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Type: Monograph
Published: 2022-10-03
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A review of the genus Brachysomus Schoenherr (Coleoptera: Curculionidae: Entiminae)

University of Oslo, Natural History Museum, Department of Zoology, P.O. Box 1172 Blindern, NO-0318 Oslo, Norway.
Coleoptera Sciaphilini taxonomy morphology identification key new species new synonym type designation new combination new record

Abstract

The genus Brachysomus Schoenherr, 1823 is reviewed. A comparative morphological study of adults from 56 Brachysomus species is provided. Redescriptions, diagnoses and identification keys are given. Descriptions of two new species are provided: B. (s. str.) podlussanyi sp. n. from Greece and B. (s. str.) pseudosetiger sp. n. from Bulgaria, which appear to be closely related to B. (s. str.) mihoki Penecke, 1914. Females of B. (s. str.) longipterus Białooki, 2007 are described. The poorly known B. (Hippomias) moczarskii Penecke, 1924 is restored from synonymy with B. (H.) oertzeni Faust, 1889. New synonyms: B. (s. str.) echinatus (Bonsdorff, 1785) = B. (s. str.) hirsutus Iablokoff-Khnzorian, 1958 syn. n. New combinations: Brachysomus (s. str.) lituratus (Reitter, 1884) comb. n. from Foucartia Jacquelin du Val, 1854; Eurosphalmus tenuicollis (Yunakov, 2006) comb. n. from Brachysomus, Eurosphalmus cribrarius (Białooki, 2007) comb. n. from Brachysomus, Nanomias skodai (Białooki, 2007) comb. n. from Brachysomus, and Chiloneus bonnairei (Hoffmann, 1942) comb. n. from Brachysomus. Lectotypes are designated for: Strophosomus hirtus Boheman, 1845; Brachysomus ornatus Stierlin, 1892; B. bensae Stierlin, 1893; Platytarsus fasciatus Stierlin, 1899; and P. setiger Gyllenhal, 1840. For each species, all known localities are presented on a distribution map. Ecological preferences of Brachysomus species are specified. First records of Brachysomus (s. str.) fremuthi Košťál, 1991 from Romania, B. (Hippomias) carpathicus Košťál, 1992 from Serbia, and B. (H.) ponticus Apfelbeck, 1898 from Greece are given.

 

References

  1. Alonso-Zarazaga, M.A. & Lyal, C.H.C. (1999) A world catalogue of families and genera of Curculionoidea (Insecta: Coleoptera) (Excepting Scolytidae and Platypodidae). Entomopraxis, Barcelona, 315 pp.
    Alonso-Zarazaga, M.A., Barrios, H., Borovec, R., Bouchard, P., Caldara, R., Colonnelli, E., Gültekin, L., Hlavác, P., Korotyaev, B., Lyal, C.H.C., Machado, A., Meregalli, M., Pierotti, H., Ren, L., Sánchez-Ruiz, M., Sforzi, A., Silfverberg, H., Skuhrovec, J., Trýzna, M., Velázquez, de Castro, A.J. & Yunakov, N.N. (2017) Cooperative Catalogue of Palaearctic Coleoptera Curculionoidea. Monografías electrónicas S.E.A., 8, 1–729.
    Anderson, R.S. (1995) An evolutionary perspective on diversity in Curculionidae. Memoirs of the Entomological Society of Washington, 14, 103–114.
    Angelov, P. (1978) Coleoptera, Curculionidae. 2. Brachiderinae, Brachicerina, Tanymecinae, Cleoninae, Curculioninae, Myorrininae. Fauna na Bulgaria. Vol. 7. Bulgarian Academy of Sciences, Sofia, 234 pp.
    Apfelbeck, V. (1898) K poznannju palearktickih curculionida (Coleoptera). Synonymicki i zoogeograpficki prinosi sa opisom novyh vrsta sa balkanskogo poluostrva [Contribution to Palaearctic Curculionidae. Synonymy and geographical data and descriptions of new species in the Balkan peninsula] Glasnik zemeljskog muzeja Bosni i Hercegovini, 10, 265–313.
    Apfelbeck, V. (1899) Zur Kenntniss der paläarktischen Curculioniden. Systematische und zoogeographische Beiträge nebst Beschreibungen neuer Arten von der Balkanhalbinsel. Otiorhynchini et Brachyderini. Wissenschaftliche Mitteilungen aus Bosnien und der Hercegowina, 6, 773–816.
    Apfelbeck, V. (1911) Contributiones ad coleopterorum faunam peninsulae balcanicae. Prinosi poznavanju faune koleoptera Balkanskog Poluostrova. II. Dio. Glasnik Zemaljskog Muzeja u Bosni i Hercegovini, 23, 209–223.
    Arnoldi, L.V., Zaslavskij, V.A. & Ter-Minasian, M.E. (1965) Curculionidae—Dolgonosiki. In: Bej-Bijenko, V.G. (Ed.), Opredelitel nasekomykh Evropeiskoi chasti SSSR. Vol 2. Zhestkokrylye i veerokrylye. Nauka, Moskva and Leningrad, pp. 485–621. [in Russian]
    Bedel, L. (1883) Faune des Coléoptères du Bassin de la Seine. Vol. VI. Rhynchophora. Annales de la Société Entomologique de France, Hors Série, Series 6, 2 (4), 17–32, pl. I. [1882]
    Benedikt, S. (2001) Brachysomus rokosensis sp. n. from Slovakia with remarks on distribution and bionomics of the genus Brachysomus in West Carpathian Mts. (Coleoptera: Curculionidae). Klapalekiana, 37, 139–146.
    Benedikt, S. (2009a) Amphigonic populations of the weevil species Brachysomus echinatus and Cathormiocerus aristatus (Coleoptera: Curculionidae: Entiminae) in the Western Carpathians. Klapalekiana, 45, 1–7.
    Benedikt, S. (2009b) Brachysomus bohemicus sp. n. from the Czechia (Coleoptera: Curculionidae). Klapalekiana, 45, 169–176.
    Białooki, P. (2007) On new taxa of Entiminae (Coleoptera: Curculionidae) from Turkey and South-Eastern Europe. Annales of Upper Silesian Museum, Entomology, 14–15, 135–195.
    Białooki, P. (2015) Descriptions of new taxa of Otiorhynchini and related tribes (Coleoptera: Curculionidae: Entiminae) from the Middle East and Balkans. Israel Journal of Entomology, 44-45, 13–50.
    Bright, D.E. & Bouchard P. (2008) The Insects and Arachnids of Canada. Part 25. Coleoptera, Curculionidae, Entiminae. NRC Research Press, Ottawa, 327 pp.
    Bonsdorff, G. (1785) s.n. In: Historia Naturalis Curculionum Sveciae. Cujus partem secundam, consent, experient. facuit, med. examini offerunt auctor Gabriel Bonsdorff, Philos. Magist, Medic. Doctor, et respondens Petrus Ant. Norlin, stockholmensis. Johan Edman, Upsaliae, pp. 19–42, 1 pl.
    Borovec, R. (2006) Five new species of Argoptochus from Greece and Macedonia (Coleoptera: Curculionidae: Entiminae: Phyllobiini). Klapalekiana, 42, 45–77.
    Borovec, R. (2009) Revision of the Palaearctic supraspecific taxa of the tribe Trachyphloeini (Coleoptera: Curculionidae: Entiminae). Klapalekiana, 45, 1–97.
    Borovec, R. & Pelletier, J. (2010) Revision of the genus Foucartia Jacquelin du Val, 1855 (Coleoptera: Curculionidae: Entiminae: Sciaphilini). Acta Societatis Zoologicae Bohemicae, 73 (2009), 1–26.
    Caldara, R. (1974) Quatro curculionidi nuovi per la fauna Italiana. Atti della Societa Italiana di scienze naturali e del Museo Civico di Storia naturale in Milano, 115, 267–270.
    Cmoluch, Z. (1960) Brachysomus strawinskii n. sp. (Coleoptera, Curculionidae). Annales Universitatis Mariae Curie-Sklodowska Lublin-Polonia, Sectio C (5), 15, 111–115.
    Cmoluh, Z. (1962) Badania nad fauną ryjkowców (Coleoptera, Curculionidae) roślinnych zespołow kserotermicznych południowo-wschodniej części Wyżyny Lublanskiej. Annales Universitatis Mariae Curie-Sklodowska Lublin-Polonia, 17 (1), 1–75.
    Dalla Torre, K.W., van Emden, M. & van Emden, F. (1937) Curculionidae: Brachyderinae II. In: Schenkling, S. (Ed.), Coleopterorum Catalogus auspiciis et auxilio. Vol. 153. W. Junk, Berlin, pp. 133–196.
    Denosmaison, J.-C. (1999) Captures de Curculionidae peu communs en Ile-de-France (Coleoptera). Entomologiste, 55 (3), 130.
    Dieckmann, L. (1980) Beiträge zur Insektenfauna der DDR: Coleoptera—Curculionidae (Brachycerinae, Otiorhynchinae, Brachyderinae). Beiträge zur Entomologie, 30 (1), 145–310. https://doi.org/10.21248/contrib.entomol.30.1.145-310
    Doyen, J.T. (1966) The Skeletal Anatomy of Tenebrio molitor (Coleoptera: Tenebrionidae). Miscellaneous publications of the Entomological Society of America, 5 (3), 103–150.
    Endrődi, S. (1963) Ormányosbogarak III. Curculionidae III. Magyarország Állatvilága. Fauna Hungariae: Vol. 10. Akadémiai Kiadó, Budapest, 104 pp.
    Fabricius, J.C. (1792) Entomologia systematica emendata et atleta. Secundum classes, ordines, genera, species adjectis synonimis, locis, observationibus, descriptionibus. Vol. 1. Proft, Hafniae, XX + 538 pp.
    Fabricius, J.C. (1801) Systema eleutheratorum secundum ordines, genera, species: adiectis synonimis, locis, observationibus, descriptionibus. Vol. 2. Bibliopoli Academici Novi, Kiliae, 687 pp.
    Faust, J. (1889) Griechische Curculioniden. Deutsche Entomologische Zeitschrift, 1, 66–91.
    Fischer von Waldheim, G. (1829) Museum Historiae Naturalis Universitatis Caesareae Mosquensis. Pars II. Insecta. Typis Universitatis Caesareae, Mosquae, 147 pp.
    Formánek, R. (1905) Zur näheren Kenntnis der Gattung Brachysomus Stephens. Wiener Entomologische Zeitung, 24, 169–193.
    Formánek, R. (1906) Bemerkungen über bekannte Rüssler und Beschreibung einer neuen Art. Wiener Entomologische Zeitung, 25, 38–40.
    Formánek, R. (1907) Eine neue Art der Rüssler-Gattung Brachysomus aus Siebenbürgen. Wiener Entomologische Zeitung, 26, 23–24.
    Formánek, R. (1909) Zwei neue Curculioniden nebst Bemerkungen über vier bekannte. Wiener Entomologische Zeitung, 28, 29–31.
    Formánek, R. (1922) Zehn neue Curculioniden nebst Bemerkungen über bekannte. Wiener Entomologische Zeitung, 39, 73–84.
    Franz, H. (1947) Über Brachysomus microphthalmus Penecke. Annalen des Naturhistorischen Museums in Wien, 55, 260–262. [1944–1947]
    GBIF.org (2019) GBIF Brachysomus Occurrence Download. Available from: https://doi.org/10.15468/dl.kav3ny (accessed 15 November 2019)
    Germann, Ch., Braunert, C. &Kakiopoulos, G. (2016) Brachysomus (Hippomias) xanthianus sp. n. from Thrace, Greece (Coleoptera, Curculionidae). Mitteilungen der Schweizerischen Entomologischen Gesellschaft, 89, 177–182. https://doi.org/10.5281/zenodo.192629
    Germar, E.F. (1824) Insectorum species: novae aut minus cognitae, descriptionibus illustratae Volumen primum Coleoptera. Impensis J.C. Hendelii et Filii, Halae, 624 pp. https://doi.org/10.5962/bhl.title.130964
    Girón, J.C. & Franz, N.M. (2010) Revision, phylogeny and historical biogeography of the genus Apodrosus Marshall, 1922 (Coleoptera: Curculionidae: Entiminae). Insect Systematics & Evolution, 41 (2010), 339–414. https://doi.org/10.1163/187631210X538799
    Girón, J.C. & Chamorro, M.L. (2020) Variability and distribution of the golden-headed weevil Compsus auricephalus (Say) (Curculionidae: Entiminae: Eustylini). Biodiversity Data Journal, 8, e55474. https://doi.org/10.3897/BDJ.8.e55474
    Gosic, R. & Łętowski, J. (2003) Ryjkowkowate (Coleoptera, Curculionoidea: Rhinomaceridae, Attelabidae, Apionidae, Curculionidae) Białej Góry k. Tomaszowa Lubelskigo. Parki Narodowe i Rezervaty Pryrodny, 22 (2), 247–266.
    Hoffmann, A. (1942) Nouveaux Curculionides Paléarctiques. Revue Française d’Entomologie, 9 (1), 12–19.
    Hoffmann, A. (1950) Coléoptères Curculionides. Vol. 1. Faune de France. 52. Librairie de la Faculte des Sciences, Paris, 486 pp.
    Iablokoff-Khnzorian, S.M. (1958) Neue Käfer aus Sovjetarmenien (Brachidae et Curculionidae). Notulae Entomologicae, 38, 73–82.
    Kataev, B.M. (1996) On similar anomalies in the structure of male genitalia of some species of Ground Beetles of the genera Harpalus Latr. and Ophonus Dej. (Coleoptera, Carabidae) and their significance for reconstructing the phylogeny of Harpali genus group. Entomologicheskoye obozrenie, 75 (9), 151–166.
    Košťál, M. (1991a) Revision of the Brachysomus subnudus group (Coleoptera, Curculionidae). Elytron, 5, 103–110.
    Košťál, M. (1991b) Revision der Brachysomus hispidus-Gruppe (Insecta: Coleoptera: Curculionidae). Annalen Naturhistorisches Museum Wien, 92, 77–85.
    Košťál, M. (1992) Revision der Brachysomus transsylvanicus-Gruppe (Insecta, Coleoptera, Curculionidae: Brachyderinae). Entomologische Abhandlungen, 55 (3), 35–49.
    Lacordaire, T. (1863) Histoire naturelle des insectes. Genera des Coléoptères ou exposé méthodique et critique de tous les genres proposés jusqu’ici dans cet ordre d’insectes: vi. Librairie encyclopédique de Roret, Paris, 608 pp.
    Łętowski, J., Gosic, R., Czarnawski, W. & Budzyńska, E. (2003) Meteriały do znajomości ryjkowkowatych (Curculionoidea, Coleoptera) Kazimierskiego Parku Krajobrazowego. Parki Narodowe i Rezervaty Pryrodny, 22 (2), 227–245.
    Linnaeus, C. (1758) Systema Naturae per régna tria naturae, secundum classes, ordines, genera, species, cum caracteribus, differentiis, synonymis. Tomus I. Edition decima, reformata. Laurentii Salvii, Holmiae, vi + 824 + [1] pp.
    May, B.M. (1993) Larvae of Curculionoidea (Insecta: Coleoptera): a systematic overview. Fauna of New Zealand. No. 28. Manaaki Whenua Press, Lincoln, 226 pp.
    McNamara, J. (1991) Family Curculionidae snout beetles or weevils. In: Bousquet, Y. (Ed.), Checklist of Beetles of Canada and Alaska. Agriculture Canada, Ottawa, pp. 329–356.
    Morimoto, K., Kojima, H. & Miyakawa, S. (2006) The insects of Japan. Volume 3. Curculionoidea: general introduction and Curculionidae: Entiminae (part 1). Phyllobiini, Polydrusini and Cyphicerini (Coleoptera). Touka Shobo Co. Ltd., Fukuoka, iv + 406 pp.
    Müller, O.F. (1776) Zoologiae danicae prodromus, seu animalium Daniae et Norvegiae indigenarum characteres, nomina, et synonymya imprimis popularium. Typis Hallageriis, Havniae, 282 pp.
    O’Brien, C.W. (1979) Thricolepoides clavisetis, a new genus and species of Peritelini from the midwestern U.S. (Otiorhynchinae, Coleoptera). The Coleopterists Bulletin, 33 (1), 71–73.
    Oberprieler, R.G. (1988) Revision of the Tanyrhynchini of continental Africa (Coleoptera: Curculionidae). Entomology memoirs of the Department of Agriculture and Water Supply Republic of South Africa, 71, 1–50.
    Penecke, K. (1910) Ein neuer Brachysomus aus der Bukovina. Wiener Entomologische Zeitung, 29, 245–247.
    Penecke, K. (1914) Brachysomus mihoki sp. n. Wiener Entomologische Zeitung, 33, 36–38.
    Penecke, K. (1924) Zwei neue Rüsselkäfer aus Thessalien. Wiener Entomologische Zeitung, 41, 13–16.
    Penecke, K. (1939) Brachysomus microphthalmus sp. n. Koleopterologische Rundschau, 25, 10–12.
    Pešić, S. (1998) The fauna of weevils (Coleoptera: Attelabidae; Apionidae; Curculionidae; Rhynchophoridae) in Kragujevac Basin (Serbia). In: Colonnelli, E., Louw, S. & Osella, G. (Eds.), Proceedings of a Symposium XX International Congress of Entomology: “Taxonomy, ecology and distribution of Curculionoidea (Coleóptera: Polyphaga, Florence, Italy, 28 August, 1996. Atti del Museo Regionale di Scienze Naturali, Torino, pp. 163–182.
    Podlussány, A. (2001) Somogy megye ormányosalkatú bogarainak katalógusa (Coleoptera: Curculionoidea). Natura Somogyiensis, 1, 237–252.
    Redtenbacher, L. (1849) Fauna Austriaca. Die Käfer nach der analytischen Methode. 27. C. Gerold, Wien, 883 pp.
    Reitter, E. (1884) Einige neue Coleopteren aus Süd-Europa. Deutsche Entomologische Zeitschrift, 28, 255–259.
    Reitter, E. (1888) Coleopteren aus Circassien, gesammelt von Hans Leder im Jahre 1887. I Theil. Wiener Entomologische Zeitung, 7, 19–26.
    Reitter, E. (1891) Übersicht der mir bekannten Foucartia-Arten. Wiener Entomologische Zeitung, 10, 214–216.
    Reitter, E. (1895) Sechster Beitrag zur Coleopteren Fauna von Europa und der angrenzenden Ländern. Wiener Entomologische Zeitung, 14, 199–204.
    Reitter, E. (1912) Bestimmungs-Schlüssel für die Unterfamilien, Tribus und Gattungen der Curculionidae. Teil 19. Bestimmungs-Tabellen der europäischen Coleopteren 68. Verhandlungen des naturforschenden Vereines in Brünn, 51, 1–33.
    Reitter, E. (1916) Fauna Germanica. Die Käfer des Deutschen Reiches. Band V. K.G. Lutz, Stuttgart, 343 pp., 16 colour pls.
    Schoenherr, C.J. (1823) Curculionides. [Tabula synoptica familiae Curculionidum]. Isis von Oken, 2 (10), cols. 1132–1146.
    Schoenherr, C.J. (1840) s.n. In: Genera et species curculionidum, cum synonymia hujus familiae. Species novae aut hactenus minus cognitae, descriptionibus a Dom. Leonardo Gyllenhal, C. H. Boheman, et entomologis aliis illustratae. Tomus quintus. Pars secunda. Supplementum continens. Rore, Parisiis and Fleischer, Lipsiae, pp. v–viii + 465–970 + 4 (Corrigenda) pp. https://doi.org/10.5962/bhl.title.8952
    Schoenherr, C.J. (1842) Genera et species Curculionidum, cum synonymia hujus familiae. Species nova ant hactenus minus cognitae, descriptionibus a Dom Leonardo Gyllenhal, C.H. Boheman, et entomologis aliis illustratae. Tomus septimus. Pars secunda. Supplementum continens. Rore, Parisiis and Fleischer, Lipsiae, [4] + 461 pp. https://doi.org/10.5962/bhl.title.8952
    Schoenherr, C.J. (1845) Genera et species curculionidum, cum synonymia hujus familiae. Species novae aut hactenus minus cognitae, descriptionibus a Dom. Leonardo Gyllenhal, C. H. Boheman, et entomologis aliis illustratae. Tomus octavus. Pars secunda. Supplementum continens. Rore, Parisiis and Fleischer, Lipsiae, viii + 504 pp., 27 pls. https://doi.org/10.5962/bhl.title.8952
    Seidlitz, G. (1868) Die Otiorhynchiden sensu str. nach den morphologischen Verwandschaftverhältnissen ihres Hautscelets. Berliner entomologische Zeitschrift, 12, 1–153.
    Seidlitz, G. (1891) Fauna Transsylvanica. Die Käfer Siebenbürgens. Hartungsche Verlagsdruckerei, Königsberg, 915 pp.
    Smreczyński, S. (1966) Ryjkovce—Curculionidae. Pidrodziny Otiorhynchinae, Brachyderinae. Klucze do oznaczania owadów Polski. Coleoptera: 19(98b.). Państwowe Wydawnictwo Naukowe, Warszawa, 130 pp.
    Smreczyński, S. (1977) Neue Rüsselkäferarten aus Griechenland und aus der Türkei(Coleoptera, Curculionidae). Acta Zoologica Cracoviensia, 22 (10), 387–396.
    Smreczyński, S. (1981) Curculionidae, 8. Unterfamilie: Brachyderinae. In: Freude, H., Harde, K.W. & Lohse, G.A. (Eds.), Die Käfer Mitteleuropas. Bruchidae, Anthribidae, Scolytidae, Platypodidae, Curculionidae. Vol. 10. Goecke & Evers, Krefeld, pp. 240–273.
    Solari, F. (1948) Tre nuove specie e quattro nuovi generi della sottofamiglia Brachyderinae e contributo alla più precisa definizione della stessa (Col. Curc.). Memorie della Società Entomologica Italiana, 27, 23–34.
    Stierlin, G. (1884) Bestimmungs-Tabellen der europäischer Rüsselkäfer. Mittheilungen der Schweizerischen entomologischen Gesellschaft, 7 (2), 57–146.
    Stierlin, G. (1892) Beschreibung einiger neuen europäischer Rüsselkäfer. Mittheilungen der Schweizerischen entomologischen Gesellschaft, 8 (9), 367–366. [1888–1893]
    Stierlin, G. (1893) Drei neue Rüsselkäfer-Arten. Mittheilungen der Schweizerischen entomologischen Gesellschaft, 9, 56–57.
    Stierlin, G. (1899) Beschreibung eniger neuer europäischer Rüsselkäfer. Mittheilungen der Schweizerischen entomologischen Gesellschaft, 10 (5), 197–204.
    Szénasi, V. & Németh T. (2019) Description of a new species of the subgenus Hippomias Yunakov, 2006 in the genus Brachysomus Schoenherr, 1823 (Coleoptera: Curculionidae: Entiminae) from Lebanon. Zootaxa, 4658 (2), 389–395. https://doi.org/10.11646/zootaxa.4658.2.13
    Thompson, R.T. (1992) Observations on the morphology and classification of weevils (Coleoptera, Curculionoidea) with a key to major groups. Journal of Natural History, 26, 835–891. https://doi.org/10.1080/00222939200770511
    UkrBIN (2019) Genus Brachysomus Schoenherr, 1823. Dataset. Ukrainian Biodiversity Information Network. Available from: http://ukrbin.com/index.php?category=25091 (asccessed 15 November 2019)
    Wanat, M. (2007) Alignment and homology of male terminalia in Curculionoidea and other Coleoptera. Invertebrate Systematics, 21, 147–171. https://doi.org/10.1071/IS05055
    Wanat, M. & Mazur, M. (2005) A new species of Brachysomus from Poland (Coleoptera: Curculionidae: Entiminae). Genus, 16 (2), 291–298.
    Yunakov, N.N. (1998) Synonymy and notes of morphology and distribution of some less known species of the genus Brachysomus Schönherr (Coleoptera, Curculionidae). Kharkov Entomological Society Gazette, 6 (2), 37–40.
    Yunakov, N.N. (1999) Contribution to the knowledge of broad-nosed weevils (Coleoptera: Curculionidae, Entiminae) of Ukraine. Kharkov Entomological Society Gazette, 7 (1), 9–16.
    Yunakov, N.N. (2005) A revision of the weevil genus Amicromias Rtt. (Coleoptera, Curculionidae, Entiminae). Entomologicheskoye obozrenie, 84 (1), 143–158.
    Yunakov, N.N. (2006) Contribution to a revision of the genus Brachysomus Schoenh. (Coleoptera: Curculionidae, Entiminae). Description of new taxa. Proceedings of the Russian Entomological Society, 77, 294–329.
    Yunakov, N.N. (2012) Afrotropical weevils of the Cadoderus Marshall generic complex (Coleoptera: Curculionidae: Entiminae: Embrithini). Part 1. Genera: Cadoderus Marshall, 1926; Sphrigodellus Marshall, 1942 and Oncophyes Marshall, 1942. Zootaxa, 3535 (1), 1–104. https://doi.org/10.11646/zootaxa.3535.1.1
    Yunakov, N.N. & Germann, C. (2012) Brachysomus (Hippomias) samos sp. n. from Greece—first island endemism in the genus (Coleoptera: Curculionidae: Entiminae). Zootaxa, 3408 (1), 59–68. https://doi.org/10.11646/zootaxa.3408.1.5
    Yunakov, N.N. & Nadein, K.S. (2006) New taxa of geophilic Entiminae (Coleoptera: Curculionidae) from the Balkan Peninsula, Caucasus and Central Asia. Entomologische Abhandlungen, 63, 77–98.
    Zwölfer, H. (1975) Rüsselkäfer und ihre Umwelt—ein Kapitel Ökologie. Stuttgarter Beiträge zur Naturkunde, 3, 19–31.

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