becki. New Zealand Journal of Zoology, 33, 235–240. https://doi.org/10.1080/03014223.2006.9518452
    Kuty, M. & Olszanowski, Z. (2008) The morphology of juvenile stages of Platynothrus altimontanus Hammer, 1958 (Acari: Oribatida: Camisiidae) with redescription of adult. Zoologischer Anzeiger, 247, 3–14. https://doi.org/10.1016/j.jcz.2006.11.002
    Łochyniska, M. (2008) Nothrid mites from Ecuador—recapitulation (Acari: Oribatida: Nothridae). Genus, 19 (1), 143–149.
    Maraun, M., Fronczek, S., Marian, F., Sandmann, D. & Scheu, S. (2013b) More sex at higher altitudes: Changes in the frequency of parthenogenesis in oribatid mites in tropical montane rain forests. Pedobiologia, 56, 185–190. https://doi.org/10.1016/j.pedobi.2013.07.001
    Maraun, M., Illig, J., Krashevska, V., Norton, R.A. & Scheu, S. (2008) Soil Fauna. In: Beck, E., Bendix, J., Kottke, I., Makeschin, F. & Mosandl, R. (Eds.), Gradients in a tropical mountain ecosystem of Ecuador. Ecological studies. Springer, Berlin and Heidelberg, pp. 181–192. https://doi.org/10.1007/978-3-540-73526-7_18
    Marian, F., Ramírez-Castillo, P., Iñiguez-Armijos, C., Günter, S.M., Maraun, M. & Scheu, S. (2020) Conversion of Andean montane forests into plantations: Effects on soil characteristics, microorganisms, and microarthropods. Biotropica, 52 (6), 1142–1154. https://doi.org/10.1111/btp.12813
    Marian, F., Sandmann, D., Krashevska, V., Maraun, M. & Scheu, S. (2018) Altitude and decomposition stage rather than litter origin structure soil microarthropod communities in tropical montane rainforests. Soil Biology and Biochemistry, 125, 263–274. https://doi.org/10.1016/j.soilbio.2018.07.017
    McGugan, J.R., Byrd, G.D., Roland, A.B., Caty, S.N., Kabir, N., Tapia, E.E., Trauger, S.A., Coloma, L.A. & O’Connell, L.A. (2016) Ant and Mite Diversity Drives Toxin Variation in the Little Devil Poison Frog. Journal of Chemical Ecology, 42, 537–551. https://doi.org/10.1007/s10886-016-0715-x
    Miko, L. (2006) Oppiide Grandjean, 1951. In: Weigmann, G. (Ed.), Hornmilben (Oribatida) [Oribatid mites (Oribatida)]. In: Dahl, F. (Series Founder), Die Tierwelt Deutschlands, The fauna of Germany, Part 76, pp. 1–520.
    Miko, L. & Ermilov, S. (2021) Parabelbella (Paradyobelba) elchacoensis subgen. nov., sp. nov. (Acari, Oribatida, Damaeidae), an unusual representative of the genus from Ecuador. Systematic and Applied Acarology, 26 (3), 599–609. https://doi.org/10.11158/saa.26.3.9
    Niedbała, W. (1988) Nouceaux Phthiracaroidea (Acari, Oribatida) Neotropicales. Redia, 71, 115–138.
    Niedbała, W. (1989) Notophthiracarus schatzii sp. nov. d’iles Galápagos (Acari, Oribatida, Phthiracaroides). Bulletin of the Polish Academy of Sciences Biological Sciences, 37 (13), 75–78.
    Niedbała, W. (2001) Ptyctimous mites (Acari: Oribatida) from Florida. Acta Zoologica Cracoviensia, 44 (3), 285–367.
    Niedbała, W. (2004) Ptyctimous mites (Acari, Oribatida) of the neotropical region. Annales Zoologici, 54 (1), 1–288.
    Niedbała, W. (2008) Supplement to the knowledge of ptyctimous mites (Acari, Oribatida) from Palaearctic Region. Genus, 19 (4), 729–818.
    Niedbała, W. & Ermilov, S. (2014) Additions to the Ecuadorian, Ethiopian and Nepalese ptyctimous mite fauna (Acari, Oribatida), with description of two new species and remarks on some known species. International Journal of Acarology, 40 (3), 254–262. https://doi.org/10.1080/01647954.2014.906499
    Niedbała, W. & Illig, J. (2007a) New species and records of ptyctimous mites (Acari, Oribatida) from Ecuador. Tropical Zoology, 20, 135–150.
    Niedbała, W. & Illig, J. (2007b) Ptyctimous mites (Acari, Oribatida) from the Ecuador rainforest. Journal of Natural History, 41 (13–16), 771–777. https://doi.org/10.1080/00222930701350159
    Niedbała, W. & Liu, D. (2018) Catalogue of ptyctimous mites (Acari, Oribatida) of the world. Zootaxa, 4393 (1), 1–238. https://doi.org/10.11646/zootaxa.4393.1.1
    Niedbała, W. & Roszkowska, M. (2016) Ptyctimous mites (Acari, Oribatida) of Ecuador (South America). International Journal of Acarology, 43 (3), 251–262. https://doi.org/10.1080/01647954.2016.1247910
    Niedbała, W. & Schatz, H. (1996) Euptyctimous mites from the Galapagos Islands, Cocos Island, and Central America (Acari: Oribatida). Genus, 7 (2), 239–317.
    Norton, R. (1994) Evolutionary aspects of oribatid mite life histories and consequences for the origin of the Astigmata. In: Houck, M. (Ed.), Mites. Springer, Boston, Massachusetts, pp. 99–135. https://doi.org/10.1007/978-1-4615-2389-5_5
    Norton, R.A. & Behan-Pelletier, V. (2009) Suborder Oribatida. In: Krantz, G.W. & Walter, D.E. (Eds.), A Manual of Acarology. 3rd Edition. Texas Tech University Press, Lubbock, Texas, pp. 430–564.
    Norton, R.A. & Behan-Pelletier, V.M. (1991) Calcium carbonate and calcium oxalate as cuticular hardening agents in oribatid mites (Acari: Oribatida). Canadian Journal of Zoology, 69, 1504–1511. https://doi.org/10.1139/z91-210
    Norton, R.A. & Ermilov, S. (2019) Anderemaeus (Acari, Oribatida)-overview, three new species from South America and reassessment of Anderemaeidae supported by ontogeny. Zootaxa, 4647 (1), 241–289. https://doi.org/10.11646/zootaxa.4647.1.17
    Ojeda, M. & Gasca-Pineda, J. (2019) Abundance and diversity of the soil microarthropod fauna from the Cuatro Ciénegas Basin. In: Álvarez, F. & Ojeda, M. (Eds.), Animal Diversity and Biogeography of the Cuatro Ciénegas Basin. Springer, Cham, pp. 29–51. https://doi.org/10.1007/978-3-030-11262-2_3
    Ojeda, M., Velez, P., Espinosa-Asuar, L., Eguiarte, L.E. & Souza, V. (2020) A new species of Trhypochthoniellus (Acari: Oribatida: Trhypochthoniidae) from Cuatro Ciénegas, Coahuila, Mexico, and a key to the world species. Systematic and Applied Acarology, 25 (6), 974–985. https://doi.org/10.11158/saa.25.6.3
    Oliveira, A.R., Argolo, P.S., De Moraes, G.J., Norton, R.A. & Schatz, H. (2017) A checklist of the oribatid mite species (Acari: Oribatida) of Brazil. Zootaxa, 4245 (1), 1–89. https://doi.org/10.11646/zootaxa.4245.1.1
    Pachl, P., Lindl, A. C., Krause, A., Scheu, S., Schaefer, I. & Maraun, M. (2017) The tropics as an ancient cradle of oribatid mite diversity. Acarologia, 57 (2), 309–322. https://doi.org/10.1051/acarologia/20164148
    Palacios-Vargas, J.G., Castaño-Meneses, G. & Estrada, D. (2011) Diversity and dynamics of microarthropods from different biotopes of Las Sardinas cave (Mexico). Subterranean Biology, 9 (1), 113–126. https://doi.org/10.3897/subtbiol.9.2514
    Palacios-Vargas, J.G. & Iglesias, R. (2004) Oribatei (Acari). In: Llorente Bousquets, J., Morrone Juan, J., Yáñez Ordóñez, O. & Vargas Fernández, I. (Eds.), Biodiversidad, taxonomía y biogeografía de artrópodos: hacia una síntesis de su conocimiento. Vol. IV. CONABIO, México, D.F., pp. 431–468.
    Pfingstl, T. & Baumann, J. (2017) Morphological diversification among island populations of intertidal mites (Acari, Oribatida, Fortuyniidae) from the Galaápagos archipelago. Experimental and Applied Acarology, 72, 115–131. https://doi.org/10.1007/s10493-017-0149-3
    Pfingstl, T. & Schatz, H. (2017) New littoral mite species (Acari, Oribatida, Fortuyniidae) from the Galápagos archipelago, with ecological and zoogeographical considerations. Zootaxa, 4244 (1), 39–64. https://doi.org/10.11646/zootaxa.4244.1.2
    Ramírez-Castillo, P., Marian, L., Marian, F., Günter, S., Espinosa, C.I., Maraun, M. & Scheu, S. (2018) Response of oribatid mites to reforestation of degraded tropical montane pastureland. European Journal of Soil Biology, 84, 35–41. https://doi.org/10.1016/j.ejsobi.2017.09.009
    Ron, R. S. (2020) Regiones naturales del Ecuador. BIOWEB, Pontificia Universidad Católica Del Ecuador, Quito, ‎Pichincha Province‎. Available from: https://bioweb.bio/faunaweb/reptiliaweb/RegionesNaturales (accessed 15 June 2022)
    Salazar-Fillipo, A. & Miko, L. (2022) Checklist of the oribatid mites (Acari: Oribatida) of the Republic of Colombia. Zootaxa, 5087 (1), 35–58. https://doi.org/10.11646/zootaxa.5087.1.2
    Schatz, H. (1991) Arrival and establishment of Acari on oceanic islands. In: Dusbábek, F. & Bukva, V. (Eds.), Modern Acarology. Academia, Prague and SPB Academic Publishing bv, The Hague, pp. 613–618.
    Schatz, H. (1993) The genus Lohmannia (Acari, Oribatida, Lohmanniidae) in the Galapagos Islands. Acarologia, 34 (1), 69–84.
    Schatz, H. (1994a) Lohmanniidae (Acari: Oribatida) from the Galapagos Islands, the Cocos Island, and Central America. Acarologia, 35 (3), 267–287.
    Schatz, H. (1994b) New records od the genus Torpacarus (Acari: Oribatida: Lohmanniidae) from the Galápagos Islands and Central America. Acarologia, 35 (2), 167–179.
    Schatz, H. (1998) Oribatid mites of the Galapagos Islands—Faunistics, ecology and speciation. Experimental and Applied Acarology, 22, 373–409. https://doi.org/10.1023/A:1006097928124
    Schatz, H. (2001) The genus Eremaeozetes (Acari: Oribatida) on the Galápagos islands. Acarologia, 41 (4), 475–493.
    Schatz, H. (2003) New Sphaerochthonius species from the Neotropical region (Acari: Oribatida). Revue Suisse de Zoologie, 110 (1), 111–124. https://doi.org/10.5962/bhl.part.80178
    Schatz, H. (2006) Catalogue of known oribatid mite species (Acari Oribatida) from the Central American landbridge (First part). Tropical Zoology, 19, 209–288.
    Schatz, H. & Palacios-Vargas, J.G. (1999) Microzetidae (Acari: Oribatida) from the Galápagos Islands. Acarologia, 40 (1), 69–83.
    Schatz, H., Behan-Pelletier, V., OConnor, B. & Norton, R.A. (2011) Suborder Oribatida van der Hammen, 1968. In: Zhang, Z. (Ed.), Animal Biodiversity: An outline of higher-level classification and survey of taxonomic richness. Zootaxa, 3148 (1), pp. 141–148. https://doi.org/10.11646/zootaxa.3148.1.26
    Scheu, S., Illig, J., Eissfeller, V., Krashevska, V., Sandmann, D. & Maraun, M. (2008) The soil fauna of a tropical mountain rainforest in southern Ecuador: Structure and functioning. Biodiversity and Ecology Series, 2, 81–98.
    Sitnikova, L.G. (1979) A new genus and new species of the mite family Cepheidae from South America. Annales Historico-Naturales Musei Nationalis Hungarici, 71, 1–14. [in Russian]
    Subías, L. (2004) Listado sistemático, sinonímico y biogeográfico de los ácaros oribátidos (Acariformes, Oribatida) del mundo (1758–2002). Graellsia, 60, 3–305. https://doi.org/10.3989/graellsia.2004.v60.iExtra.218
    Subías, L. (2022) Listado sistemático , sinonímico y biogeográfico de los ácaros oribátidos (Acariformes: Oribatida) del mundo (excepto fósiles), 17a actualización. Available from: http://bba.bioucm.es/cont/docs/RO_1.pdf (accessed 10 June 2022)
    Vázquez, M., May, D. & Alamilla, E. (2016) Riqueza específica y biodiversidad de Cozumel, Quintana Roo, México. Teoría y Praxis, 12 (19), 137–171. https://doi.org/10.22403/UQROOMX/TYP19/07
    Villagomez, F. & Palacios-Vargas, J.G. (2017) Los ácaros oribátidos (Acari: Oribatei) de los estratos edáficos y arbóreos de la selva alta perennifolia de los Tuxtlas, Veracruz. Entomología Mexicana, 4, 28–34.
    Villagomez, F., Palacios-Vargas, J.G., Castaño-Meneses, G. & Castellanos-Vargas, I. (2019) Effect of tree identity, temporal variation and edaphic parameters on the structure of the edaphic community of oribatid mites in an evergreen tropical forest of Mexico. Applied Ecology and Environmental Research, 17 (6), 14621–14639. https://doi.org/10.15666/aeer/1706_1462114639
    Zhu, X., Gao, B., Yuan, S. & Hu, Y. (2010) Community structure and seasonal variation of soil arthropods in the forest-steppe ecotone of the mountainous region in Northern Hebei, China. Journal of Mountain Science, 7 (2), 187–196. https://doi.org/10.1007/s11629-010-0198-0