Skip to main content Skip to main navigation menu Skip to site footer
Published: 2022-08-05

The family Melicharidae (Acari, Mesostigmata) in Slovakia, with description of new species, annotated faunal synopsis and identification keys of species from Europe

Institute of Zoology, Slovak Academy of Sciences, Dúbravská cesta 9, 845 06 Bratislava, Slova­kia
Acari Mites systematics identification keys morphology taxonomy fauna


The mites of the family Melicharidae (Acari: Mesostigmata) collected in Slovakia are reviewed. Sixty-one species belonging to five genera were recorded, of which thirty-three are new to science and fifteen are new to the Slovak fauna. All species are revised and rediagnosed, and an extensive set of photomicrographs (more than 1,300 images arranged in 337 Plates) is provided for their identification. Information on habitat and regional distribution is provided for each species. Proctolaelaps (fifty-three species) is the most species-rich and morphologically heterogeneous genus in the family.

                The following thirty-three new species are proposed and described: Mycomelichares triplacis sp. nov., Proctolaelaps appendiculatus sp. nov., Proctolaelaps bivalvifer sp. nov., Proctolaelaps bombi sp. nov., Proctolaelaps bucinator sp. nov., Proctolaelaps calycinus sp. nov., Proctolaelaps carnifex sp. nov., Proctolaelaps conifericola sp. nov., Proctolaelaps coralifer sp. nov., Proctolaelaps cryptopteryx sp. nov., Proctolaelaps dendroctonoides sp. nov., Proctolaelaps dorci sp. nov., Proctolaelaps dorcisimilis sp. nov., Proctolaelaps fagi sp. nov., Proctolaelaps falcarius sp. nov., Proctolaelaps fragilis sp. nov., Proctolaelaps fusifer sp. nov., Proctolaelaps hemisphaericus sp. nov., Proctolaelaps laeviusculus sp. nov., Proctolaelaps myrmecophilus sp. nov., Proctolaelaps orolaelapoides sp. nov., Proctolaelaps oxyodon sp. nov., Proctolaelaps pcolai sp. nov., Proctolaelaps pelticola sp. nov., Proctolaelaps populi sp. nov., Proctolaelaps quadridens sp. nov., Proctolaelaps sacculiger sp. nov., Proctolaelaps sagittarius sp. nov., Proctolaelaps serpentarius sp. nov., Proctolaelaps spanius sp. nov., Proctolaelaps speluncarum sp. nov., Proctolaelaps tubisaccus sp. nov., and Proctolaelaps tulipifer sp. nov.

                Nineteen new species are recognized in the present work, which were previously thought to belong to a single species, Proctolaelaps pygmaeus (J. Müller, 1859). The species boundaries of these species are often only weakly determined by some external morphological characters, but well definable and recognizable by the morphology of the internal structures of the sperm access system in the female (the same is true for the species related to Proctolaelaps hystrix).

                Proctolaelaps bulbicolus (Oudemans, 1929), Proctolaelaps convivus (Berlese, 1916), Proctolaelaps fissuratus (Berlese, 1916), Proctolaelaps laevisternus (Berlese, 1916), Proctolaelaps sopalit (Samšiňák, 1958), Proctolaelaps vulgaris (Pinchuk, 1972), and Proctolaelaps ovatus (Ma, Ning & Wei, 2003), previously suspected to be junior synonyms of Proctolaelaps pygmaeus (J. Müller, 1859), are here considered as unrecognizable species, relegated to the species incertae sedis, or species related within the invalid genus Proctofissus Karg, 1979.

                The very broad concept of the genus Proctolaelaps does not reflect the phylogenetic relationships within the apparently morphologically heterogeneous species, and it needs a thorough revision. As documented in present work, the genus includes species with different types of sperm access system: for example, species in the hystrix group possess the “laelapoid-type” of the sperm access system, whereas species related to Proctolaelaps pygmaeus (here placed in the pygmaeus group) have a highly modified sperm access system derived from the “phytoseioid-type”, with two separate parts each associated with coxa III and clearly not connected by a medial structure called the “sacculus foemineus” (a specific component of the laelapoid-type sperm access system).

                The taxonomy includes four new specific synonyms as follows: Proctolaelaps arctorotundus Nikolsky, 1984 = Proctolaelaps epuraeae Hirschmann, 1962, Proctolaelaps pruni Karg, 1988 = Proctolaelaps eccoptogasteris (Vitzthum, 1923), Proctolaelaps pseudofiseri Nikolsky, 1984 = Proctolaelaps fiseri Samšiňák, 1960, and Proctolaelaps ventrianalis Karg, 1971 = Proctolaelaps intermedius Athias-Henriot, 1959. Melichares biebrzae Gwiazdowicz, 2005 is newly placed in Proctolaelaps Berlese, 1923.

                The genus Orolaelaps De Leon, 1963, previously known from North and South America, is reported from Europe for the first time, introduced by an invasive cosmopolitan species Stelidota geminata (Say), the strawberry sap beetle of the family Nitidulidae.

                The following species are reported from Slovakia for the first time, namely Orolaelaps quisqualis De Leon, 1963, Proctolaelaps bickleyi (Bram, 1956), Proctolaelaps biebrzae (Gwiazdowicz, 2005), Proctolaelaps cossi (Dugès, 1834), Proctolaelaps cossoides Faraji, Sakenin-Chelav & Karg, 2007, Proctolaelaps hystricoides Lindquist & Hunter, 1965, Proctolaelaps izabelae Faraji, 2011, Proctolaelaps jueradeus (Schweizer, 1949), Proctolaelaps longanalis (Westerboer, 1963), Proctolaelaps rotundus Hirschmann, 1962, and Proctolaelaps xyloteri Samšiňák, 1960. The records of Proctolaelaps bombophilus (Westerboer, 1963), Proctolaelaps epuraeae Hirschmann, 1962, Proctolaelaps pomorum (Oudemans, 1929), and Proctolaelaps scolyti Evans, 1958 were previously based on misidentified specimens and are now supported by new reliable findings [in contrast to Proctolaelaps hystrix (Vitzthum, 1923) and Proctolaelaps pini Hirschmann, 1963, which were not definitely confirmed by the present work in Slovakia].

                New dichotomous keys are given for the identification of sixty-eight species from seven genera found so far in Europe (Melichares Hering, 1838, Mucroseius Lindquist, 1962, Mycolaelaps Lindquist, 1995, Mycomelichares Mašán & Joharchi, 2021, Orolaelaps De Leon, 1963, Orthadenella Athias-Henriot, 1973, and Proctolaelaps Berlese, 1923).



  1. Abo-Shnaf, R.I. A. & Moraes, G.J. (2016) Proctolaelaps species (Acari: Mesostigmata: Melicharidae) from Egypt, with description of a new species and complementary descriptions of other five species. Zootaxa, 4162, 479–503.
    Ambros, M. (1984) Poznámky k výskytu roztočov (Acari: Mesostigmata) drobných zemných cicavcov (Insectivora, Rodentia) pohoria Tríbeč. Rosalia (Nitra), 1, 33–46. [in Slovak]
    Ambros, M. (1985) Ektoparazity drobných zemných cicavcov okresu Považská Bystrica (Biele Karpaty, Javorníky, Strážovské vrchy, Považské podolie). 2. Roztoče (Acari: Mesostigmata). In: Galvánek, J. & Gregor, J. (Eds), XIX. tábor ochrancov prírody, Manínska tiesňava 1983. Prehľad odborných výsledkov, Považská Bystrica, pp. 78–88. [in Slovak]
    Ambros, M. (1986a) Príspevok k poznaniu fauny roztočov (Acarina: Mesostigmata) drobných cicavcov Považského Inovca a ŠPR Chynoranský luh. In: Gregor, J. (Ed.), Zborník odborných prác III. Západoslovenského tábora ochrancov prírody. Zväzok II, Krajský ústav štátnej pamiatkovej starostlivosti a ochrany prírody, Bratislava, pp. 59–71. [in Slovak]
    Ambros, M. (1986b) Roztoče (Acari: Mesostigmata) – ektoparazity piskora vrchovského (Sorex alpinus Schinz, 1837) z územia Západných Karpát. Biologia (Bratislava), 41, 185–192. [in Slovak]
    Ambros, M. (1987a) Ektoparazitofauna drobných zemných cicavcov (Insectivora, Rodentia) Pohronského Inovca (Západné Karpaty). 2. Roztoče (Mesostigmata). Rosalia (Nitra), 4, 231–244. [in Slovak]
    Ambros, M. (1987b) Poznámky k faune roztočov (Acari: Mesostigmata) hmyzožravcov a hlodavcov okresu Lučenec (Revúcka vrchovina, Cerová vrchovina). In: Ambros, M. (Ed.), XXII. Tábor ochrancov prírody. Prehľad odborných výsledkov, p. 127–133. [in Slovak]
    Ambros, M. (1988a) Fauna roztočov (Acari: Mesostigmata) drobných zemných cicavcov (Insectivora, Rodentia) južného Slovenska. In: Ambros, M. (Ed.), Zborník odborných prác V. Západoslovenského tábora ochrancov prírody, Bratislava, p. 51–62. [in Slovak]
    Ambros, M. (1988b) Ektoparazity drobných zemných cicavcov (Insectivora, Rodentia) Chránenej krajinnej oblasti Štiavnické vrchy. 2. Roztoče (Acarina: Mesostigmata). Ochrana Prírody, 9, 151–167. [in Slovak]
    Ambros, M. (1989a) Doplnky k faune roztočov (Acari: Mesostigmata) drobných cicavcov z niektorých lokalít Malých Karpát, Borskej nížiny a Chvojnickej pahorkatiny. In: Ambros, M. (Ed.), XXIV. Tábor ochrancov prírody. Prehľad odborných výsledkov (Lúky pod Korlátkom, Rozbehy, 9.–17. júla 1988), Okresný národný výbor – odbor kultúry a Slovenský zväz ochrancov prírody a krajiny, Ústredný výbor, Bratislava – Senica, p. 213–224. [in Slovak]
    Ambros, M. (1989b) Ektoparazity drobných zemných cicavcov (Insectivora, Rodentia) ŠPR Chabenec v Nízkych Tatrách. 2. Roztoče (Acari: Mesostigmata). In: Ambros, M. (Ed.), XXV. TOP (Tále 1989), Prehľad odborných výsledkov, p. 129–138. [in Slovak]
    Ambros, M. (1990) Fauna roztočov (Acari: Mesostigmata) z drobných cicavcov Chránenej krajinnej oblasti Ponitrie. 1. časť Tríbeč. Rosalia (Nitra), 6, 229–252. [in Slovak]
    Ambros, M. (1991) Fauna roztočov (Acari: Mesostigmata) z drobných cicavcov Chránenej krajinnej oblasti Ponitrie. 2. časť: Vtáčnik. Rosalia (Nitra), 7, 153–167. [in Slovak]
    Ambros, M. (1993) Ektoparazitické článkonožce – roztoče (Acari: Mesostigmata) drobných zemných cicavcov Chránenej krajinnej oblasti Slovenský kras. 2. Jelšavský kras, Silická planina a Turnianska kotlina. Ochrana prírody – Naturae Tutela, 2, 221–228. [in Slovak]
    Ambros, M. & Kalúz, S. (1985) Príspevok k poznaniu ektoparazitov drobných cicavcov (Insectivora, Rodentia) z územia Bratislavy so zameraním na parazitické roztoče čeľade Dermanyssidae (Acari: Mesostigmata). In: Zborník referátov z konferencie “Zoocenózy urbánnych a suburbánnych celkov so zvláštnym akcentom na podmienky Bratislavy”, Slovenská zoologická spoločnosť pri SAV a Zoologický ústav Univerzity Komenského, Bratislava, p. 198–202. [in Slovak]
    Ambros, M. & Kalúz, S. (1987) Roztoče (Acari: Mesostigmata) – ektoparazity drobných cicavcov (Insectivora, Rodentia) z územia Bratislavy. Acta Rerum Naturalium Musei Nationalis Slovaci, 33, 119–128. [in Slovak]
    Ambros, M. & Stanko, M. (1993) Fauna roztočov (Acari: Mesostigmata) drobných cicavcov (Insectivora, Rodentia) Spiša. In: Nullasz, L. (Ed.), XVI. Východoslovenský tábor ochrancov prírody (Červený Kláštor, 25. 7.–1. 8. 1992). Prehľad odborných výsledkov, Okresný výbor Slovenského zväzu ochrancov prírody a krajiny, Poprad a Správa Pieninského národného parku, Červený Kláštor, p. 60–66. [in Slovak]
    Ambros, M., Krištofík, J. & Šustek, Z. (1992) The mites (Acari, Mesostigmata) in the birds nests’ in Slovakia. Biologia (Bratislava), 47, 369–381.
    Ambros, M., Dudich, A., Kováčik, J. & Štollmann, A. (1982) Teriologický a parazitologický výskum na 18. Tábore ­­­ochrancov prírody. In: Vestenický, K. & Čuboňová, K. (Eds), XVIII. Tábor ochrancov prírody 1982. Prehľad odborných výsledkov, Tlačiarne SNP, Martin, p. 52–69. [in Slovak]
    Andreev, E.A. (1985) Mites associated with caterpillar galleries of the common goat moth (Cossus cossus L.), an important trunk pest in southern Turkmenistan. Biologicheskie Nauki, 1985 (8), 34–38.
    Athias-Henriot, C. (1959) Phytoseiidae et Aceosejidae (Acarina, Gamasina) d’Algerie. III Contribution aux Aceosejinae. Bulletin de la Société d’Historie Naturalle de L’Afrique du Nord, 50, 158–195.
    Athias-Henriot, C. (1961) Mesostigmates (Urop. Excl.) édaphiques meditérranéens (Acaromorpha, Anactinotrichida) (collect. Prof. H. Franz et C. Athias-Henriot). Pemière Série. Acarologia, 3, 381–509.
    Athias-Henriot, C. (1973) Observations sur les genres Neojordensia Evans et Orthadenella n.g. en Europe Occidentale (Gamasides, Dermanyssina, Ascidae). Acarologia, 15, 18–32.
    Bai, X.-L. & Gu, Y.-M. (1994) A new species of the genus Paraproctolaelaps from China (Acari: Ascidae). Acta Zootaxonomica Sinica, 19, 75–77. [in Chinese]
    Bai, X.-L., Yin, S.-G. & Gu, Y.-M. (1993) A new species and a new record of the genus Proctolaelaps from China (Acari: Blattisocidae). Acta Zootaxonomica Sinica, 18, 174–176. [in Chinese]
    Bai, X.-L., Yan, L.-M., Wei, H. & Qi, R.-J. (2008) Two new gamasid species from Ningxia, China (Acari, Gamasina). Acta Zootaxonomica Sinica, 33, 756–760. [in Chinese]
    Bai, X.-L., Zang, T., Yan, Y., & Yang, P. (2015) A new species of the genus Proctolaelaps Berlese from Ningxia, China (Acari, Ascidae). Acta Arachnologica Sinica, 24, 27–30. [in Chinese]
    Baker, E.W. & Yunker, C.E. (1964) New blattisociid mites (Acarina: Mesostigmata) recovered from neotropical flowers and hummingbirds’ nares. Annals of the Entomological Society of America, 57, 103–126.
    Berlese, A. (1916) Centuria prima di Acari nuovi. Redia, 12, 19–67.
    Berlese, A. (1918) Centuria quarta di Acari nuovi. Redia, 13, 115–192.
    Berlese, A. (1923) Centuria sesta di Acari nuovi. Redia, 15, 237–262.
    Bhattacharyya, A.K., Sanyal, A.K. & Bhattacharyya, T. (1997) Three new ascid mites (Acarina: Mesostigmata: Ascidae) from a nest of five-striped squirrel Funambulus pennanti Wroughton in West Bengal, India. Acarina, 5, 37–43.
    Bibin, A.R. (2017) Invasive sap beetles Epuraea ocularis and Stelidota geminata (Coleoptera, Nitidulidae) in the Russian Black Sea region. Russian Journal of Biological Invasions, 8, 301–302.
    Braig, H.R. & Perotti, M.A. (2009) Carcasses and mites. Experimental & Applied Acarology, 49, 45–84.
    Bram, R.A. (1956) A new predatory mite from insect culture. Proceedings of the Entomological Society of Washington, 58, 292–294.
    Bregetova, N.G. (1977) Family Aceosejidae Baker and Wharton, 1952 (sensu Evans, 1958). In: Ghilyarov, M.S. & Bregetova, N.G. (Eds.), Key to the Soil-Inhabiting Mites. Mesostigmata. Nauka, Leningrad, pp. 169–226. [in Russian]
    Connell, W.A. (1980) Stelidota geminata (Say) infestations of strawberries (Coleoptera: Nitidulidae). Entomological News, 91 (2), 55–56.
    Chant, D.A. (1963) The subfamily Blattisocinae Garman (Aceosejinae Evans) (Acarina: Blattisocidae Garman) (Aceosejidae Baker and Wharton) in North America, with descriptions of a new species. Canadian Journal of Zoology, 41, 243–305.
    Christian, A. & Karg, W. (2006) The predatory mite genus Lasioseius Berlese, 1916 (Acari, Gamasina). Abhandlungen und Berichte des Naturkundemuseums Görlitz, 77, 99–250.
    Costa, M. (1963) The mesostigmatic mites associated with Copris hispanus (L.) (Coleoptera, Scarabaeidae) in Israel. Journal of the Linnean Society of London, Zoology, 45, 25–45.
    Cyprich, D., Fenďa, P., Krumpál, M., Lukáš, J. & Ambruš, B. (2000) Hniezdna fauna zebričky červenozobej (Taeniopygiaguttata castanotis). Tichodroma, 13, 189–200. [in Slovak]
    Davydova, M.S. (1988) New species of gamasid mites associated with bumble bees. Novye i Maloizvestnye Vidy Fauny Sibiri, 20, 18–32. [in Russian]
    De Leon, D. (1963) New genus and twelve new species of mites from Mexico and southeast United States (Acarina: Blattisociidae). The Florida Entomologist, 46, 197–207.
    Dudich, A. (1988) Náčrt spoločenstiev mikromammálií a ich ektoparazitocenóz prírodných regiónov okresu Vranov nad Topľou. In: Ambros, M. (Ed.), XI. Východoslovenský tábor ochrancov prírody (Hermanovce 25. 7.–1. 8. 1987). Zborník odborných výsledkov, Krajský ústav štátnej pamiatkovej starostlivosti a ochrany prírody v Prešove, Okresný národný výbor, odbor kultúry vo Vranove nad Topľou, Prešov – Vranov nad Topľou, pp. 131–146. [in Slovak]
    Dudich, A., Ambros, M., Štollmann, A. & Kováčik, J. (1987) Ektoparazitické článkonožce drobných zemných cicavcov Chránenej krajinnej oblasti Slovenský kras. I. Plešivecká planina a Koniar. Ochrana Prírody, 8, 99–122. [in Slovak]
    Dugès, A. (1834) Recherches sur l’order des Acarines en général et la famille des Trombidiés en particular. I. Annales des Sciences Naturelles, Zoologie, 2, 5–46.
    El-Banhawy, E.M. & Nasr, A.K. (1986) Incidence of mites in bee hives in the state of New York with descriptions of two new species. Bulletin of the Zoological Society of Egypt, 34, 76–82.
    EPPO – European and Mediterranean Plant Protection Organization (2010) Stelidota geminata: another invasive species in Europe? EPPO Reporting Service – Pests & Diseases, Paris, 187, 10–11.
    Evans, G.O. (1954) Some new and rare species of Acarina. Proceedings of the Zoological Society of London, 123, 793–811.
    Evans, G.O. (1957) An introduction to the British Mesostigmata (Acarina) with keys to families and genera. Journal of the Linnean Society, Zoology, 43, 203–259.
    Evans, G.O. (1958) A revision of the British Aceosejinae (Acarina: Mesostigmata). Proceedings of the Zoological Society of London, 131, 177–229.
    Evans, G.O. (1963) Observations on the chaetotaxy of the legs in the free-living Gamasina (Acari: Mesostigmata). Bulletin of the British Museum (Natural History), Zoology, 10, 277–303.
    Evans, G.O. & Till, W.M. (1979) Mesostigmatic mites of Britain and Ireland (Chelicerata: Acari-Parasitiformes). An introduction to their external morphology and classification. Transactions of the Zoological Society of London, 35, 139–270.
    Fain, A., Hyland, K.E. & Aitken, T.H.G (1977) Flower mites of the family Ascidae phoretic in nasal cavities of birds (Acarina: Mesostigmata). Acta Zoologica et Pathologica Antverpiensia, 69, 99–154.
    Faraji, F. (2011) A new species of Proctolaelaps Berlese (Acari: Melicharidae) associated with the nest of European starling Sturnus vulgaris. Revista Ibérica de Acarologia, 19, 23–26.
    Faraji, F., Sakenin-Chelav, H. & Karg, W. (2007) A new species of Proctolaelaps Berlese from Iran (Acari: Ameroseiidae). Abhandlungen und Berichte des Naturkundemuseums Görlitz, 78, 107–112.
    Fenďa, P. (2010) Mites (Mesostigmata) inhabiting bird nests in Slovakia (Western Carpathians). In: Sabelis, M.W. & Bruin, J. (Eds), Trends in Acarology. Proceedings of the 12th International Congress. 1st Edition. Springer Science+Business Media B.V., Dordrecht, Heidelberg, London, New York, p. 199–205.
    Fenďa, P. & Kalúz, S. (2009) Distribution and ecology of the ascid mites in Slovakia (Acari, Mesostigmata, Ascidae). In: Tajovský, K., Schlaghamerský, J. & Pižl, V. (Eds), Contributions to Soil Zoology in Central Europe III. Institute of Soil Biology Academy of Sciences of the Czech Republic, České Budějovice, pp. 33–40.
    Fenďa, P. & Košel, V. (2000) Roztoče (Acarina: Mesostigmata) jaskýň Slovenského raja. In: Mock, A., Kováč, Ľ. & Fulín, M. (Eds), Fauna jaskýň (Cave Fauna), Východoslovenské múzeum, Košice, pp. 21–30. [in Slovak]
    Fenďa, P. & Košel, V. (2004) Mites (Acarina: Mesostigmata) inhabiting caves of the Belianske Tatry Mountains (Northern Slovakia). Biologia (Bratislava), 59, 35–40.
    Fenďa, P. & Košel, V. (2005) Roztoče (Acarina, Mesostigmata) centrálnych Západných Karpát I. Belianske Tatry, jaskynná fauna. Entomofauna Carpathica, 17, 40–47. [in Slovak]
    Fenďa, P., Krumpál, M. & Cyprich, D. (1998) The soil fauna in the birds’ nests in Slovakia. In: Pižl, V. & Tajovský, K. (Eds), Soil Zoological Problems in Central Europe, ISB AS CR, České Budějovice, pp. 23–30.
    Fenďa, P. & Lengyel, J. (2007) Roztoče (Acarina, Mesostigmata) v hniezdach orliaka morského (Haliaeetus albicilla) na Slovensku. Entomofauna Carpathica, 19, 48–50 [in Slovak]
    Fenďa, P. & Lukáš, J. (2014) First records of mites (Acari: Mesostigmata) from Slovakia. Folia Faunistica Slovaca, 19, 171–175.
    Fenďa, P. & Mašán, P. (2003) Roztoče – Acari (Parasitiformes, ex.Uropodina). In: Mašán, P. & Svatoň, J. (Eds), Pavúkovce Národného parku Poloniny – Arachnids of the Poloniny National Park (Arachnida: Araneae, Pseudoscorpiones, Opiliones, Acari – Parasitiformes). Štátna ochrana prírody SR, Banská Bystrica a Správa Národného parku Poloniny, Snina, pp. 143–205. [in Slovak]
    Fenďa, P. & Mašán, P. (2009) Roztoče – Acari (Parasitiformes, ex. Uropodina). In: Mašán, P. & Mihál, I. (Eds), Pavúkovce Cerovej vrchoviny – Arachnids of the Cerová vrchovina (Araneae, Opiliones, Pseudoscorpiones, Acari). ŠOP SR Banská Bystrica, Správa CHKO Cerová vrchovina Rimavská Sobota, ÚZ SAV Bratislava, ÚEL SAV Zvolen, pp. 153–208. [in Slovak]
    Fenďa, P. & Schniererová, E. (2005) Mites (Acarina, Gamasida) in littoral zone of Jakubov fishponds (Slovakia). In: Tajovský, K., Schlaghamerský, J. & Pižl, V. (Eds), Contributions to Soil Zoology in Central Europe I. ISB AS CR, České Budějovice, pp. 9–14.
    Fenďa, P. & Schniererová, E. (2010) Mites (Acari: Mesostigmata) of the birds’ nests on the water in southwestern Slovakia. Folia Faunistica Slovaca, 15, 55–60.
    Fenďa, P., Kucman, P., Bačíková, S., Országhová, Z., Puchala, P., Sobeková, K., Jánošková, V. & Melišková, M. (2011) Roztoče (Acari, Mesostigmata) v hniezdach vrabca poľného (Passer montanus) v PR Šúr (JZ Slovensko). Folia Faunistica Slovaca, 16, 35–42. [in Slovak]
    Franz, H. (1943) Die Landtierwelt der mittleren hohen Tauern. Denkschriften der Akademie der Wissenschaften in Wien, Mathematisch-Naturwissenschaftliche Klasse, 107, 1–552, 14 plates, 11 maps.
    Gočaltovská, E. (1963) Výskum akarofauny (Gamasoidea) v rezervácii Jurského Šúru. PhD Thesis, Faculty of Natural Sciences, Comenius University, Bratislava, 75 pp. [in Slovak]
    Gwiazdowicz, D.J. (2005) Melichares biebrzae sp. n. (Acari, Ascidae), a new ascid mite from Poland. Biologia, Bratislava, 60, 479–481.
    Gwiazdowicz, D.J. (2007) Ascid mites (Acari, Mesostigmata) from Selected Forest Ecosystems and Microhabitats in Poland. Wydawnictwo Akademii Rolniczej Im. Augusta Cieszkowskiego, Poznań, 248 pp.
    Gwiazdowicz, D.J. & Rakowski, R. (2010) Redescription of Proctolaelaps parvanalis (Thor, 1930) (Acari: Ascidae) from Spitsbergen. Entomologica Fennica, 20, 281–286.
    Gwiazdowicz, D.J., Blaszak, C. & Ehrnsberger, R. (2008) Die Milben in der Zoologischen Staatssammlung München. Teil 11. Gattungen: Arctoseius Thor, 1930, Cheiroseius Berlese, 1916, Melichares Hering, 1839, Proctolaelaps Berlese, 1923. Spixiana, 31, 29–33.
    Gwiazdowicz, D.J., Marchenko, I.I. & Teodorowicz, E. (2014) Description of Orthadenella coulsoni sp. nov. (Acari, Mesostigmata, Melicharidae) from Siberia with a key to the females of Orthadenella. Journal of Natural History, 49, 1659–1671.
    Halbert, J.N. (1920) The Acarina of the seashore. Royal Irish Academy, London, Section B, Nº 7, 35, 106–152.
    Halliday, R.B. (2001) Mesostigmatid mite fauna of Jenolan Caves, New South Wales (Acari: Mesostigmata). Australian Journal of Entomology, 40, 299–311.
    Halliday, R.B., Walter, D.E. & Lindquist, E.E. (1998) Revision of the Australian Ascidae (Acarina: Mesostigmata). Invertebrate Taxonomy, 12, 1–54.
    Haragsim, O., Samšiňák, K. & Vobrázková, E. (1978) The mites inhabiting the bee-hives in ČSR. Zeitschrift für Angewandte Entomologie, 87, 52–67.
    Heliövaara, K., Peltonen, M., Mannerkoski, I. & Siitonen, J. (1998) Suomen kaarnakuoriaiset. (Coleoptera: Scolytidae). University of Helsinki. Department of Applied Zoology. Reports 25. Yliopistopaino, Helsinki, Finland, 91 pp.
    Hering, E. (1838) Die Krätzmilben der Thiere und einige verwandte Arten, nach eigenen Untersuchungen beschrieben. Nova Acta Physico-Medica Academiae Caesareae Leopoldino-Carolinae Naturae Curiosorum, 18, 573–624.
    Hirschmann, W. (1962) Gangsystematik der Parasitiformes. Teil 5. Gamasiden Rückenhaarbesttimmungstafeln von 260 Typhlodromus-Arten der Erde. Gänge, Chaetotaxie Porotaxie, Mundwerkzeuge von Typhlodromus und verwandten Gattungen von Proctolaelaps, Melichares, Lasioseius, Iphidozercon, Sejus, Rhodacarellus, Rhodacarus, Gamasellus, Veigaia, Macrocheles ivanovi. Erstversuch der Aufstellung eines Gangsystems der Gamasiden aufgrund der Gnathosoma-unterseite. Acarologie. Schriftenreihe für Verglenchende Milbenkunde, 5, 1–56.
    Ibrahim, G.A. & Taha, H.A. (1989) Proctolaelaps metwalyi, a new species from Egypt associated with stored products (Acari: Gamasida, Ascidae). Azhar Journal of Agricultural Research (Egypt), 10, 87–91.
    Ishikawa, K. (1968) Studies on the mesostigmatid mites associated with the insects in Japan (I). Reports of Research Matsuyama Shinonome Junior College, 3, 197–218.
    Jelínek, J. (2014) Coleoptera: Sphindidae, Kateretidae, Nitidulidae. Folia Heyrovskyana, Series B, 21, 1–29.
    Joharchi, O. & Tolstikov, A.V. (2022) A new species of Orolaelaps De Leon (Acari: Mesostigmata: Melicharidae) from Cuba. Acarina, 30, 79–88.
    Joharchi, O., Döker, I. & Khaustov, A.A. (2021) Rediscovery of two gamasid mites (Acari: Mesostigmata) associated with beetles in Western Siberia, Russia. International Journal of Acarology, 47, 327‒338.
    Kalúz, S. (1996) Pôdne roztoče (Acarina) a iné článkonožce (Arthropoda) v podmienkach zavlažovania. Entomofauna Carpathica, 8, 43–51. [in Slovak]
    Kalúz, S. (1998a) Pôdne roztoče (Acarina) Grečovho vrchu (Slovenský kras). Entomofauna Carpathica, 10, 73–79. [in Slovak]
    Kalúz, S. (1998b) Roztoče (Acarina) machov Národnej prírodnej rezervácie Rozsutec v Malej Fatre. In: Korňan, M. (Ed.), Výskum a ochrana Krivánskej Fatry. Správa Národného parku Malá Fatra, Varín, p. 62–65. [in Slovak]
    Kandil, M.M. (1979) Revision of the genus Neojordensia Evans, 1957 with description of a new species from Hungary (Acari: Phytoseioidea). Folia Entomologica Hungarica, 32, 15–19.
    Karg, W. (1965) Larvalsystematische und phylogenetische Untersuchung sowie Revision des Systems der Gamasina Leach, 1915 (Acarina, Parasitiformes). Mitteilungen aus dem Zoologischen Museum in Berlin, 41, 193–340.
    Karg, W. (1971) Acari (Acarina), Milben, Unterordnung Anactinochaeta (Parasitiformes). Die freilebenden Gamasina (Gamasides), Raubmilben. Die Tierwelt Deutschlands, 59, 1–475.
    Karg, W. (1976) To the knowledge of the superfamily Phytoseioidea Karg, 1965. Zoologische Jahrbücher Abteilung für Systematik, Ökologie und Geographie der Tiere, 103, 505–546.
    Karg, W. (1979) Zur Kenntnis der Milbengattungen Lasioseius Berlese, 1916, Proprioseiopsis Muma, 1961, Podocinum Berlese, 1882 und Proctolaelaps Berlese, 1923 (Acarina, Parasitiformes). Deutsche Entomologische Zeitschrift, Neue Folge, 26, 1–8.
    Karg, W. (1985) Die Raubmilbengattung Proctolaelaps Berlese, 1923. Zoologische Jahrbücher Abteilung für Systematik, Ökologie und Geographie der Tiere, 112, 185–206.
    Karg, W. (1988a) Zur Kenntnis der Gattung Proctolaelaps Berlese, 1923 (Acarina, Parasitiformes). Zoologische Jahrbücher Abteilung für Systematik, Ökologie und Geographie der Tiere, 115, 441–454.
    Karg, W. (1988b) Eine neue Raubmilbenart der Gattung Proctolaelaps Berlese, 1923 (Acarina, Parasitiformes) am Grossen Obstbaumsplintkäfer (Scolytus (Eccoptogaster) mali Bechest.). Archiv für Phytopathologie und Pflanzenschutz, 24, 515–517.
    Karg, W. (1993) Acari (Acarina), Milben Parasitiformes (Anactinochaeta) Cohors Gamasina Leach, Raubmilben. Die Tierwelt Deutschlands und der angrenzenden Meeresteile nach ihren Merkmalen und ihrer Lebensweise, 59. Teil, 2., überarbeitete Auflage, Gustav Fischer Verlag, Jena, Stuttgart, New York, 523 pp.
    Karg, W. (1994) Raubmilben der Ascidae Ascidae, Ameroseiidae, Rhodacaridae und Macrochelidae auf dem Galapagos Archipel (Acarina, Parasitiformes). Mitteilungen aus dem Zoologischen Museum in Berlin, 70, 113–131.
    Karg, W. (2005) Neue Erkenntnisse zur weit verbreiteten Milbenfamilie Ameroseiidae Evans (Acarina, Parasitiformes). Abhandlungen und Berichte des Naturkundemuseums Görlitz, 77, 57–75.
    Karg, W. (2006) The systematics of Parasitiformes, especially of Gamasina Leach (Acarina), with new species from Ecuador. Mitteilungen aus dem Museum für Naturkunde in Berlin, Zoologische Reihe, 82, 140–169.
    Karg, W. & Rodriguez, N. (1984) Zwei neue Raubmilbenarten der Gattung Proctolaelaps Berlese, 1923 (Acarina, Gamasina) aus der Republik Kuba. Archiv für Phytopathologie und Pflanzenschutz, 20, 357–359.
    Karg, W., Baker, R.A. & Jenkinson, L. (1995) Mite infestation in Drosophila cultures and description of a new mite species of the genus Proctolaelaps Berlese with notes on its control in the laboratory (Acarina: Parasitiformes). Deutsche Entomologische Zeitschrift, Neue Folge, 42, 187–190.
    Klimov, P.B., OConnor, B., Ochoa, R., Bauchan, G.R., Redford, A.J. & Scher, J. (2016) Bee Mite ID: Bee-Associated Mite Genera of the World. USDA APHIS Identification Technology Program (ITP), Fort Collins, CO. [date of access on September 10, 2021].
    Koçak, A.O. & Kemal, M. (2008) Nomenclatural notes on the genus group names of the order Acarina. Centre for Entomological Studies Ankara, Miscellaneous Works, 145, 1–6.
    Kolibáč, J. (2005) A review of the Trogositidae. Part 1: Morphology of the genera (Coleoptera, Cleroidea). Entomologica Basiliensia et Collectionis Frey, 27, 39–159.
    Kolibáč, J. (2013) Trogossitidae: A review of the beetle family, with a catalogue and keys. ZooKeys, 366, 1–194.
    Kováč, Ľ., Mock, A., Ľuptáčik, P., Hudec, I., Košel, V. & Fenďa, P. (2002) Článkonožce (Arthropoda) Belianskej jaskyne (Belianske Tatry). Aragonit, 7: 27–29. [in Slovak]
    Kováč, Ľ., Mock, A., Ľuptáčik, P., Košel, V., Fenďa, P., Svatoň, J. & Mašán, P. (2005) Terrestrial arthropods of the Domica Cave system and the Ardovská Cave (Slovak Karst) – principal microhabitats and diversity. In: Tajovský, K., Schlaghamerský, J. & Pižl, V. (Eds), Contributions to Soil Zoology in Central Europe I, ISB AS CR, České Budějovice, pp. 61–70.
    Kováč, Ľ., Mock, A., Ľuptáčik, P., Višňovská, Z. & Fenďa, P. (2006) Bezstavovce (Evertebrata) Dobšinskej ľadovej jaskyne (Slovenský raj). In: Bella, P. (Ed.), Výskum, využívanie a ochrana jaskýň. 5. Vedecká konferencia s medzinárodnou účasťou, Zborník referátov. Správa slovenských jaskýň, Liptovský Mikuláš, pp. 179–186. [in Slovak]
    Kováč, Ľ., Schnitzerová, E., Miklisová, D. & Mati, R. (1999) Gamasina communities (Acari, Parasitiformes) of arable soils with two different soil types. Pedobiologia, 43, 54–63.
    Krantz, G.W. (1962) Acari: free-living Mesostigmata. II. The family Aceosejidae. Parc National de La Garamba, 34, 3–29.
    Krištofík, J., Mašán, P. & Šustek, Z. (1996) Ectoparasites of bee-eater (Merops apiaster) and arthropods in its nests. Biologia (Bratislava), 51, 557–570.
    Krištofík, J., Mašán, P. & Šustek, Z. (2001) Mites (Acari), beetles (Coleoptera) and fleas (Siphonaptera) in the nests of great reed warbler (Acrocephalus arundinaceus) and reed warbler (A. scirpaceus). Biologia (Bratislava), 56, 525–536.
    Krištofík, J., Mašán, P., Šustek, Z. & Gajdoš, P. (1993) Arthropods in the nests of penduline tit (Remiz pendulinus). Biologia (Bratislava), 48, 493–505.
    Krištofík, J., Mašán, P., Šustek, Z. & Karaska, D. (2009) Arthropods in the nests of lesser spotted eagle (Aquila pomarina). Biologia (Bratislava), 64, 974–980.
    Krištofík, J., Mašán, P., Šustek, Z. & Kloubec, B. (2003) Arthropods (Pseudoscorpionida, Acari, Coleoptera, Siphonaptera) in nests of the tengmalm’s owl, Aegolius funereus. Biologia (Bratislava), 58, 231–240.
    Krištofík, J., Mašán, P., Šustek, Z. & Nuhličková, S. (2013) Arthropods (Acarina, Coleoptera, Siphonaptera) in nests of hoopoe (Upupa epops) in Central Europe. Biologia (Section Zoology), 68, 155–161.
    Krištofík, J., Šustek, Z. & Mašán, P. (2002) Arthropods (Pseudoscorpionida, Acari, Coleoptera, Siphonaptera) in the nests of red-backed shrike (Lanius collurio) and lesser grey shrike (Lanius minor). Biologia (Bratislava), 57, 603–613.
    Krumpál, M., Cyprich, D. & Fenďa, P. (1998) Predbežný prehľad fauny roztočov (Acarina) a bĺch (Siphonaptera) hniezd niektorých druhov vtákov (Aves) a cicavcov (Mammalia) v Malej Fatre. In: Korňan, M. (Ed.), Výskum a ochrana Krivánskej Fatry, Správa národného parku Malá Fatra, Varín, pp. 52–61. [in Slovak]
    Lindquist, E.E. (1962) Mucroseius monochami, a new genus and species of mite (Acarina: Blattisociidae) symbiotic with sawyer beetles. The Canadian Entomologist, 94, 972–980.
    Lindquist, E.E. (1964) A Revision of Mites of Subfamily Blattisociinae (Acarina: Blattisociidae) in America North of Mexico. Ph.D. Dissertation, University of California, Berkeley, 413 pp.
    Lindquist, E.E. (1994) Some observations on the chaetotaxy of the caudal body region of gamasine mites (Acari: Mesostigmata), with a modified notation for some ventrolateral body setae. Acarologia, 35, 323–326.
    Lindquist, E.E. (1995) Remarkable convergence between two taxa of ascid mites (Acari: Mesostigmata) adapted to living in pore tubes of bracket fungi in North America, with description of Mycolaelaps new genus. Canadian Journal of Zoology, 73, 104–128.
    Lindquist, E.E. & Evans, G.O. (1965) Taxonomic concepts in the Ascidae, with a modified setal nomenclature for the idiosoma of the Gamasina (Acarina: Mesostigmata). Memoirs of the Entomological Society of Canada, 47, 1–65.
    Lindquist, E.E. & Hunter, P.E. (1965) Some mites of the genus Proctolaelaps Berlese (Acarina: Blattisociidae) associated with forest insect pests. The Canadian Entomologist, 97, 15–32.
    Lindquist, E.E. & Moraza, M.L. (2008) A new genus of flower-dwelling melicharid mites (Acari: Mesostigmata: Ascoidea) phoretic on bats and insects in Costa Rica and Brazil. Zootaxa, 1685, 1–37.
    Lindquist, E.E. & Wu, K.W. (1991) Review of mites of the genus Mucroseius (Acari: Mesostigmata: Ascidae) associated with sawyer beetles (Cerambycidae: Monochamus and Mecynippus) and pine wood nematodes [Aphelenchoididae: Bursaphelenchus xylophilus (Steiner and Buhrer) Nickle], with descriptions of six new species from Japan and North America, and notes on their previous misidentification. The Canadian Entomologist, 123, 875–927.
    Lindquist, E.E., Krantz, G.W. & Walter, D.E. (2009) Order Mesostigmata. In: Krantz, G.W. & Walter, D.E. (Eds.), A Manual of Acarology. 3rd Edition. Texas Tech University Press, Lubbock, Texas, pp. 1–807.
    Lindquist, E.E., OConnor, B.M., Shaw, M.D. & Sidorchuk, E.A. (2020) Review of the genera Berlesia Canestrini, 1884, and Katydiseius Fain & Lukoschus, 1983, the subfamily Katydiseiinae Fain & Lukoschus, 1983, and their family group relationships (Acari: Mesostigmata: Gamasina), with description of three new species parasitic on gryllacridid crickets (Orthoptera). Zootaxa, 4857, 5–70.
    Loughner, R.L., Loeb, G.M., Schloemann, S. & Demchak, K. (2008) Evaluation of cultural practices for potential to control strawberry sap beetle (Coleoptera: Nitidulidae). Journal of Economic Entomology, 101, 850–858.
    Lyonet, P. (1760) Traiteì anatomique de la chenille, qui ronge le bois de saule. Pierre De Hondt, Marc Michel Rey, Th. Becket & P. A. De Hondt, La Haye, Amsterdam et Londres, 665 pp.
    Lyonet, P. (1829) Anatomie des différentes espèces d´insects. Memoires du Muséum d´Histoire Naturelle, 18, 233–312, 7 plates.
    Ma, L.-M. (1996) Descriptions of Cheiroseius wuwenzheni sp. nov. and Proctolaelaps longichelicerae sp. nov. (Acari: Aceosejidae). Acta Arachnologica Sinica, 5, 36–41. [in Chinese]
    Ma, L.-M. (2006) New synonyms of gamasid mites (Acari: Mesostigmata). Acta Arachnologica Sinica, 15, 23–26. [in Chinese]
    Ma, L.-M., Zhang, A.H. & Li, Y.R. (2003) Two new species of the genus Hypoaspis and a new species of the genus Melichares associated with insects (Acari: Gamasina: Laelapidae and Aceosejidae). Acta Arachnologica Sinica, 12, 72–78. [in Chinese]
    Ma, Y., Ning, G. & Wei, Y.-W. (2003) A new species of the genus Hypoaspis from Qinghai, China (Acari, Laelapidae). Acta Zootaxonomica Sinica, 28, 256–257. [in Chinese]
    Makarova, O.L. (2004) Gamasid mites (Parasitiformes, Mesostigmata) dwelling in bracket fungi at the Pechoro-Ilychskii Reserve (Republic of Komi). Entomological Review, 84, 667–672. [Originally published in Zoologichesky Zhurnal, 2004, 83, 1335‒1340.]
    Makarova, O.L. (2009) The fauna of free-living gamasid mites (Parasitiformes, Mesostigmata) in the northern taiga: an analysis of the zonal specificity. Entomological Review, 89, 1177–1193. [Originally published in Zoologichesky Zhurnal, 2009, 88, 1039–1054.]
    Makarova, O.L. (2011) A review of gamasid mites (Parasitiformes, Mesostigmata) dwelling in the taiga of the Pechoro-Ilychskii Nature Reserve (northern Cis-Ural Region) with analysis of their assemblages in spruce forests. Entomological Review, 91, 915–931. [Originally published in Zoologichesky Zhurnal, 2011, 90, 649–664.]
    Mašán, P. (1994) The mesostigmatic mites (Acarina, Mesostigmata) associated with the dung beetles (Coleoptera, Scarabaeidae) in South Slovakia. Biologia (Bratislava), 49, 201–205.
    Mašán, P. (1998) Two new mesostigamatic mites (Acarina, Proctolaelaps, Hypoaspis) associated with erotylid and melolonthine beetles (Coleoptera: Erotylidae, Scarabaeidae) from Slovakia. Entomological Problems, 29, 19–22.
    Mašán, P. (2001a) First records of mesostigmatid mite species (Acarina) in Slovakia. Biologia, Bratislava, 56, 216–218.
    Mašán, P. (2001b) First records of mites (Acari) from Slovakia. Biologia, Bratislava, 56, 496, 502.
    Mašán, P. & Halliday, B. (2009) Mesostigmatid mites associated with the dung beetle Copris lunaris (Coleoptera: Scarabaeidae). European Journal of Entomology, 106, 545–550.
    Mašán, P. & Krištofík, J. (1993) Mites and ticks (Acarina: Mesostigmata et Ixodida) from the nests of Riparia riparia L. in South Slovakia. Biologia (Bratislava), 48, 155–162.
    Mašán, P. & Krištofík, J. (1995) Mesostigmatid mites (Acarina:Mesostigmata) in the nests of penduline tit (Remiz pendulinus). Biologia (Bratislava), 50, 481–485.
    Mašán, P. & Krištofík, J. (1996) Mites (Acarina, Gamasoidea) and fleas (Siphonaptera) from the nests of edible dormouse (Glis glis, Myoxidae). Entomofauna Carpathica, 8, 135–140.
    Mašán, P. & Országhová, Z. (1995) Mesostigmatic mites (Acarina) in the winter nests of Hirundo rustica in the vicinity of Bratislava (Slovakia). Acta Zoologica Universitatis Comenianae, 39, 33–37.
    Mašán, P. & Stanko, M. (2005) Mesostigmatic mites (Acari) and fleas (Siphonaptera) associated with nests of mound-building mouse, Mus spicilegus Petényi, 1882 (Mammalia, Rodentia). Acta Parasitologica, 50, 228–234.
    Mašán, P., Joharchi, O. & Abramov, V.V. (2021) A new genus and two new species of melicharid mites (Acari: Mesostigmata) associated with wood-destroying fungi and mycophagous erotylid beetles (Coleoptera: Erotylidae) in Europe. Zootaxa, 4980, 157–173.
    Mašán, P., Kalúz, S. & Babjaková, A. (1994) Mites (Acarina) from the winter nests of the common mole (Talpa europaea L.) in south Slovakia. Biologia (Bratislava), 49, 667–673.
    Mašán, P., Perotti, M.A., Saloña-Bordas, M.I. & Braig, H.R. (2013) Proctolaelaps euserratus, an ecologically unusual melicharid mite (Acari, Mesostigmata) associated with animal and human decomposition. Experimental and Applied Acarology, 61, 415–429.
    McGraw, J.R. & Farrier, M.H. (1969) Mites of the superfamily Parasitoidea (Acarina: Mesostigmata) associated with Dendroctonus and Ips (Animalia: Curculionidae). North Carolina Agricultural Experiment Station, Technical Bulletin, 192, 1–162.
    Mineiro, J.L.C., Lindquist, E.E. & Moraes, G.J. (2009) Edaphic ascid mites (Acari: Mesostigmata: Ascidae) from the state of São Paulo, Brazil, with description of five new species. Zootaxa, 2024, 1–32.
    Mock, A., Ľuptáčik, P., Fenďa, P., Svatoň, J., Országh, I. & Krumpál, M. (2005) Terrestrial arthropods inhabiting caves near Veľký Folkmar (Čierna hora Mountains, Slovakia). In: Tajovský, K., Schlaghamerský, J. & Pižl, V. (Eds), Contributions to Soil Zoology in Central Europe I, ISB ASCR, České Budějovice, pp. 95–101.
    Moraes, G.J., Britto, E.P.J, Mineiro, J.L.C. & Halliday, B. (2016) Catalogue of the mite families Ascidae Voigts & Oudemans, Blattisociidae Garman and Melicharidae Hirschmann (Acari: Mesostigmata). Zootaxa, 4112 (1), 1–299.
    Moraza, M.L. & Lindquist, E.E. (2011) A new genus of fungus-inhabiting blattisociid mites (Acari: Mesostigmata: Phytoseioidea) from Middle America, with a key to genera and subgenera of the subfamily Blattisociinae. Zootaxa, 2758, 1–25.
    Moser, J.C., Wilkinson, R.C. & Clark, E.W. (1974) Mites associated with Dendronctonus frontalis Zimmerman (Scolytidae: Coleoptera) in Central America and Mexico. Turrialba, 24, 379–381.
    Mrciak, M. (1958) Ďalšie nálezy roztočov radu Parasitiformes z drobných cicavcov ČSR. Biologia (Bratislava), 13, 311–314. [in Slovak]
    Mrciak, M. (1963) O vzťahoch niektorých druhov roztočov (Gamasoidea) k malým cicavcom a biotopom v Ondavskej vrchovine. Acta Facultatis Rerum Naturalium Universitatis Comenianae, Zoologia, 7, 437–532. [in Slovak]
    Mrciak, M. (1977) K faune roztočov (Acarina) drobných cicavcov okolia retenčnej nádrže Zemplínska Šírava po zavodnení. Acta Facultatis Rerum Naturalium Universitatis Comenianae, Zoologia, 22, 87–108. [in Slovak]
    Mrciak, M., Daniel, M. & Rosický, B. (1966) Parasites and nest inhabitants of small mammals in the western Carpathians. I. Mites of the superfamily Gamasoidea (Parasitiformes). Acta Facultatis Rerum Naturalium Universitatis Comenianae, Zoologia, 13, 81–116.
    Müller, J. (1859) Beitrag zur Höhlenfauna Mährens. Lotos, Zeitschrift für Naturwissenschaften, 9, 26–33.
    Narayanan, E.S. & Ghai, S. (1963) A new species of Melichares (Melichares) Hering (Aceosejidae) associated with the fig insects. Proceedings of the National Institute of Sciences of India, 29B, 547–550.
    Naskrecki, P. & Colwell, R.K. (1995) New genus and two new species of Melicharini from Venezuela (Acari: Mesostigmata: Ascidae). Annals of the Entomological Society of America, 88, 284–293.
    Negm, M.W. & Gotoh, T. (2021) Ontogenetic description of Proctolaelaps bickleyi (Bram) (Acari:), newly recorded from lepidopteran insect cultures in Japan. Systematic & Applied Acarology, 26(7), 1314–1326.
    Nesbitt, H.H.J. (1951) A taxonomic study of the Phytoseiinae (family Laelaptidae) predaceous upon Tetranychidae of economic importance. Zoologische Verhandelingen, 12, 1–64, 32 plates.
    Nikolsky, V.V. (1984) New species of gamasid mites (Parasitiformes, Gamasina) from Siberia. Novye i Maloizvestnye Vidy Fauny Sibiri, 14, 26–33. [in Russian]
    OConnor, R. & Klimov, P.B. (2012) Genus Proctolaelaps Berlese, 1923. [date of access on September 10, 2021].
    Országhová, Z., Hrvoľ, J., Benická, V. & Sobeková, K. (2006) Fauna Bratislavy. Hniezdna biológia lastovičky domovej (Hirundo rustica). Univerzita Komenského Bratislava vo vydavateľstve UK, Bratislava, 126 pp. [in Slovak]
    Ostovan, H. & Faraji, F. (2014) A new species of Orthadenella (Athias-Henriot) (Acari: Mesostigmata: Melicharidae) from Iran with comments on O. lawrencei (Evans). International Journal of Acarology, 40, 605–610.
    Oudemans, A.C. (1902) New list of Dutch Acari, second part. With remarks on known and descriptions of a new subfamily, new genera and species. Tijdschrift voor Entomologie, 45, 1–52, 6 plates.
    Oudemans, A.C. (1905) Acarologische Aanteekeningen XVII. Entomologische Berichten, 1 (23), 222–226.
    Oudemans, A.C. (1913) Acarologische Aanteekeningen XLIX. Entomologische Berichten, 4 (73), 2–18.
    Oudemans, A.C. (1916) Acarologische Aanteekeningen LIX. Entomologische Berichten, 4 (90), 296–299.
    Oudemans, A.C. (1929a) Acarologische Aanteekeningen XCIX. Entomologische Berichten, 8 (169), 11–20.
    Oudemans, A.C. (1929b) Acarologische Aanteekeningen C. Entomologische Berichten, 8 (170), 28–36.
    Oudemans, A.C. (1936) Kritisch Historisch Overzicht der Acarologie, Deerde Gedeelte, 1805–1850. Band A, Acari, Holothyroidea, Mesostigmata. E.J. Brill, Leiden, xx + 430 pp.
    Papáč, V., Fenďa, P., Ľuptáčik, P., Mock, A., Svatoň, J. & Christophoryová, J. (2009) Terestrické bezstavovce (Evertebrata) jaskýň vo vulkanitoch Cerovej vrchoviny. Aragonit, 14, 32–42. [in Slovak]
    Papáč, V., Kováč, Ľ., Mock, A., Košel, V. & Fenďa, P. (2006) Terestrické článkonožce (Arthropoda) vybraných jaskýň Silickej planiny. In: Bella, P. (Ed.), Výskum, využívanie a ochrana jaskýň. 5. Vedecká konferencia s medzinárodnou účasťou, Zborník referátov. Správa slovenských jaskýň, Liptovský Mikuláš, pp. 187–199. [in Slovak]
    Papáč, V., Ľuptáčik, P. & Fenďa, P. (2007) Terestrické článkonožce Obrovskej priepasti (Slovenský kras, Dolný vrch). Aragonit, 12, 51–53. [in Slovak]
    Pinchuk, L.M. (1972) New species of gamasoid mites (Parasitiformes: Gamasoidea) from nests of Moldavian mammals. Izvestiya Akademii Nauk Moldavskoi SSR, Seriya Biologicheskikh i Khimicheskikh Nauk, Kishinev, 5, 67–72. [in Russian]
    Pinchuk, L.M. (1976) Gamasina Mites of Small Mammals of Dnestr-Prut Region. Akademia Nauk Moldavskoi SSR, Institut Zoologii, Shtiinza, Kishinev, 174 pp. [in Russian]
    Rittner, O. & Nir, A. (2013) First record of Myrrha octodecimguttata (Coleoptera: Coccinellidae) and Epuraea ocularis (Coleoptera: Nitidulidae) from Israel. Zootaxa, 3609, 349–350.
    Samšiňák, K. (1958) Roztoči ze slunečnicových semen (Acari). Československá Parasitologie, 5, 185–198.
    Samšiňák, K. (1960) Über einige forstwirtschaftlich wichtige Milben der Gattung Proctolaelaps Berlese 1923. Československá Parasilogie, 7, 297–307.
    Samšiňák, K. (1964) Termitophile Milben aus der VR China. I. Mesostigmata. Entomologische Abhandlungen Staatliches Museum für Tierkunde in Dresden, 32 (3), 33–52.
    Samšiňák, K., Najvarová, Š. & Vobrázková, E. (1979) Roztočí zvířena lůžek dětských léčeben na Štrbském Plese. Práce Slovenskej Entomologickej Spoločnosti (Bratislava), 1, 243–246. [in Czech]
    Sanyal, A.K. & Gupta, S.K. (2005) Acari from Antarctic Continent: One new species of Proctolaelaps (Mesostigmata: Ascidae) and some new reports (Prostigmata, Astigmata, Mesostigmata, Cryptostigmata). Proceedings of the Zoological Society, Calcutta, 58 (2), 79–84.
    Schweizer, J. (1949) Die Landmilben des Schweizerischen Nationalparkes. 1. Teil: Parasitiformes Reuter 1909. Ergebnisse der Wissenschaftlichen Untersuchung des Schweizerischen Nationalparks, Neue Folge, 2 (21), 1–99.
    Sidor, V. (1986) Akarofauna veľkochovu hospodárskych zvierat Agrokomplexu v Nitre. Poľnohospodárstvo, 32, 737–747. [in Slovak]
    Sourassou, N.F., Moraes, G.J. & Santos, J.C. (2015) Orolaelaps (Acari: Mesostigmata: Melicharidae): description of two new species, redescription of Orolaelaps quisqualis and new characterisation of the genus. Zootaxa, 4039 (2), 312–322.
    Stanko, M. (1988) Poznámky k faune ektoparazitov (Siphonaptera, Mesostigmata, Ixodoidea) drobných cicavcov doliny Delňa (Slanské vrchy). In: Ambros, M. (Ed.), XII. Východoslovenský tábor ochrancov prírody (Sigord – Kokošovce), Prehľad odborných výsledkov, Prešov, pp. 102–113. [in Slovak]
    Stanko, M. (1989) Roztoče (Acarina, Mesostigmata) drobných cicavcov dvoch územných celkov Západných Karpát. Biologia (Bratislava), 44, 449–512. [in Slovak]
    Stanko, M. (1990) Ektoparazity (Acarina: Mesostigmata, Ixodoidea) drobných zemných cicavcov zoologickej záhrady v Košiciach. Zborník Východoslovenského Múzea v Košiciach, Prírodné Vedy, 31, 159–171. [in Slovak]
    Stanko, M. (1995) Súčasný stav poznania fauny ektoparazitov (Siphonaptera, Anoplura, Ixodida, Mesostigmata) drobných cicavcov územia biosférickej rezervácie Východné Karpaty. Natura Carpatica, 26, 61–70. [in Slovak]
    Stanko, M. (1999) Ektoparazity drobných cicavcov národnej prírodnej rezervácie Latorický luh (Východoslovenská nížina). 2. Roztoče (Mesostigmata). Natura Carpatica, 40, 101–108. [in Slovak]
    Stanko, M. & Fričová, J. (1996) Príspevok k poznaniu fauny drobných cicavcov (Insectivora, Rodentia) a ich ektoparazitov v okolí prírodnej pamiatky Plavečské štrkoviská (Spišskošarišské medzihorie). Ochrana Prírody, 14, 143–151. [in Slovak]
    Stanko, M. & Miklisová, D. (1995) Interactions of small mammal communities of windbreaks and adjacent fields with respect to epidemiological aspect. Ekológia (Bratislava), 14, 3–16.
    Stanko, M., Fričová, J., Miko, L. & Miklisová, D. (1993) Spoločenstvá článkonožcov (Acarina, Insecta) na drobnýchcicavcoch (Insectivora, Rodentia) dolného toku Ondavy (Východoslovenská nížina). Správy Slovenskej Entomologickej Spoločnosti Pri SAV, 4, 9–16. [in Slovak]
    Stone, C. (1988) A new species of Proctolaelaps (Acari: Ascidae) from New South Wales, Australia. Acarologia, 29, 319–327.
    Švaňa, M., Fenďa, P. & Országhová, Z. (2006) Roztoče (Acarina, Mesostigmata) v hniezdach vtákov JZ Slovenska. Folia Faunistica Slovaca, 11, 39–42. [in Slovak]
    Szabó, A., Ripka, G., Hajdu, Zs., Tempfli, B., Varga, M., Mészáros, I., Kutasi, Cs., Németh, T. & Pénzes, B. (2013) New data on the mesostigmatid mite fauna of Hungary (Acari: Mesostigmata). Acta Phytopathologica et Entomologica Hungarica, 48, 149–154.
    Thor, S. (1930) Beiträge zur Kenntnis der Invertebraten Fauna von Svalbard. Skrifter om Svalbard og Ishavet, 27, 1–156.
    Trach, V.A. & Khaustov, A.A. (2017) Mites of the genus Proctolaelaps Berlese, 1923 (Acari: Mesostigmata: Melicharidae) associated with bark beetles in Asian Russia. Acarina, 25, 151–163.
    Trach, V.A., Khaustov, A.A. & Lindquist, E.E. (2019) A new unique species of Mucroseius Lindquist, 1962 (Acari: Mesostigmata: Melicharidae) mites associated with sawyer beetles (Cerambycidae: Monochamus Dejean, 1821) from the Palaearctic Region. Journal of Natural History, 53, 13-14, 761-778.
    Trach, V.A., Marchenko, I.I. & Joharchi, O. (2019) Redescription of the female of bumblebee-associated gamasid mite Proctolaelaps sibiriensis (Davydova, 1988) (Acari: Mesostigmata: Melicharidae) from North Asia. Acarologia, 59, 531‒541.
    Treat, A.E. & Niederman, L. (1967) Three species of Proctolaelaps (Acarina: Mesostigmata) from noctuid moths. American Museum Novitates, 2312, 1–12.
    Turk, F.A. & Turk, S.M. (1952) Studies of Acari – 7th series: records and descriptions of mites new to the British fauna, together with short notes on the biology of sundry species. Annals and Magazine of Natural History, Series 12, 5, 475–506.
    Várfalvyová, D., Miklisová, D. & Stanko, M. (2010) Charakteristika spoločenstiev mezostigmátnych roztočov (Mesostigmata) v hniezdach Mus spicilegus (Rodentia, Muridae) na Slovensku. Folia Faunistica Slovaca, 15, 13–17. [in Slovak]
    Vávra, J.Ch., Průdek, P., Hauck, D. & Čížek, L. (2016) Faunistic records from the Czech Republic – 403. Coleoptera: Ptinidae, Nitidulidae, Monotomidae. Klapalekiana, 52, 99–101.
    Vitzthum, H.G. (1923) Acarologische Beobachtungen. 7 Reiche. Archiv für Naturgeschichte, A 89 (2), 97–181.
    Walter, D.E. & Proctor, H.C. (2013) Mites: Ecology, Evolution & Behaviour. Life at a Microscale. Second Edition. Springer, Dordrecht, 494 pp.
    Westerboer, I. (1963) Die Familie Podocinidae Berlese 1916. Abschnitt IV. In: Stammer, H.-J. (Ed.), Beiträge zur Systematik und Ökologie mitteleuropäischer Acarina. Band II. Mesostigmata I. Akademische Verlagsgesellschaft Geest & Portig K.- G., Leipzig, pp. 179–450.
    Wilson, D.S. (1980) The Natural Selection of Populations and Communities. The Benjamin/Cummings Publishing Company, Inc. Menlo Park, California, 186 pp.
    Wiśniewski, J. (1980) Zwei neue mit Proctolaelaps xyloteri Samš. (Mesostigmata, Blattisociidae) verwandte Proctolaelaps-Arten aus Polen. Acarologia, 21, 3–8.
    Womersley, H. (1956) On some new Acarina-Mesostigmata from Australia, New Zealand and New Guinea. The Journal of the Linnean Society of London, Zoology, 42, 505–599.
    Zeman, P. & Jurík, M. (1981) A contribution to the knowledge of fauna and ecology of gamasoid mites in cavity nests of birds in Czechoslovakia. Folia Parasitologica (Prague), 28, 265–271.