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Type: Article
Published: 2021-03-09
Page range: 415–424
Abstract views: 101
PDF downloaded: 10

Changes in the genus Beana Walker (Beaninae Zahiri & Holloway) with two new species and removing Beana nitida Tams to a new genus Beanoides (gen. nov.)

175, M1 Muban Phichai, Mueang Lampang, Lampang, 52000 THAILAND.
Department of Biology, Sölvegatan 37, Lund University, SE-223 62 Lund, SWEDEN.


Two new Beana species (B. mutaneni sp. nov and B. penniuncus sp. nov) are described here. Based on molecular data, Beana nitida Tams is removed to its own genus Beanoides gen. nov. and placed in the subfamily Chloephorinae. Morphological characteristics and DNA barcodes are provided for the new species, with a comparison of morphological structures and genetic distances to Beana terminigera Walker. The current number of valid species in the genus is six.



  1. Guindon, S., Dufayard, J.F., Lefort, V., Anisimova, M., Hordijk, W. & Gascuel, O. (2010) New algorithms and methods to estimate maximum-likelihood phylogenies: assessing the performance of PhyML 3.0. Systematic biology, 59 (3), 307–321.

    Hadley, A. (2012) CombineZP, released 6 June 2010, open-source image processing software for creating extended depth of field images. Available from: (accessed 9 May 2017)

    Hoang, D.T., Chernomor, O., von Haeseler, A., Minh, B.Q. & Vinh, L.S. (2018) UFBoot2: improving the ultrafast bootstrap approximation. Molecular Biology and Evolution, 35 (2), 518–522.

    Holloway, J.D. (2003) The Moths of Borneo, Nolidae. Part 18. Southdene Sdn. Bhd., Kuala Lumpur, 279 pp.

    Holloway, J.D. & Miller, S.E. (1995) Status of Rhyncopalpus brunellus in the Hawaiian Islands, with comments on the systematics of the Nolinae (Lepidoptera: Noctuidae). Bishop Museum Occasional Papers, 42, 31–34.

    Keegan, K., Lafontaine, J.D., Wahlberg, N. & Wagner, D. (2019) Towards resolving and redefining Amphipyrinae: a massively polyphyletic taxon (Lepidoptera, Noctuoidea, Noctuidae). Systematic Entomology, 44, 451–464.

    Klots, A.B. (1970) Lepidoptera. In: Tuxen, S.L. (Ed.), Taxonomists‘ glossary of genitalia in insects. Munksgaard, Copenhagen, pp. 115–130.

    Kononenko, V.S. & Pinratana, A. (2013) Moth of Thailand. Vol. 3. Part 2. Noctuoidea. An illustrated Catalogue of Erebidae, Nolidae, Euteliidae and Noctuidae (Insecta, Lepidoptera) in Thailand. Brothers of St. Gabriel, Bangkok, 625 pp.

    Mutanen, M., Kivelä, S.M., Vos, R.A., Doorenweerd, C., Ratnasingham, S., Hausmann, A., Huemer, P., Dinca, V., Nieukerken, E.J. van, Lopez-Vaamonde, C., Vila, R., Aarvik, L., Decaëns, T., Efetov, K.A., Hebert, P.D.N., Johnsen, A., Karsholt, O., Pentinsaari, M., Rougerie, R., Segerer, A., Tarmann, G., Zahiri, R. & Godfray, H.C.J. (2016) Species-level para- and polyphyly in DNA barcode gene trees: strong operational bias in European Lepidoptera. Systematic Biology, 65, 1024–1040.

    Nguyen, L.-T., Schmidt, H.A., von Haeseler, A. & Minh, B.Q. (2015) IQ-TREE: A fast and effective stochastic algorithm for estimating maximum likelihood phylogenies. Molecular Biology and Evolution, 32, 268–274.

    Pellinen, M.J. & Wahlberg, N. (2015) A new species of Niganda Moore, 1879 from Thailand, with description of variation in male genitalia and female facies of N. radialis Moore (Lepidoptera: Notodontidae, Ceirinae). Zootaxa, 4033 (1), 94–102.

    Tams, W.H.T. (1924) List of the moth, collected in Siam by E. J. Godfrey B. SC., F. E. S. Journal of theNatural History Society of Siam, 6, 229–289.

    Wahlberg, N. & Wheat, C.W. (2008) Genomic Outposts Serve the Phylogenomic Pioneers: Designing Novel Nuclear Markers for Genomic DNA Extractions of Lepidoptera. Systematic Biology, 57 (2), 231–242.

    Walker, F. (1862) Catalogue of the Heterocerous Lepidopterous insects collected at Sarawak, in Borneo, by Mr A. R. Wallace, with descriptions of new species. Journal of the proceedings of the Linnean Society, Zoology, 6, 183.

    Zahiri, R., Holloway, J.D., Kitching, I.J., Lafontaine, D., Mutanen, M. & Wahlberg, N. (2012) Molecular phylogenetics of Erebidae (Lepidoptera, Noctuoidea). Systematic Entomology, 37, 102–124.

    Zahiri, R., Kitching, I.J., Lafontaine, J.D., Mutanen, M., Kaila, L., Holloway, J.D. & Wahlberg, N. (2011) A new molecular phylogeny offers hope for a stable family-level classification of the Noctuoidea (Lepidoptera). Zoologica Scripta, 40, 158–173.

    Zahiri, R., Lafontaine, J.D., Holloway, J.D., Kitching, I.J., Schmidt, B.C., Kaila, L. & Wahlberg, N. (2013) Major lineages of Nolidae (Lepidoptera, Noctuoidea) elucidated by molecular phylogenetics. Cladistics, 29, 337–359.