Skip to main content Skip to main navigation menu Skip to site footer
Type: Correspondence
Published: 2022-02-04
Page range: 495-500
Abstract views: 430
PDF downloaded: 18

A new fossil of Ceratophrys Wied-Neuwied (Ceratophryidae: Anura) from Southeastern Brazil

Programa de Pós-Graduação em Evolução e Diversidade, CCNH, Universidade Federal do ABC, Al. da Universidade s/n (09606-045), São Bernardo do Campo, São Paulo, Brazil
Programa de Pós-Graduação em Evolução e Diversidade, CCNH, Universidade Federal do ABC, Al. da Universidade s/n (09606-045), São Bernardo do Campo, São Paulo, Brazil


  1. Araújo, C.D.O., Condez, T.H., Bovo, R.P., Centeno, F.D.C. & Luiz, A.M. (2010) Amphibians and reptiles of the Parque Estadual Turístico do Alto Ribeira (PETAR), SP: An Atlantic Forest remnant of southeastern Brazil. Biota Neotropica, 10 (4), 257–274.
    Barcelos, L.A., Almeida-Silva, D., Santos, C.M.D. & Verdade, V.K. (2020) Description of a new species of fossil Ceratophrys (Anura: Ceratophryidae) from Versalles Cave, São Paulo, Brazil. Journal of Vertebrate Paleontology, 40 (3), e1811293.
    de Almeida, F.F.M., Amaral, G., Cordani U.G. & Kawashita, K. (1973) The Precambrian Evolution of the South American Cratonic Margin South of the Amazon River. In: Nairn A.E.M. & Stehli, F.G. (Eds.), The South Atlantic. Springer, Boston, Massachusetts, pp 411–446.
    Fabrezi, M. (2006) Morphological evolution of Ceratophryinae (Anura, Neobatrachia). Journal of Zoological Systematics and Evolutionary Research, 44, 153–166.
    Fabrezi, M. & Quinzio, S.I. (2008) Morphological evolution in Ceratophryinae frogs (Anura, Neobatrachia): the effects of heterochronic changes during larval development and metamorphosis. Zoological Journal of the Linnean Society, 154, 752−780.
    Faivovich, J., Nicoli, L., Blotto, B.L., Pereyra, M.O., Baldo, D., Barrionuevo, J.S., Fabrezi, M., Wild E.R. & Haddad, C.F.B. (2014) Big, bad, and beautiful: phylogenetic relationships of the horned frogs (Anura: Ceratophryidae). South American Journal of Herpetology, 9, 207–227.
    Frost, D.R. (2021) Amphibian Species of the World: An Online Reference. Version 6.0. American Museum of Natural History, New York, New York. Available from: (accessed 25 June 2021)
    Gómez, R.O. & Turazzini, G.F. (2021) The fossil record and phylogeny of South American horned frogs (Anura, Ceratophryidae). Journal of Systematic Palaeontology, 19 (2), 91–130.
    Günther, A.C.L.G. (1859) Sexual Differences Found in Bones of Some Recent and Fossil Species of Frogs and Fishes. Annals and Magazine of Natural History, 3, 377–386.
    Lynch, J.D. (1971) Evolutionary relationships, osteology, and zoogeography of leptodactyloid frogs. University Kansas Museum Natural History, Miscellaneous. Publications 53. University Kansas Museum Natural History, Lawrence, Kansas, 238 pp.
    Lynch, J.D. (1982) Relationships of the frogs of the genus Ceratophrys (Leptodactylidae) and their bearing on hypotheses of Pleistocene forest refugia in South America and punctuated equilibria. Systematic Biology, 31, 166–179.
    Maciel, N.M., Kokubum, M.N.C., Braga, P.H.P., Queiróz Jr., A.T. & Matsushita, R.H. (2013) Distribution extension, new state record and geographic distribution map of Ceratophrys joazeirensis Mercadal, 1986 (Anura: Ceratophryidae). Herpetology Notes, 6, 447–450.
    Mercadal, I.T. (1986) Ceratophrys joazeirensis sp. n. (Ceratophryidae, Anura) del noreste de Brasil. Amphibia-Reptilia, 7 (4), 313–334.
    Nicoli, L. (2019) The fossil record of Ceratophrys Wied-Neuwied (Anura: Ceratophryidae): a revision and update of fossil South American horned frogs. Zootaxa, 4658 (1), 37–68.
    Nicoli, L., Tomassini, R.L. & Montalvo, C.I. (2017) The oldest record of Ceratophrys (Anura, Ceratophryidae) from the Late Miocene of central Argentina. Journal of Vertebrate Paleontology, 37, e1261360, 1–5.
    Oliveira A.M. (2018) Megafauna Cenozóica do continente Americano: considerações sobre ecomorfologia, paleoecologia, evolução e paleobiogeografia. Ph.D. dissertation. Universidade Federal do ABC, Santo André. [unknown pagination]
    Simon, M.N., Machado, F.A. & Marroig, G. (2016) High evolutionary constraints limited adaptive responses to past climate changes in toad skulls. Proceedings of the Royal Society B: Biological Sciences, 283 (1841), 20161783.
    Vieira, K.S., Silva, A.Z. & Arzabe, C. (2006) Cranial morphology and karyotypic analysis of Ceratophrys joazeirensis (Anura: Ceratophryidae, Ceratophrynae): taxonomic considerations. Zootaxa, 1320 (1), 57–68.
    Vieira, K.S., Montenegro, P.F.G., Santana, G.G. & da Silva-Vieira, W.L. (2018) Effect of climate change on distribution of species of common horned frogs in South America. PloS one, 13 (9), e0202813.
    Wild, E. (1997) The ontogeny and phylogeny of Ceratophryine frogs (Anura: Leptodactylidae). Ph.D. dissertation. University of Kansas, Lawrence, Kansas, 24 pp.