Skip to main content Skip to main navigation menu Skip to site footer
Type: Article
Published: 2021-03-30
Page range: 201–247
Abstract views: 202
PDF downloaded: 62

Report on hydrozoans (Cnidaria), excluding Stylasteridae, from the Emperor Seamounts, western North Pacific Ocean

Research Associate, Royal British Columbia Museum, 675 Belleville Street, Victoria, British Columbia, Canada V8W 9W2.
School of Life Sciences, 216 Edmondson Hall, University of Hawaii at Manoa, Honolulu, Hawaii 96822, USA.
Anthoathecata Hydroidolina hydroids Leptothecata marine invertebrates Medusozoa taxonomy Trachymedusae zoological nomenclature

Abstract

Fourteen species of hydroids, collected during August 2019 by ROV SuBastian of the Schmidt Ocean Institute, are reported from the Emperor Seamount chain in the western North Pacific Ocean. Two others, Candelabrum sp. and Eudendrium sp., were observed only on videos taken by the ROV. From collections and video observations, eight species of hydroids were found at Jingū Seamount, three at Yomei, Nintoku, and Annei seamounts, and one at Koko Seamount and Hess Rise. At Suiko and Godaigo seamounts, hydroids were seen in videos but they could not be identified. Latebrahydra schulzei, an endobiotic associate of the hexactinellid sponge Walteria flemmingii Schulze, 1886 from Annei Seamount and Hess Rise, is described as a new genus and species tentatively attributed to Hydractiniidae L. Agassiz, 1862. Another new species, Hydractinia galeai, is described from Jingū Seamount. Among its distinctive characters is a zooid termed a sellectozooid, likely serving in both food capture and defence. Hydroids examined from Yomei, Nintoku, and Jingū seamounts are elements of a cold-water fauna occurring in the North Pacific Boreal Bathyal province, while those of Annei and Koko seamounts, and Hess Rise, are part of the biota of the Central North Pacific Bathyal province. Hydroids identified as Bouillonia sp., from Nintoku Seamount, represent the first record of this predominantly deep water tubulariid genus in the North Pacific Ocean. Bonneviella superba Nutting, 1915, from Jingū Seamount, is reported for the first time outside the Aleutian Islands. Bonneviella cf. gracilis Fraser, 1939, known elsewhere only from Dease Strait in the western Canadian Arctic, was also collected on Jingū. In addition to hydroids, medusae of Ptychogastria polaris Allman, 1878 were observed on videos from Nintoku, Jingū, Annei, and Koko seamounts at depths between 2423–1422 m. An unidentified siphonophore was observed near bottom at 2282 m on Nintoku Seamount.

 

References

  1. Agassiz, A. (1865) Illustrated catalogue of the Museum of Comparative Zoölogy, at Harvard College. No. II. North American Acalephae. Sever & Francis, Cambridge, Massachusetts, 234 pp.

    Agassiz, L. (1862) Contributions to the natural history of the United States of America. Vol. IV. Little, Brown & Company, Boston, Massachusetts, 380 pp.

    Alder, J. (1856) A notice of some new genera and species of British hydroid zoophytes. Annals and Magazine of Natural History, Series 2, 18, 353–362.

    https://doi.org/10.1080/00222935608697652

    Alder, J. (1859) Descriptions of three new species of sertularian zoophytes. Annals and Magazine of Natural History, Series 3, 3, 353–356.

    Alder, J. (1862) Observations on British zoophytes. Edinburgh New Philosophical Journal, New Series, 15, 144.

    Allman, G.J. (1863) Notes on the Hydroida. I. On the structure of Corymorpha nutans. II. Diagnoses of new species of Tubularidae obtained, during the autumn of 1862, on the coasts of Shetland and Devonshire. Annals and Magazine of Natural History, Series 3, 11, 1–12.

    https://doi.org/10.1080/00222936308681369

    Allman, G.J. (1876) Diagnoses of new genera and species of Hydroida. Journal of the Linnean Society, Zoology, 12, 251–284.

    https://doi.org/10.1111/j.1096-3642.1876.tb00682.x

    Allman, G.J. (1878) Hydrozoa. In: Nares, G.S. (Ed.), Narrative of a voyage to the Polar Sea during 1875–6 in H.M. Ships ‘Alert’ and ‘Discovery.’ With notes on the natural history edited by H.W. Feilden, F.G.S., C.M.Z.S., F.R.G.S. Naturalist to the expedition. Vol. II. Sampson Low, Marston, Searle, & Rivington, London, pp. 290–292.

    Allman, G.J. (1888) Report on the Hydroida dredged by H.M.S. Challenger during the years 1873–76. Part II—The Tubularinae, Corymorphinae, Campanularinae, Sertularinae, and Thalamophora. Report on the Scientific Results of the Voyage of H.M.S. Challenger during the Years 1873–76, Zoology, 7 (23), 1–90.

    Antsulevich, A.E. (1980) O dvukh dalnevostochnykh vidakh gidroidov (Leptolida, Thecaphora). Zoologicheskiï Zhurnal, 59, 615–617. [in Russian]

    Antsulevich, A.E. (1987) Gidroidy shelfa Kurilskikh Ostrovov. Akademiya Nauk SSSR, Zoologicheskii Institut, Leningrad, 165 pp. [in Russian]

    Antsulevich, A.E. (2009) Ob izuchennosti fauny gidroidov Beringova Morya. Vestnik Sankt-Peterburgskogo Universiteta, 3 (3), 14–30. [in Russian]

    Antsulevich, A.E. (2015) Hydrozoa (hydroids and hydromedusae) of Russian seas. St. Petersburg University Press, St. Petersburg, 860 pp. [in Russian]

    Antsulevich, A. & Vervoort, W. (1993) Some little-known species of hydroids (Cnidaria: Hydrozoa: Lafoeidae) and description of Papilionella pterophora gen. nov., spec. nov. (Sertulariidae). Zoologische Mededelingen, Leiden, 67, 431–443.

    Aurivillius, C.W.S. (1895) Das Plankton der Baffins Bay und Davis’ Strait. Eine thiergeographische Studie. Zoologiska Studier: Festskrift Wilhelm Lilljeborg tillegnad på hans åttionde Födelsedag af Svenska zoologer, 1895, 179–212.

    Aurivillius, C.W.S. (1899) Animalisches Plankton aus dem Meere zwischen Jan Mayen, Spitzbergen, K. Karls Land und der Nordkuste Norwegens. Kongliga Svenska Vetenskaps-Akademiens Handlingar, Ny Följd, 32 (6), 1–71.

    Bale, W.M. (1888) On some new and rare Hydroida in the Australian Museum collection. Proceedings of the Linnean Society of New South Wales, Series 2, 3, 745–799.

    https://doi.org/10.5962/bhl.title.26267

    Beaulieu, S.E. (2001) Life on glass houses: sponge stalk communities in the deep sea. Marine Biology, 138, 803–817.

    https://doi.org/10.1007/s002270000500

    Bedot, M. (1912) Matériaux pour servir à l’histoire des hydroïdes. 4me période (1872 à 1880). Revue Suisse de Zoologie, 20, 213–469.

    Bedot, M. (1916) Matériaux pour servir à l’histoire des hydroïdes. 5e période (1881 à 1890). Revue Suisse de Zoologie, 24, 1–349.

    Bedot, M. (1918) Matériaux pour servir à l’histoire des hydroïdes. 6e période (1891 à 1900). Revue Suisse de Zoologie, 26, Fascicule Supplémentaire, 1–376.

    Bedot, M. (1925) Matériaux pour servir à l’histoire des hydroïdes. 7e période (1901 à 1910). Revue Suisse de Zoologie, 32, Fascicule Supplémentaire, 1–657.

    Bergmann, M., Dannheim, J., Bauerfeind, E. & Klages, M. (2009) Trophic relationships along a bathymetric gradient at the deep-sea observatory HAUSGARTEN. Deep-Sea Research I, 56, 408–424.

    https://doi.org/10.1016/j.dsr.2008.10.004

    Beyer, F. (1958) A new, bottom-living trachymedusa from the Oslofjord. Description of the species, and a general discussion of the life conditions and fauna of the fjord deeps. Nytt Magasin for Zoologi, 6, 121–143.

    Bigelow, H.B. (1909) Reports on the scientific results of the Expedition to the Eastern Tropical Pacific, in charge of Alexander Agassiz, by the U.S. Fish Commission Steamer “Albatross,” from October, 1904, to March, 1905, Lieut. Commander L.M. Garrett, U.S.N., commanding. XVI. The Medusae. Memoirs of the Museum of Comparative Zoölogy at Harvard College, 37, 1–243.

    Bigelow, H.B. (1913) Medusae and Siphonophorae collected by the U. S. Fisheries Steamer “Albatross” in the northwestern Pacific, 1906. Proceedings of the United States National Museum, 44, 1–119.

    https://doi.org/10.5479/si.00963801.44-1946.1

    Billard, A. (1929) Note sur un genre nouveau et quelques espèces nouvelles d’Halecidae. Bulletin de la Société Zoologique de France, 54, 305–307.

    Blainville, H.M.D. de (1830) Zoophytes, Zoophyta. In: Levrault, F.G. (Ed.), Dictionnaire des Sciences Naturelles...par plusieurs Professeurs du Jardin du Roi, et des Principales écoles de Paris. Tome 60. Le Normant, Paris, pp. 1–548.

    Blanco, R., Shields, M.A. & Jamieson, A.J. (2013) Macrofouling of deep-sea instrumentation after three years at 3690 m depth in the Charlie Gibbs fracture zone, mid-Atlantic ridge, with emphasis on hydroids (Cnidaria: Hydrozoa). Deep-Sea Research II, 98, 370–373.

    https://doi.org/10.1016/j.dsr2.2013.01.019

    Bonnevie, K. (1898) Zur Systematik der Hydroiden. Zeitschrift für Wissenschaftliche Zoologie, 63, 465–495.

    Bouillon, J., Gravili, C., Pagès, F., Gili, J.-M. & Boero, F. (2006) An introduction to Hydrozoa. Mémoires du Muséum National d’Histoire Naturelle, 194, 1–591.

    Brinckmann-Voss, A. (1983) British Columbia marine faunistic report on the Hydrozoa: Part II. Hydroids. Canadian Technical Report of Fisheries and Aquatic Sciences, 1185, 1–20.

    Brinckmann-Voss, A. (1996) Seasonality of hydroids (Hydrozoa, Cnidaria) from an intertidal pool and adjacent subtidal habitats at Race Rocks, off Vancouver Island, Canada. Scientia Marina, 60, 89–97.

    Broch, H. (1907) Hydroiden und Medusen. Report of the Second Norwegian Arctic Expedition in the “Fram” 1898–1902, 2 (12), 1–12.

    Broch, H. (1909) Hydroiduntersuchungen II. Zur Kenntnis der Gattungen Bonneviella und Lictorella. Nyt Magazin for Naturvidenskaberne, 47, 195–206.

    Broch, H. (1910) Die Hydroiden der Arktischen Meere. Fauna Arctica, 5, 127–248.

    Broch, H. (1918) Hydroida (Part II.). The Danish Ingolf Expedition, 5 (7), 1–205.

    Broch, H. (1928) Hydrozoa I. In: Grimpe, G. & Wagler, E. (Eds.), Die Tierwelt der Nord-und Ostsee, 3.b, pp. 1–100.

    Broch, H. (1948) Post-glacial interchange between Pacific and Atlantic deeper living boreal faunas? Norsk Geografisk Tidsskrift, 12, 126–141.

    https://doi.org/10.1080/00291954808551657

    Browne, E.T. (1907) The hydroids collected by the “Huxley” from north side of the Bay of the Bay of Biscay in August, 1906. Journal of the Marine Biological Association of the U.K., 8, 15–36.

    https://doi.org/10.1017/S002531540004371X

    Calder, D.R. (1970) Thecate hydroids from the shelf waters of northern Canada. Journal of the Fisheries Research Board of Canada, 27, 1501–1547.

    https://doi.org/10.1139/f70-175

    Calder, D.R. (1996) Hydroids (Cnidaria: Hydrozoa) recorded from depths exceeding 3000 m in the abyssal western North Atlantic. Canadian Journal of Zoology, 74, 1721–1726.

    https://doi.org/10.1139/z96-190

    Calder, D.R. (1997) Synopsis of hydroids from 1000 m and deeper in the western North Atlantic. In: den Hartog, J.C. (Ed.), Proceedings of the 6th International Conference on Coelenterate Biology. Nationaal Natuurhistorisch Museum, Leiden, pp. 85–90.

    Calder, D.R. (2010) Some anthoathecate hydroids and limnopolyps (Cnidaria, Hydrozoa) from the Hawaiian archipelago. Zootaxa, 2590 (1), 1–91.

    https://doi.org/10.11646/zootaxa.2590.1.1

    Calder, D.R. (2017) Additions to the hydroids (Cnidaria, Hydrozoa) of the Bay of Fundy, northeastern North America, with a checklist of species reported from the region. Zootaxa, 4256 (1), 1–86.

    https://doi.org/10.11646/zootaxa.4256.1.1

    Calder, D.R. & Choong, H.H.C. (2018) Names of hydroids (Cnidaria, Hydrozoa) established by Charles McLean Fraser (1872–1946), excluding those from Allan Hancock Expeditions. Zootaxa, 4487 (1), 1–83.

    https://doi.org/10.11646/zootaxa.4487.1.1

    Calder, D.R. & Vervoort, W. (1998) Some hydroids (Cnidaria: Hydrozoa) from the Mid-Atlantic Ridge, in the North Atlantic Ocean. Zoologische Verhandelingen, Leiden, 319, 1–65.

    Calder, D.R., Vervoort, W. & Hochberg, F.G. (2009) Lectotype designations of new species of hydroids (Cnidaria, Hydrozoa), described by C.M. Fraser, from Allan Hancock Pacific and Caribbean Sea expeditions. Zoologische Mededelingen, Leiden, 83, 919–1058.

    Carter, H.J. (1877) On the close relationship of Hydractinia, Parkeria, and Stromatopora; with descriptions of new species of the former, both recent and fossil. Annals and Magazine of Natural History, Series 4, 19, 44–76.

    https://doi.org/10.1080/00222937708682095

    Carter, H.J. (1878) On new species of Hydractiniidae, recent and fossil, and on the identity in structure of Millepora alcicornis with Stromatopora. Annals and Magazine of Natural History, Series 5, 1, 298–311.

    https://doi.org/10.1080/00222937808682336

    Clark, S.F. (1876) The hydroids of the Pacific coast of the United States, south of Vancouver Island. With a report upon those in the Museum of Yale College. Transactions of the Connecticut Academy of Arts and Sciences, 3, 249–264.

    Clark, S.F. (1877) Report on the hydroids collected on the coast of Alaska and the Aleutian Islands, by W.H. Dall, U.S. Coast Survey, and party, from 1871 to 1874 inclusive. Proceedings of the Academy of Natural Sciences of Philadelphia, 1876, 28, 209–238.

    Cockerell, T.D.A. (1911) The nomenclature of the hydromedusae. Proceedings of the Biological Society of Washington, 24, 77–86.

    Collins, A.G. (2000) Towards understanding the phylogenetic history of Hydrozoa: hypothesis testing with 18S gene sequence data. Scientia Marina, 64 (Suplemento 1), 5–22.

    https://doi.org/10.3989/scimar.2000.64s15

    Collins, A.G., Winkelmann, S., Hadrys, H. & Schierwater, B. (2005) Phylogeny of Capitata and Corynidae (Cnidaria, Hydrozoa) in light of mitochondrial 16S rDNA data. Zoologica Scripta, 34, 91–99.

    https://doi.org/10.1111/j.1463-6409.2005.00172.x

    Cornelius, P.F.S. (1975) A revision of the species of Lafoeidae and Haleciidae (Coelenterata: Hydroida) recorded from Britain and nearby seas. Bulletin of the British Museum (Natural History), Zoology, 28, 375–426.

    Cornelius, P.F.S. (1982) Hydroids and medusae of the family Campanulariidae recorded from the eastern North Atlantic, with a world synopsis of genera. Bulletin of the British Museum (Natural History), Zoology, 42, 37–148.

    Cornelius, P.F.S. (1992) Medusa loss in leptolid Hydrozoa (Cnidaria), hydroid rafting, and abbreviated life-cycles among their remote-island faunae: an interim review. Scientia Marina, 56, 245–261.

    Cunha, A.F., Collins, A.G. & Marques, A.C. (2017) Phylogenetic relationships of Proboscoida Broch, 1910 (Cnidaria, Hydrozoa): are traditional morphological diagnostic characters relevant for the delimitation of lineages at the species, genus, and family levels? Molecular Phylogenetics and Evolution, 106, 118–135.

    https://doi.org/10.1016/j.ympev.2016.09.012

    Cunha, A.F., Collins, A.G. & Marques, A.C. (2020) When morphometry meets taxonomy: morphological variation and species boundaries in Proboscoida (Cnidaria: Hydrozoa). Zoological Journal of the Linnean Society, 190, 417–447.

    https://doi.org/10.1093/zoolinnean/zlz166

    Dunbar, M.J. (1942) Marine macroplankton from the Canadian eastern Arctic. II. Medusae, Siphonophora, Ctenophora, Pteropoda, and Chaetognatha. Canadian Journal of Research, 20D, 71–77.

    https://doi.org/10.1139/cjr42d-008

    Dvoretsky, V.G. & Dvoretsky, A.G. (2010) Checklist of fauna found in zooplankton samples from the Barents Sea. Polar Biology, 33, 991–1005.

    https://doi.org/10.1007/s00300-010-0773-4

    Dvoretsky, V.G. & Dvoretsky, A.G. (2018) Состав и распределение летнего зоопланктона полярного фронта (Баренцево море). [Composition and distribution of summer zooplankton of the Polar Front (Barents Sea)]. Вестник Кольского научного центра РАН, [Bulletin of the Kola Scientific Center of the Russian Academy of Sciences], 2 (10), 25–37. [in Russian]

    Edwards, C. (1973) The medusa Modeeria rotunda and its hydroid Stegopoma fastigiatum, with a review of Stegopoma and Stegolaria. Journal of the Marine Biological Association of the U.K., 53, 573–600.

    https://doi.org/10.1017/S0025315400058781

    Ehrenberg, C.G. (1834) Beiträge zur physiologischen Kenntniss der Corallenthiere im allgemeinen, und besonders des rothen Meeres, nebst einem Versuche zur physiologischen Systematik derselben. Abhandlung der Königlichen Akademie der Wissenschaften zu Berlin, 1, 225–380.

    Fabricius, O. (1780) Fauna Groenlandica. Rothe, Hafniae & Lipsiae, 452 pp.

    Feniuk, V. (1947) Some materials on the hydroids (Thecaphora) from the Ochotsk Sea. Bulletin de la Société des Naturalistes de Moscou, Section Biologique, Nouvelle Série, 52 (2), 3–13. [in Russian, with English summary]

    Fewkes, J.W. (1889) New Invertebrata from the coast of California. Bulletin of the Essex Institute, 21, 99–146.

    Fleming, J. (1820) Observations on the natural history of the Sertularia gelatinosa of Pallas. Edinburgh Philosophical Journal, 2, 82–89.

    Fraser, C.M. (1911) The hydroids of the west coast of North America with special reference to those of the Vancouver Island region. Bulletin from the Laboratories of Natural History of the State University of Iowa, 6 (1), 3–91.

    Fraser, C.M. (1912) Some hydroids of Beaufort, North Carolina. Bulletin of the United States Bureau of Fisheries, 30, 339–387.

    Fraser, C.M. (1914a) Some hydroids of the Vancouver Island region. Transactions of the Royal Society of Canada, Series 3, Section 4, 8, 99–216.

    Fraser, C.M. (1914b) Notes on some Alaskan hydroids. Transactions of the Royal Society of Canada, Series 3, Section 4, 8, 217–222.

    Fraser, C.M. (1931) Biological and oceanographic conditions in Hudson Bay. 3. Hydroids of Hudson Bay and Hudson Strait. Contributions to Canadian Biology and Fisheries, 6, 477–481.

    https://doi.org/10.1139/f31-025

    Fraser, C.M. (1935a) Hydroids from the west coast of Vancouver Island. Canadian Field-Naturalist, 49, 143–145.

    Fraser, C.M. (1935b) Some Japanese hydroids, mostly new. Transactions of the Royal Society of Canada, Series 3, Section 5, 29, 105–112.

    Fraser, C.M. (1936) Hydroid distribution in the vicinity of the Queen Charlotte Islands. Canadian Field-Naturalist, 50, 122–126.

    Fraser, C.M. (1937) Hydroids of the Pacific coast of Canada and the United States. University of Toronto Press, Toronto, 207 pp.

    https://doi.org/10.3138/9781487595548

    Fraser, C.M. (1938a) Hydroids of the 1934 Allan Hancock Pacific Expedition. Allan Hancock Pacific Expeditions, 4, 1–105.

    Fraser, C.M. (1938b) Hydroids of the 1932, 1933, 1935, and 1938 Allan Hancock Pacific Expeditions. Allan Hancock Pacific Expeditions, 4, 129–153.

    Fraser, C.M. (1939) Hydroids of the western Canadian Arctic region, 1935–1937. Canadian Journal of Research, Section D, Zoological Sciences, 17 (3), 59–61.

    https://doi.org/10.1139/cjr39d-008

    Fraser, C.M. (1944) Hydroids of the Atlantic coast of North America. University of Toronto Press, Toronto, 451 pp.

    Fraser, C.M. (1947) Distribution and relationship in American hydroids. University of Toronto Press, Toronto, 464 pp. [dating of this work follows Calder & Choong 2018]

    Fraser, C.M. (1948) Hydroids of the Allan Hancock Pacific Expeditions since March, 1938. Allan Hancock Pacific Expeditions, 4, 179–343.

    Galea, H.R., Roder, C., Walcher, C., Warmuth, M., Kohlberg, E. & Fischer, P.F. (2016) Glaciambulata neumayeri gen. et sp. nov., a new Antarctic trachymedusa (Cnidaria: Hydrozoa), with a revision of the family Ptychogastriidae. European Journal of Taxonomy, 252, 1–30.

    https://doi.org/10.5852/ejt.2016.252

    Galea, H.R. & Schories, D. (2012) Some hydrozoans (Cnidaria) from King George Island, Antarctica. Zootaxa, 3321, 1–21.

    https://doi.org/10.11646/zootaxa.3321.1.1

    Gasbarro, R., Wan, D. & Tunnicliffe, V. (2018) Composition and functional diversity of macrofaunal assemblages on vertical walls of a deep northeast Pacific fjord. Marine Ecology Progress Series, 597, 47–64.

    https://doi.org/10.3354/meps12599

    Gili, J.-M., Vervoort, W. & Pagès, F. (1989) Hydroids from the West African coast: Guinea Bissau, Namibia and South Africa. Scientia Marina, 53, 67–112.

    Goldfuss, G.A. (1818) Ueber die Classification der Zoophyten. Isis oder Encyclopädische Zeitung von Oken, 1818, 1008–1013.

    Govindarajan, A.F., Boero, F. & Halanych, K.M. (2006) Phylogenetic analysis with multiple markers indicates repeated loss of the adult medusa stage in Campanulariidae (Hydrozoa, Cnidaria). Molecular Phylogenetics and Evolution, 38, 820–834.

    https://doi.org/10.1016/j.ympev.2005.11.012

    Grange, L.J. & Smith, C.R. (2013) Megafaunal communities in rapidly warming fjords along the West Antarctic Peninsula: hotspots of abundance and beta diversity. PLoS ONE, 8 (11), e77917.

    https://doi.org/10.1371/journal.pone.0077917

    Grange, L.J., Smith, C.R., Lindsay, D.J, Bentlage, B. & Youngbluth, M.J. (2017) High abundance of the epibenthic trachymedusa Ptychogastria polaris Allman, 1878 (Hydrozoa, Trachylina) in subpolar fjords along the West Antarctic Peninsula. PLoS ONE, 12 (1), e0168648.

    https://doi.org/10.1371/journal.pone.0168648

    Grönberg, G. (1898) Die Hydroid-Medusen des arktischen Gebiets. Zoologische Jahrbücher, Abtheilung für Systematik, Geographie und Biologie der Thiere, 11, 451–468.

    Haeckel, E. (1866) Generelle morphologie der Organismen. Allgemeine Grundzüge der organischen Formen-Wissenschaft, mechanisch begründet durch die von Charles Darwin reformirte Descendenz-Theorie. Band 2. Georg Reimer, Berlin, LIX pp.

    https://doi.org/10.5962/bhl.title.3953

    Haeckel, E. (1879) Das System der Medusen. Erster Theil einer Monographie der Medusen. Denkschriften der Medicinisch-Naturwissenschaftlichen Gesellschaft zu Jena, 1, 1–360.

    Haeckel, E. (1882) Report on the deep-sea medusæ dredged by H.M.S. Challenger during the years 1873–1876. Report on the Scientific Results of the Voyage of H.M.S. Challenger during the Years 1873–76, Zoology, 4 (12), 1–154.

    Hartlaub, C. (1904) Hydroiden. Expédition Antarctique Belge. Résultats du Voyage du S. Y. Belgica en 1897–1898–1899. Rapports Scientifiques, Zoologie, 1–19.

    Hertwig, R. (1879) Der Organismus der Radiolarien. Gustav Fischer, Jena, 277 pp.

    Heywood, J., Kent, D., Schultz, J., Gibbs, D., Borden, L., Neale, M., Kulcsar, C., Banwait, R. & Trethewey, L. (2018) 2017 Arctic Marine Ecology Benchmarking Program. Final report. Ocean Wise Conservation Association, Vancouver, 45 pp.

    Hickson, S.J. & Gravely, F.H. (1907) Hydroid zoophytes. National Antarctic Expedition 1901–1904, Natural History 3, Coelentera 2, 1–34.

    Hincks, T. (1861) A catalogue of the zoophytes of south Devon and South Cornwall. Annals and Magazine of Natural History, Series 3, 8, 251–262.

    https://doi.org/10.1080/00222936108697413

    Hincks, T. (1869) A history of the British hydroid zoophytes. John van Voorst, London, 338 pp. [dating of this work follows Williams (2018), who discovered that it was published in March 1869 and not 1868 as per the title-pages]

    Hincks, T. (1874) Notes on Norwegian Hydroida from deep water. Annals and Magazine of Natural History, Series 4, 13, 125–137.

    https://doi.org/10.1080/00222937408680824

    Hirohito, The Showa Emperor (1984) A new hydroid Hydractinia bayeri n. sp. (family Hydractiniidae) from the Bay of Panama. Publications of the Biological Laboratory. Imperial Household, Tokyo, 8 pp.

    Hirohito, The Showa Emperor (1988) The hydroids of Sagami Bay. Part 1. Athecata. Biological Laboratory, Imperial Household, Tokyo, 179 pp.

    Hirohito, The Showa Emperor (1995) The hydroids of Sagami Bay. Part II. Thecata. Biological Laboratory, Imperial Household, Tokyo, 355 pp.

    Hodgson, M.M. (1950) A revision of the Tasmanian Hydroida. Papers and Proceedings of the Royal Society of Tasmania, 1949, 1–65.

    Ijima, I. (1896) Notice of new Hexactinellida from Sagami Bay. Zoologischer Anzeiger, 19, 249–254.

    Ijima, I. (1927) The Hexactinellida of the Siboga Expedition. Siboga-Expeditie, 6 , 1–383.

    Inaba, M. (1890) Hydroida obtained in Misaki, Miura, Soshu. Dobutsugaku Zasshi, 2, 143–149. [in Japanese]

    https://doi.org/10.4044/joma1889.2.4_143

    Inaba, M. (1892) Hydroida obtained in Shima. Dobutsugaku Zasshi, 4, 345–351. [in Japanese]

    https://doi.org/10.5026/jgeography.4.345

    International Commission on Zoological Nomenclature (1999) International code of zoological nomenclature. 4th Edition. International Trust for Zoological Nomenclature, London, 306 pp.

    Jäderholm, E. (1902) Die Hydroiden der schwedischen zoologischen Polarexpedition 1900. Bihang till Kongliga Svenska Vetenskaps-Akademiens Handlingar, 28, Afd 4, 12, 1–11.

    https://doi.org/10.5962/bhl.part.9812

    Jäderholm, E. (1907) Zur Kenntnis der Hydroidenfauna des Beringsmeeres. Arkiv för Zoologi, 4 (8), 1–8.

    Jäderholm, E. (1908) Die Hydroiden des sibirischen Eismeeres, gesammelt von der Russischen Polar-Expedition 1900–1903. Zapiski Imperatorskoï Akademii Nauk po Fiziko- Matematicheskomu Otdeleniyu, Series 8, 18 (12), 1–28.

    Jäderholm, E. (1909) Northern and Arctic invertebrates in the collection of the Swedish State Museum (Riksmuseum). IV. Hydroiden. Kungliga Svenska Vetenskapsakademiens Handlingar, Ny Följd, 45 (1), 1–124.

    Jäderholm, E. (1919) Zur Kenntnis der Hydroidenfauna Japans. Arkiv för Zoologi, 12 (9), 1–34.

    https://doi.org/10.5962/bhl.part.789

    Johnston, G. (1837) A catalogue of the zoophytes of Berwickshire. History of the Berwickshire Naturalists’ Club, 1, 107–108.

    Koreneva, E.V. (1980) 14. Palynological study of samples from sites 430, 431, 432, 433. Initial Reports of the Deep-Sea Drilling Project, 55, 409–413.

    https://doi.org/10.2973/dsdp.proc.55.114.1980

    Kosobokova, K.N., Hopcroft, R.R. & Hirche, H.-J. (2011) Patterns of zooplankton diversity through the depths of the Arctic’s central basins. Marine Biodiversity, 41, 29–50.

    https://doi.org/10.1007/s12526-010-0057-9

    Kramp, P.L. (1911) Report on the hydroids collected by the Danmark Expedition at north-east Greenland. Meddelelser om Grønland, 45, 341–396.

    Kramp, P.L. (1914) Meduser og siphonophorer. Catalogue of the medusae and Siphonophora of Greenland. Meddelelser om Grønland, 43, 381–456.

    Kramp, P.L. (1942) The Godthaab Expedition 1928. Medusae. Meddelelser om Grønland, 81 (1), 1–168. [not seen]

    Kramp, P.L. (1943) The zoology of East Greenland. Medusae, Siphonophora and Ctenophora. Meddelelser om Grønland, 121 (12), 1–20. [not seen]

    Kramp, P.L. (1947) Medusae. Part III. Trachylina and Scyphozoa, with zoogeographical remarks on all the medusae of the northern Atlantic. Danish Ingolf Expedition, 5 (14), 1–66.

    Kramp, P.L. (1955) A revision of Ernst Haeckel’s determinations of a collection of medusae belonging to the Zoological Museum of Copenhagen. Deep-Sea Research, 3 (Supplement), 149–168.

    Kramp, P.L. (1957) Hydromedusae from the Discovery Collections. Discovery Reports, 29, 1–128.

    https://doi.org/10.5962/bhl.part.12484

    Kramp, P.L. (1961) Synopsis of the medusae of the world. Journal of the Marine Biological Association of the United Kingdom, 40, 1–469.

    https://doi.org/10.1017/S0025315400007347

    Kühn, A. (1913) Entwicklungsgeschichte und Verwandtschaftsbeziehungen der Hydrozoen. I. Teil: Die Hydroiden. Ergebnisse und Fortschritte der Zoologie, 4, 1–284.

    Kussakin, O.G. & Kostina, E.E. (1996) The intertidal biota of volcanic Yankich Island (Middle Kuril Islands). Publications of the Seto Marine Biological Laboratory, 37, 201–225.

    https://doi.org/10.5134/176267

    Lamarck, J.B.P.A. de (1816) Histoire naturelle des animaux sans vertèbres. Tome 2. Verdière, Paris, 568 pp.

    Lamouroux, J.V.F. (1812) Extrait d’un mémoire sur la classification des polypiers coralligènes non entièrement pierreux. Nouveau Bulletin des Sciences, par la Société Philomatique de Paris, 3, 181–188.

    Lamouroux, J.V.F. (1821) Exposition méthodique des genres de l’ordre des polypiers, avec leur description et celle des principales espèces, figurées dans 84 planches; les 63 premières appartenant à l’histoire naturelle des zoophytes d’Ellis et Solander. Agasse, Paris, 115 pp.

    https://doi.org/10.5962/bhl.title.11328

    Larson, R.J., Matsumoto, G.I., Madin, L.P. & Lewis, L.M. (1992) Deep-sea benthic and benthopelagic medusae: recent observations from submersibles and a remotely operated vehicle. Bulletin of Marine Science, 51, 277–286.

    Leclère, L., Schuchert, P., Cruaud, C., Couloux, A. & Manuel, M. (2009) Molecular phylogenetics of Thecata (Hydrozoa, Cnidaria) reveals long-term maintenance of life history traits despite high frequency of recent character changes. Systematic Biology, 58, 509–526.

    https://doi.org/10.1093/sysbio/syp044

    Leloup, E. (1938) Quelques hydropolypes de la Baie de Sagami, Japon. Bulletin du Musée Royal d’Histoire Naturelle de Belgique, 14 (28), 1–22.

    Levinsen, G.M.R. (1893) Meduser, ctenophorer og hydroider fra Grønlands vestkyst, tilligemed bemærkninger om hydroidernes systematik. Videnskabelige Meddelelser fra Dansk Naturhistorisk Forening i Kjøbenhavn, Series 5, 4, 143–220.

    Linko, A.K. (1904) Zoologische Studien im Barents-Meere. Zoologischer Anzeiger, 28, 210–220.

    Linko, A.K. (1911) Gidroidy (Hydroidea). Tom I. Haleciidae, Lafoëidae, Bonneviellidae i Campanulariidae. Fauna Rossii i Supredelnykh Stran, 1, 1–250.

    https://doi.org/10.5962/bhl.title.46270

    Linnaeus, C. (1758) Systema naturae per regna tria naturae, secundum classes, ordines, genera, species cum characteribus, differentiis, synonymis, locis. Editio decima, reformata. Laurentii Salvii, Holmiae, 823 pp.

    https://doi.org/10.5962/bhl.title.542

    Lorenz, L. von (1886) Polypomedusen von Jan Mayen. Gesammelt von Dr. F. Fischer. Die Internationale Polarforschung 1882–1883. Die Österreichische Polarstation Jan Mayen, 3, 25–28.

    Maas, O. (1893) Die craspedoten Medusen der Plankton-Expedition. Ergebnisse der Plankton-Expedition der Humboldt-Stiftung, 2, K. c., 1–107.

    Maas O. (1905) Die Craspedoten Medusen der Siboga-Expeditie. Siboga-Expeditie. 10, 1–84.

    Maas, O. (1906) Die arktischen Medusen (ausschliesslich der Polypomedusen). Fauna Arctica, 4 (3), 479–526.

    Macgillivray, J. (1842) Catalogue of the marine zoophytes of the neighbourhood of Aberdeen. Annals and Magazine of Natural History, Series 1, 9 (60), 462–469.

    https://doi.org/10.1080/03745484209445365

    Mackie, G.O. (1985) Midwater macroplankton of British Columbia studied by submersible PISCES IV. Journal of Plankton Research, 7, 753–777.

    https://doi.org/10.1093/plankt/7.6.753

    Mammen, T.A. (1963) On a collection of hydroids from south India. I. Suborder Athecata. Journal of the Marine Biological Association of India, 5, 27–61.

    Marenzeller, E. von (1902) Südjapanische Anneliden. III. Aphroditea, Eunicea. Denkschriften der Kaiserlichen Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, 72, 563–582.

    Marktanner-Turneretscher, G. (1895) Zoologische Ergebnisse der im Jahre 1889 auf Kosten der Bremer geographischen Gesellschaft von Dr. Willy Kükenthal und Dr. Alfred Walter ausgeführten Expedition nach Ost-Spitzbergen. Hydroiden. Zoologische Jahrbücher, Abtheilung für Systematik, Geographie und Biologie der Thiere, 8, 391–438.

    Maronna, M.M., Miranda, T.P., Peña Cantero, Á.L., Barbeitos, M.S. & Marques, A.C. (2016) Towards a phylogenetic classification of Leptothecata (Cnidaria, Hydrozoa). Scientific Reports, 6, 18075.

    https://doi.org/10.1038/srep18075

    Martynov, A.V. (1914) Notice sur quelques formes nouvelles de trichoptères, provenant de différentes localités. Annuaire du Musée Zoologique de l’Académie Impériale des Sciences de St.-Pétersbourg, 19, 125–133. [in Russian and English]

    Mayer, A.G. (1910) Medusae of the world. Vol. II. The hydromedusae. Carnegie Institution of Washington Publication 109. Carnegie Institution of Washington, Washington, D.C. pp. 231–498.

    McCormick, J.M. (1965) Some aspects of the ecology of hydroids off the Oregon coast. Northwest Science, 39, 139–147.

    Miglietta, M.P. (2006) Hydractinia antonii sp. nov.: a new, partially calcified hydractiniid (Cnidaria: Hydrozoa: Hydractiniidae) from Alaska. Journal of the Marine Biological Association of the United Kingdom, 86, 993–996.

    https://doi.org/10.1017/S0025315406013968

    Millard, N.A.H. (1975) Monograph on the Hydroida of southern Africa. Annals of the South African Museum, 68, 1–513.

    Millard, N.A.H. (1977) The South African Museum’s Meiring Naude cruises. Part 3. Hydroida. Annals of the South African Museum, 73, 105–131.

    Miyake, H., Lindsay, D.J. & Kubota, S. (2004) Midwater and bentho-pelagic animals on the south slope of Shiribeshi Seamount off the west coast of Hokkaido. JAMSTEC Journal of Deep Sea Research, 24, 37–42.

    Motz-Kossowska, S. (1905) Contribution à la connaissance des hydraires de la Méditerranée occidentale. I.—Hydraires gymnoblastiques. Archives de Zoologie Expérimentale et Générale, 4e Série, 3, 39–98.

    Moura, C.J., Cunha, M.R., Porteiro, F.M. & Rogers, A.D. (2012) Polyphyly and cryptic diversity in the hydrozoan families Lafoeidae and Hebellidae (Cnidaria: Hydrozoa). Invertebrate Systematics, 25, 454–470.

    https://doi.org/10.1071/IS11045

    Moura, C.J., Harris, D.J., Cunha, M.R. & Rogers, A.D. (2008) DNA barcoding reveals cryptic diversity in marine hydroids (Cnidaria, Hydrozoa) from coastal and deep-sea environments. Zoologica Scripta, 37, 93–108.

    https://doi.org/10.1111/j.1463-6409.2007.00312.x

    Namikawa, H. (2009) Preliminary report on deep water hydrozoan species collected from the Pacific Ocean off northern Japan. In: Fujita, T. (Ed.), Deep-sea fauna and pollutants off Pacific coast of northern Japan. National Museum of Nature and Science Monographs, 39, pp. 19–23.

    Naumov, D.V. (1951) Nekotorye osobennosti anatomii semeistva Bonneviellidae v svyazi s voprosom o pervichnom pokolenii u gidrozoev. Doklady Akademii Nauk SSSR, 81, 1151–1154. [not seen]

    Naumov, D.V. (1960) Gidroidy i gidromeduzy morskikh, solonovatovodnykh i presnovodnykh basseinov SSSR. Akademiya Nauk SSSR, Opredeliteli po Faune SSSR, 70, 1–626. [in Russian]

    Naumov, D.V. (1966) Phylum Coelenterata. In: Pavlovskii, E.N. (Ed.), Atlas of the invertebrates of the Far Eastern Seas of the USSR. Academy of Sciences of the U.S.S.R., Zoological Institute, Israel Program for Scientific Translations No. 1672. Zoological Institute, Academy of Sciences of the U.S.S.R., Jerusalem, pp. 70–95.

    Nutting, C.C. (1901) Papers from the Harriman Alaska Expedition. XXI. The hydroids. Proceedings of the Washington Academy of Sciences, 3, 157–216.

    Nutting, C.C. (1905) Hydroids of the Hawaiian Islands collected by the steamer Albatross in 1902. Bulletin of the United States Fish Commission for 1903, 1905, pp. 931–959. [dating of this article follows Calder (2017: 68)]

    Nutting, C.C. (1915) American hydroids. Part III. The Campanularidae and the Bonneviellidae. Smithsonian Institution, United States National Museum Special Bulletin, 4 (3), 1–126.

    Oken, L. (1815) Okens Lehrbuch der Naturgeschichte. III. Theil. Zoologie. Volume 1. Oken, Jena, 842 pp.

    Owen, R. (1841) Description of a new genus and species of sponge (Euplectella aspergillum). Proceedings of the Zoological Society of London, 9, 3–5.

    Owen, R. (1843) Lectures on the comparative anatomy and physiology of the invertebrate animals: delivered at the Royal College of Surgeons, in 1843. Longman, Brown, Green, & Longmans, London, 392 pp.

    Palerud, R., Gulliksen, B., Brattegard, T. Sneli, J.-A. & Vader, W. (2004) The marine macro-organisms in Svalbard waters. In: Prestrud, R., Strøm, H. & Goldman, H.V. (Eds.), A catalogue of the terrestrial and marine animals of Svalbard. Norsk Polarinstitutt Skrifter, 201, 5–56.

    Panteleeva, N.N., Frolova, E.A. & Sheiko, O.V. (1999) New records of the benthic medusa Ptychogastria polaris Allman, 1878 (Trachylida, Hydroidea) in the Barents Sea and off the Kurile Islands (Pacific Ocean). Polar Biology, 22, 372–378.

    https://doi.org/10.1007/s003000050431

    Park, J.H. (1990) Systematic study on the marine hydroids (Cnidaria, Hydrozoa) in Korea I. Korean Journal of Systematic Zoology, 6, 71–86.

    Park, J.H. (1991) Systematic study on the marine hydroids (Cnidaria, Hydrozoa) in Korea II. The families Sphaerocorynidae, Eudendriidae, Haleciidae and Lafoëidae. Korean Journal of Zoology, 34, 541–547.

    Park, J.H. (1993) Marine hydroids (Cnidaria: Hydrozoa: Hydroida) from Cheju Island, Korea. Korean Journal of Systematic Zoology, 9, 261–280.

    Park, J.H. & Song, J.-I. (2000) Hydroids (Cnidaria: Hydrozoa) of Dokdo Islands, Korea. Korean Journal of Systematic Zoology, 16, 55–63.

    Peña Cantero, Á.L. (2012) Filling biodiversity gaps: benthic hydroids from the Bellingshausen Sea (Antarctica). Polar Biology, 35, 851–865.

    https://doi.org/10.1007/s00300-011-1130-y

    Peña Cantero, Á.L. (2019) Benthic hydroids off Scott Island and the shelf and slope of the Ross Sea (Antarctica) collected during the IPY-CAML TAN0802 voyage by R/V Tangaroa. Marine Biodiversity, 49, 863–885.

    https://doi.org/10.1007/s12526-018-0865-x

    Peña Cantero, Á.L., Boero, F. & Piraino, S. (2013) Shallow-water benthic hydroids from Tethys Bay (Terra Nova Bay, Ross Sea, Antarctica). Polar Biology, 36, 731–753.

    https://doi.org/10.1007/s00300-013-1299-3

    Peña Cantero, Á.L. & Horton, T. (2017) Benthic hydroids (Cnidaria, Hydrozoa) from bathyal and abyssal depths of the northeast Atlantic held in the modern Discovery Collections. Zootaxa, 4347 (1), 1–30.

    https://doi.org/10.11646/zootaxa.4347.1.1

    Peña Cantero, Á.L., Sentandreu, V. & Latorre, A. (2010) Phylogenetic relationships of the endemic Antarctic benthic hydroids (Cnidaria, Hydrozoa): what does the mitochondrial 16S rRNA tell us about it? Polar Biology, 33, 41–57.

    https://doi.org/10.1007/s00300-009-0683-5

    Pennycuik, P.R. (1959) Faunistic records from Queensland. Part V.—Marine and brackish water hydroids. Papers from the Department of Zoology, University of Queensland, 1, 141–210.

    Pequegnat, W.E. (1964) The epifauna of a California siltstone reef. Ecology, 45, 272–283.

    https://doi.org/10.2307/1933840

    Petersen, K.W. (1990) Evolution and taxonomy in capitate hydroids and medusae (Cnidaria: Hydrozoa). Zoological Journal of the Linnean Society, 100, 101–231.

    https://doi.org/10.1111/j.1096-3642.1990.tb01862.x

    Piepenburg, D., Chernova, N.V., von Dorrien, C.F., Gutt, J., Neyelov, A.V., Rachor, E., Saldanha, L. & Schmid, M.K. (1996) Megabenthic communities in the waters around Svalbard. Polar Biology, 16, 431–446.

    https://doi.org/10.1007/BF02390425

    Prokopchuk, I.P. & Trofimov, A.G. (2019) Interannual dynamics of zooplankton in the Kola Section of the Barents Sea during the recent warming period. ICES Journal of Marine Science, 76 (Supplement 1), i10–i23.

    https://doi.org/10.1093/icesjms/fsz206

    Proudfoot, B., Devillers, R., Brown, C.J., Edinger, E. & Copeland, A. (2020) Seafloor mapping to support conservation planning in an ecologically unique fjord in Newfoundland and Labrador, Canada. Journal of Coastal Conservation, 24, 36, 1–17.

    Prudkovsky, A.A., Nikitin, M.A., Berumen, M.L., Ivanenko, V.N. & Reimer, J.D. (2016) On the paraphyly of Cytaeididae and placement of Cytaeis within the suborder Filifera (Hydrozoa: Anthoathecata). Marine Biodiversity, 18 June 2016, 1–8.

    https://doi.org/10.1007/s12526-016-0534-x

    Puce, S., Calcinai, B., Bavestrello, G., Cerrano, C., Gravili, C. & Boero, F. (2005) Hydrozoa (Cnidaria) symbiotic with Porifera: a review. Marine Ecology, 26, 73–81.

    https://doi.org/10.1111/j.1439-0485.2005.00050.x

    Ramil, F. & Vervoort, W. (1992) Report on the Hydroida collected by the “BALGIM” expedition in and around the Strait of Gibraltar. Zoologische Verhandelingen, Leiden, 277, 1–262.

    Rees, W.J. (1953) Records of hydroids and medusae taken at Herdla, Bergen in 1937. Universitetet i Bergen Årbok 1952. Naturvitenskapelig Rekke, 16, 1–8.

    Regelous, M., Hofmann, A.W., Abouchami, W. & Galer, S.J.G. (2003) Geochemistry of lavas from the Emperor Seamounts, and the geochemical evolution of Hawaiian magmatism from 85 to 42 Ma. Journal of Petrology, 44, 113–140.

    https://doi.org/10.1093/petrology/44.1.113

    Reiswig, H.M. & Kelly, M. (2018) The marine fauna of New Zealand. Euplectellid glass sponges (Hexactinellida, Lyssacinosida, Euplectellidae). National Institute of Water and Atmospheric Research Biodiversity Memoir, 130, 1–170.

    Rho, B.J. & Chang, S.R. (1974) On the classification and the distribution of the marine benthic animals in Korea. I. Hydroids. Journal of Korean Research Institute for Better Living, Ewha Womans University, 12, 133–158.

    Ritchie, J. (1907) The hydroids of the Scottish National Antarctic Expedition. Transactions of the Royal Society of Edinburgh, 45, 519–545.

    https://doi.org/10.1017/S0080456800022821

    Ritchie, J. (1909) Supplementary report on the hydroids of the Scottish National Antarctic Expedition. Transactions of the Royal Society of Edinburgh, 47, 65–101.

    https://doi.org/10.1017/S0080456800011881

    Ronowicz, M. & Schuchert, P. (2007) Halecium arcticum (Cnidaria: Hydrozoa), a new species of hydroid from Spitzbergen. Zootaxa, 1549 (1), 55–62.

    https://doi.org/10.11646/zootaxa.1549.1.4

    Russell, F.S. (1940) On the nematocysts of hydromedusae. III. Journal of the Marine Biological Association of the United Kingdom, 24, 515–523.

    https://doi.org/10.1017/S0025315400045422

    Ruthensteiner, B., Reinicke, G.-B. & Straube, N. (2008) The type material of Hydrozoa described by Eberhard Stechow in the Zoologische Staatssammlung München. Spixiana, 31, 3–27.

    Rybakova, E., Galkin, S., Gebruk, A., Sanamyan, N. & Martynov, A. (2020) Vertical distribution of megafauna on the Bering Sea slope based on ROV survey. PeerJ, 8, e8628

    https://doi.org/10.7717/peerj.8628

    Rybakova, E., Kremenetskaia, A., Vedenin, A., Boetius, A. & Gebruk, A. (2019) Deep-sea megabenthos communities of the Eurasian Central Arctic are influenced by ice-cover and sea-ice algal falls. PLoS ONE, 14 (7), e0211009.

    https://doi.org/10.1371/journal.pone.0211009

    Samyn, Y. (2014) Return to sender: Hydrozoa collected by Emperor Hirohito of Japan in the 1930s and studied in Brussels. Archives of Natural History, 41, 17–24.

    https://doi.org/10.3366/anh.2014.0207

    Sanamyan, K. & Sanamyan, N. (2020) Флора и фауна острова Матуа (средние Курильские острова) Атлас-определитель Том 1. Море. Русское географическое общество Камчатский филиал Тихоокеанского института географии ДВО РАН, Камчатский, 495 pp. [Flora and fauna of Matua island (middle Kuril Islands). Identifier Atlas. Vol. 1. Sea. Russian Geographical Society, Kamchatka Branch of the Pacific Institute of Geography FEB RAS, Kamchatka]

    Sars, G.O. (1874) Bidrag til kundskaben om norges hydroider. Forhandlinger i Videnskabs-Selskabet i Christiania, 1873, 91–150.

    Sars, M. (1850) Beretning om en i Sommeren 1849 foretagen zoologisk Reise i Lofoten og Finmarken. Nyt Magazin for Naturvidenskaberne, 6, 121–211.

    Schuchert, P. (2001) Hydroids of Greenland and Iceland (Cnidaria, Hydrozoa). Meddelelser om Grønland, Bioscience, 53, 1–184.

    Schuchert, P. (2007) The European athecate hydroids and their medusae (Hydrozoa, Cnidaria): Filifera Part 2. Revue Suisse de Zoologie, 114, 195–396.

    https://doi.org/10.5962/bhl.part.80395

    Schuchert, P. (2012) North-west European athecate hydroids and their medusae. Synopses of the British Fauna, New Series, 59, 1–364.

    Schuchert, P. (2021) World Hydrozoa Database. Accessed through: World Register of Marine Species at: https://www.marinespecies.org/aphia.php?p=taxdetails&id=117001 (accessed 11 November 2020)

    Schuchert, P. & Reiswig, H.M. (2006) Brinckmannia hexactinellidophila, n. gen., n. sp.: a hydroid living in tissues of glass sponges of the reefs, fjords and seamounts of Pacific Canada and Alaska. Canadian Journal of Zoology, 84, 564–572.

    https://doi.org/10.1139/z06-031

    Schuchert, P., Sanamyan, N. & Sanamyan, K. (2016) Observations on two large athecate hydroids (Cnidaria: Hydrozoa) from the Kamchatka Peninsula (NW Pacific). Revue Suisse de Zoologie, 123, 165–178.

    Schulze, F.E. (1880) On the structure and arrangement of the soft parts in Euplectella aspergillum. Transactions of the Royal Society of Edinburgh, 29, 661–673.

    https://doi.org/10.1017/S0080456800026181

    Schulze, F.E. (1886) Über den Bau und das System der Hexactinelliden. Abhandlungen der Königlichen Akademie der Wissenschaften zu Berlin, Physikalisch-Mathematische Classe, 1886, 1–97.

    Schulze, F.E. (1887) Report on the Hexactinellida collected by H.M.S. Challenger during the years 1873–76. Report on the Scientific Results of the Voyage of H.M.S. Challenger during the years 1873–76, Zoology, 21 (53), 1–514.

    Segonzac, M. & Vervoort, W. (1995) First record of the genus Candelabrum (Cnidaria, Hydrozoa, Athecata) from the Mid-Atlantic Ridge: a description of a new species and a review of the genus. Bulletin du Museum National d’Histoire Naturelle, Paris, Série 4, 17, 31–64.

    Sheiko, O. & Stepanjants, S. (1997) Medusozoa (Cnidaria: Anthozoa excepted) from the Commander Islands, faunistic composition and biogeography. Proceedings of the 6th International Conference on Coelenterate Biology, 1995, 437–445.

    Sirenko, B.I. (2001) List of species of free-living invertebrates of Eurasian Arctic seas and adjacent deep waters. Russian Academy of Sciences, Zoological Institute, Explorations of the Fauna of the Seas, 51 (59), 1–131.

    Sirenko, B.I., Markhaseva, E.L., Buzhinskaya, G.N., Golikov, A.A., Menshutkina, T.V., Petryashov, V.V., Semenova, T.N., Stepanjants, S.D. & Vassilenko, S.V. (1996) Preliminary data on suprabenthic invertebrates collected during the RV Polarstern cruise in the Laptev Sea. Polar Biology, 16, 345–352.

    https://doi.org/10.1007/BF02342182

    Spassky, N. (1929) K faune gidroidov Kol’skogo zaliva i yugo-zapadnoǐ chasti Barentseva morya. Raboty Murmanskoi Biologicheskoi Stantsii, 3, 1–48. [in Russian]

    Sswat, M., Gulliksen, B., Menn, I., Sweetman, A.K. & Piepenburg, D. (2015) Distribution and composition of the epibenthic megafauna north of Svalbard (Arctic). Polar Biology, 38, 861–877.

    https://doi.org/10.1007/s00300-015-1645-8

    Starmans, A. (1997) Vergleichende Untersuchungen zur Ökologie und Biodiversität des Mega-Epibenthos der Arktis und Antarktis. Berichte zur Polarforschung, 250, 1–150.

    Stechow, E. (1907) Neue japanische Athecata und Plumularidae aus der Sammlung Dr. Doflein. Zoologischer Anzeiger, 32, 192–200.

    Stechow, E. (1913a) Neue Genera thecater Hydroiden aus der Familie der Lafoeiden und neue Species von Thecaten aus Japan. Zoologischer Anzeiger, 43, 137–144. [published 02 December 1913 (Vervoort 1995)]

    Stechow, E. (1913b) Hydroidpolypen der japanischen Ostküste. II. Teil: Campanularidae, Halecidae, Lafoeidae, Campanulinidae und Sertularidae, nebst Ergänzungen zu den Athecata und Plumularidae. Abhandlungen der Mathematisch-Physikalischen Klasse der Königlichen Bayerischen Akademie der Wissenschaften, 3 (Supplement-Band 2), 1–162.

    https://doi.org/10.5962/bhl.title.11621

    Stechow, E. (1919) Zur Kenntnis der Hydroidenfauna des Mittelmeeres, Amerikas und anderer Gebiete, nebst Angaben über einige Kirchenpauer’sche Typen von Plumulariden. Zoologische Jahrbücher, Abteilung für Systematik, Geographie und Biologie der Tiere, 42, 1–172.

    Stechow, E. (1921a) Neue Gruppen skelettbildender Hydrozoen und Verwandtschaftsbeziehungen rezenter und fossiler Formen. Verhandlungen der Deutschen Zoologischen Gesellschaft, 26, 29–31.

    Stechow, E. (1921b) Neue Genera und Species von Hydrozoen und anderen Evertebraten. Archiv für Naturgeschichte, Abteilung A, 3 (87), 248–265.

    Stechow, E. (1923a) Die Hydroidenfauna der Japanischen Region. Journal of the College of Science, Imperial University of Tokyo, 44 (8), 1–23.

    Stechow, E. (1923b) Über Hydroiden der Deutschen Tiefsee-Expedition, nebst Bemerkungen über einige andre Formen. Zoologischer Anzeiger, 56, 97–119.

    Stechow, E. (1923c) Zur Kenntnis der Hydroidenfauna des Mittelmeeres, Amerikas und anderer Gebiete. II. Teil. Zoologische Jahrbücher, Abteilung für Systematik, Ökologie und Geographie der Tiere, 47, 29–270.

    Stechow, E. (1925) Hydroiden der Deutschen Tiefsee-Expedition. Wissenschaftliche Ergebnisse der Deutschen Tiefsee-Expedition auf dem Dampfer „Valdivia” 1898–1899, 17, 383–546.

    Stechow, E. (1962) Über skelettbildende Hydrozoen. Zoologischer Anzeiger, 169, 416–428.

    Stechow, E. & Uchida, T. (1931) Report of the Biological Survey of Mutsu Bay. 21. Hydroiden von Mutsu-Bai, Nord-Japan. Science Reports of the Tôhoku Imperial University, Series 4 (Biology), 6, 545–571.

    Stepanjants, S.D. (1979) Gidroidy vod Antarktiki i Subantarktiki. Rezultaty biologicheskikh Issledovaniy Sovetskikh Antarkticheskikh Ekspeditsiy, 6. Issledovaniya Fauny Morei, 22 (30), 1–199. [in Russian]

    Stepanjants, S. (1989) Hydrozoa of the Eurasian Arctic seas. In: Herman, Y. (Ed.), The Arctic seas. Climatology, oceanography, geology, and biology. Van Nostrand Reinhold Co., New York, pp. 397–430.

    https://doi.org/10.1007/978-1-4613-0677-1_16

    Stepanjants, S.D. (2013a) Deep-water Hydrozoa (Cnidaria: Medusozoa) in the Sea of Japan, collected during the 51st cruise of R/V Akademik M.A. Lavrentyev, with description Opercularella angenikae, sp. nov. Deep-Sea Research II, 86–87, 231–237.

    https://doi.org/10.1016/j.dsr2.2012.08.014

    Stepanjants, S.D. (2013b) Class Hydrozoa. In: Sirenko, B.I. (Ed.) Check-list of species of free-living invertebrates of the Russian Far Eastern seas. Russian Academy of Sciences, Zoological Institute, Explorations of the Fauna of the Seas, 75 (83), pp. 38–43.

    Stepanjants, S.D. & Chernyshev, A.V. (2015) Deep-sea epibiotic hydroids from the abyssal plain adjacent to the Kuril–Kamchatka Trench with description of Garveia belyaevi sp. nov. (Hydrozoa, Bougainvilliidae). Deep-Sea Research II, 111, 44–48.

    https://doi.org/10.1016/j.dsr2.2014.07.015

    Stepanjants, S.D., Cortese, G., Kruglikova, S.B. & Bjørklund, K.R. (2006) A review of bipolarity concepts: history and examples from Radiolaria and Medusozoa (Cnidaria). Marine Biology Research, 2, 200–241.

    https://doi.org/10.1080/17451000600781767

    Stimpson, W. (1858) Prodromus descriptionis animalium evertebratorum, quae in Expeditione ad Oceanum Pacificum Septentrionalem, a Republica Federata missa, Cadwaladaro Ringgold et Johanne Rogers Ducibus, observavit et descripsit. Pars VII. Crustacea Anomoura. Proceedings of the Academy of Natural Sciences of Philadelphia, 10, 225–252.

    https://doi.org/10.5962/bhl.title.51447

    Stimpson, W. (1907) Report on the Crustacea (Brachyura and Anomura) collected by the North Pacific Exploring Expedition, 1853–1856. Smithsonian Miscellaneous Collections, 49 (3), 5–240.

    https://doi.org/10.5962/bhl.title.51448

    Stübing, D. & Piepenburg, D. (1998) Occurrence of the benthic trachymedusa Ptychogastria polaris Allman, 1878 (Cnidaria: Hydrozoa) off northeast Greenland and in the northern Barents Sea. Polar Biology, 19, 193–197.

    https://doi.org/10.1007/s003000050234