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Published: 2020-12-17
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Catalogue of rhysodine beetles (Coleoptera: Carabidae) from East Asia

Engineering Research Center for Forest and Grassland Disaster Prevention and Reduction, Mianyang Normal University, 166 Mianxing West Road, Mianyang 621000, Sichuan Province, P. R. China.
Coleoptera Rhysodinae bibliography diversity distribution key

Abstract

An annotated catalogue for 22 extant species of the subfamily Rhysodinae (Coleoptera: Carabidae) from East Asia is presented, with 7 genera classified into 3 tribes: I. Clinidiini Bell & Bell, 1978: Clinidium Kirby, 1830 (1 sp.) and Rhyzodiastes Fairmaire, 1895 (4 spp.); II. Omoglymmiini Bell & Bell, 1978: Arrowina Bell & Bell, 1978 (1 sp.), Yamatosa Bell & Bell, 1979 (6 spp.), Shyrodes Grouvelle, 1903 (1 sp.), and Omoglymmius Ganglbauer, 1891 (8 spp.); III. Rhysodini Laporte, 1840: Rhysodes Germar, 1822 (1 sp.). For each genus- and species-group name, synonymy, type locality, type depository, distribution, and bibliography are provided where appropriate. A modified key to genera of rhysodine beetles from East Asia is compiled.

 

References

  1. Agassiz, J.L.R. (1848) Nomenclatoris zoologici. Index universalis, continens nomina systematica classium, ordinum, familiarum et generum animalium omnium, tam viventium quam fossilium, secundum ordinem alphabeticum unicum disposita, adjectis homonymiis plantarum. Jent et Gassmann, Soloduri, p. 943.

    https://doi.org/10.5962/bhl.title.1819

    Arnett, R.H. Jr. & Ivie, M.A. (2000) 5. Rhysodidae Laporte, 1840. In: Arnett, R.H., Jr. & Thomas, M.C. (Eds.), American Beetles. Volume 1. Archostemata, Myxophaga, Adephaga, Polyphaga: Staphyliniformia. CRC Press, Boca Raton, London, New York and Washington, pp. 30–31.

    https://doi.org/10.1201/9781482274325

    Arrow, G.J. (1901) Remarks upon the genus Rhysodes, with descriptions of some new Oriental species. Annals and Magazine of Natural History, Series 7, 7 (37), 83–89.

    https://doi.org/10.1080/00222930108678444

    Arrow, G.J. (1942) The beetle family Rhysodidae, with some new species and a key to those at present known. The Proceedings of the Royal Entomological Society of London, Series B, Taxonomy, 11, 171–183.

    https://doi.org/10.1111/j.1365-3113.1942.tb00720.x

    Baca, S.M., Alexander, A., Gustafson, G.T. & Short, A.E.Z. (2017) Ultraconserved elements show utility in phylogenetic inference of Adephaga (Coleoptera) and suggest paraphyly of ‘Hydradephaga’. Systematic Entomology, 42 (4), 786–795.

    https://doi.org/10.1111/syen.12244

    Baehr, M. (1979) Vergleichende Untersuchungen am Skelett und an der Coxalmuskulatur des Prothorax der Coleoptera, ein Beitrag zur Klärung der phylogenetischen Beziehungen der Adephaga (Coleoptera, Insecta). Zoologica, 44 (130), 1–76.

    Baehr, M. & Will, K. (2019) 12. Carabidae Latreille, 1802. In: Ślipiński, S.A. & Lawrence, J.F. (Eds.), Australian Beetles. Volume 2. Archostemata, Myxophaga, Adephaga, Polyphaga (part). CSIRO Publishing, Clayton South, Victoria, pp. 61–217.

    Bell, R.T. (1967) Coxal cavities and the classification of the Adephaga (Coleoptera). Annals of the Entomological Society of America, 60 (1), 101–107.

    https://doi.org/10.1093/aesa/60.1.101

    Bell, R.T. (1970) The Rhysodini of North America, Central America, and the West Indies (Coleoptera: Carabidae or Rhysodidae). Miscellaneous Publications of the Entomological Society of America, 6 (6), 289–324.

    Bell, R.T. (1975) Omoglymmius Ganglbauer, a separate genus (Coleoptera: Carabidae or Rhysodidae). The Coleopterists Bulletin, 29 (4), 351–352.

    Bell, R.T. (1977) Ergebnisse der Bhutan-Expedition 1972 des Naturhistorischen Museums in Basel. Coleoptera: Fam. Rhysodidae. Entomologica Basiliensia, 2, 151–158.

    Bell, R.T. (1991) Rhysodidae (Adephaga). In: Stehr, F.W. (Ed.), Immature Insects. Volume 2. Kendall/Hunt Publishing Company, Dubuque, Iowa, pp. 304–305.

    Bell, R.T. (1998) Where do the Rhysodini (Coleoptera) belong? In: Ball, G.E., Casale, A. & Vigna Taglianti, A. (Eds.), Phylogeny and Classification of Caraboidea (Coleoptera: Adephaga). Proceedings of a Symposium, Florence, Italy, 28 August, 1996. XX International Congress of Entomology. Museo Regionale di Scienze Naturali, Torino, pp. 261–272.

    Bell, R.T. (1999) Rhysodini. Wrinkled Bark Beetles. Version 14 December 1999. The Tree of Life Web Project. Available from: http://tolweb.org/Rhysodini/67/1999.12.14 (20 October 2019)

    Bell, R.T. (2003) Family Rhysodidae Laporte, 1840. In: Löbl, I. & Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 1. Archostemata—Myxophaga—Adephaga. Apollo Books, Stenstrup, p. 78.

    Bell, R.T. & Bell, J.R. (1962) The taxonomic position of the Rhysodidae. The Coleopterists Bulletin, 16 (4), 99–106.

    Bell, R.T. & Bell, J.R. (1975) Two new taxa of Clinidium (Coleoptera: Rhysodidae or Carabidae) from the eastern U.S., with a revised key to U.S. Clinidium. The Coleopterists Bulletin, 29 (2), 65–68.

    Bell, R.T. & Bell, J.R. (1978) Rhysodini of the world. Part I. A new classification of the tribe, and a synopsis of Omoglymmius subgenus Nitiglymmius, new subgenus (Coleoptera: Carabidae or Rhysodidae). Quaestiones Entomologicae, 14 (1), 43–88.

    Bell, R.T. & Bell, J.R. (1979) Rhysodini of the world. Part II. Revisions of the smaller genera (Coleoptera: Carabidae or Rhysodidae). Quaestiones Entomologicae, 15 (4), 377–446.

    Bell, R.T. & Bell, J.R. (1981) Insects of Micronesia. Coleoptera: Rhysodidae. Bernice P. Bishop Museum Insects of Micronesia, 15 (2), 51–67.

    Bell, R.T. & Bell, J.R. (1982) Rhysodini of the world. Part III. Revision of Omoglymmius Ganglbauer (Coleoptera: Carabidae or Rhysodidae) and substitutions for preoccupied generic names. Quaestiones Entomologicae, 18 (1–4), 127–259.

    Bell, R.T. & Bell, J.R. (1985) Rhysodini of the world. Part IV. Revisions of Rhyzodiastes Fairmaire and Clinidium Kirby, with new species in other genera (Coleoptera: Carabidae or Rhysodidae). Quaestiones Entomologicae, 21 (1), 1–172.

    Bell, R.T. & Bell, J.R. (1987) Rhysodine beetles in the Geneva collection: a new species of Yamatosa, and a major range extension for Omoglymmius sakuraii Nakane (Coleoptera: Carabidae or Rhysodidae). Revue suisse de Zoologie, 94 (4), 683–686.

    https://doi.org/10.5962/bhl.part.79543

    Bell, R.T. & Bell, J.R. (1989) Rhysodine beetles in the Geneva collection II: new species of Yamatosa and Omoglymmius, descriptions of undescribed sexes in other species, and some major range extensions (Coleoptera: Carabidae or Rhysodidae). Revue suisse de Zoologie, 96 (3), 637–642.

    https://doi.org/10.5962/bhl.part.82053

    Bell, R.T. & Bell, J.R. (1991) The Rhysodini of Australia (Insecta: Coleoptera: Carabidae or Rhysodidae). Annals of Carnegie Museum, 60 (3), 179–210.

    Bell, R.T. & Bell, J.R. (1993) Rhysodine beetles (Insecta: Coleoptera: Carabidae or Rhysodidae): new species, new data, and revised keys to Omoglymmius (subgenera Omoglymmius and Pyxiglymmius). Annals of Carnegie Museum, 62 (2), 165–185.

    Bell, R.T. & Bell, J.R. (2000) Rhysodine beetles (Insecta: Coleoptera: Carabidae): new species, new data. II. Annals of Carnegie Museum, 69 (2), 69–91.

    Bell, R.T. & Bell, J.R. (2002) Two new species of Rhysodini (Coleoptera: Carabidae) with revised keys to Yamatosa Bell and Bell and Omoglymmius (Pyxiglymmius) Bell and Bell. Stuttgarter Beiträge zur Naturkunde, Serie A (Biologie), 636, 1–7.

    Bell, R.T. & Bell, J.R. (2009) Rhysodine beetles (Insecta: Coleoptera: Carabidae): new species, new data III. Annals of Carnegie Museum, 78 (1), 45–77.

    https://doi.org/10.2992/007.078.0104

    Bell, R.T. & Bell, J.R. (2011) Four new species of Rhysodini (Coleoptera: Carabidae) with revised keys to Grouvellina Bell & Bell and the mishmicus group of Rhyzodiastes Fairmaire. Stuttgarter Beiträge zur Naturkunde A, Neue Serie, 4, 129–135.

    Beutel, R.G. (1990) Metathoracic features of Omoglymmius hamatus and their significance for classification of Rhysodini (Coleoptera: Adephaga). Entomologia Generalis, 15 (3), 185–201.

    https://doi.org/10.1127/entom.gen/15/1990/185

    Beutel, R.G. (1992a): Phylogenetic analysis of thoracic structures of Carabidae (Coleoptera). Zeitschrift für zoologische Systematik und Evolutionsforschung, 30 (1), 53–74.

    https://doi.org/10.1111/j.1439-0469.1992.tb00390.x

    Beutel, R.G. (1992b) Larval head structures of Omoglymmius hamatus and their implications for the relationships of Rhysodidae (Coleoptera: Adephaga). Entomologica Scandinavica, 23 (2), 169–184.

    https://doi.org/10.1163/187631292X00281

    Beutel, R.G. (1993) Phylogenetic analysis of Adephaga (Coleoptera) based on characters of the larval head. Systematic Entomology, 18 (2), 127–147.

    https://doi.org/10.1111/j.1365-3113.1993.tb00658.x

    Beutel, R.G. (1995) The Adephaga (Coleoptera): phylogeny and evolutionary history. In: Pakaluk, J. & Ślipiński, S.A. (Eds.), Biology, Phylogeny, and Classification of the Coleoptera: Papers Celebrating the 80th Birthday of Roy A. Crowson. Muzeum i Instytut Zoologii PAN, Warszawa, pp. 173–217.

    Beutel, R.G. (1998) Trachypachidae and the phylogeny of Adephaga (Coleoptera). In: Ball, G.E., Casale, A. & Vigna Taglianti, A. (Eds), Phylogeny and Classification of Caraboidea (Coleoptera: Adephaga). Proceedings of a Symposium (28 August, 1996, Florence, Italy) XX International Congress of Entomology. Museo Regionale di Scienze Naturali, Torino, pp. 81–106.

    Beutel, R.G. (2005) 7.9. Family Rhysodidae Laporte, 1840 (= Rhysodini s. Bell). In: Beutel, R.G. & Leschen, R.A.B. (Eds.), Handbook of Zoology. Volume IV Arthropoda: Insecta Part 38. Coleoptera, Beetles. Volume 1: Morphology and Systematics (Archostemata, Adephaga, Myxophaga, Polyphaga partim). Walter de Gruyter, Berlin, pp. 146–152.

    https://doi.org/10.1515/9783110904550

    Beutel, R.G. (2016) 7.11 Family Rhysodidae Laporte, 1840 (= Rhysodini s. Bell). In: Beutel, R.G. & Leschen, R.A.B. (Eds.), Handbook of Zoology. Arthropoda: Insecta. Coleoptera, Beetles. Vol. 1. Morphology and Systematics (Archostemata, Adephaga, Myxophaga, Polyphaga partim).2nd Edition. Walter de Gruyter, Berlin, pp. 190–197.

    https://doi.org/10.1515/9783110373929

    Beutel, R.G., Ribera, I., Fikáček, M., Vasilikopoulos, A., Misof, B. & Balke, M. (2020) The morphological evolution of the Adephaga (Coleoptera). Systematic Entomology, 45 (2), 378–395.

    https://doi.org/10.1111/syen.12403

    Beutel, R.G. & Roughley, R.E. (1988) On the systematic position of the family Gyrinidae (Coleoptera: Adephaga). Zeitschrift für Zoologische Systematik und Evolutionsforschung, 26 (5), 380–400.

    https://doi.org/10.1111/j.1439-0469.1988.tb00324.x

    Bousquet, Y. (2012) Catalogue of Geadephaga (Coleoptera, Adephaga) of America, north of Mexico. ZooKeys, 245, 1–1722.

    https://doi.org/10.3897/zookeys.245.3416

    Bousquet, Y. (2016) Litteratura Coleopterologica (1758–1900): a guide to selected books related to the taxonomy of Coleoptera with publication dates and notes. ZooKeys, 583, 1–776.

    https://doi.org/10.3897/zookeys.583.7084

    Burakowski, B. (1975) Descriptions of larva and pupa of Rhysodes sulcatus (F.) (Col: Rhysodidae) and notes on the bionomy of this species. Annales Zoologici, Warsaw, 32 (12), 271–287.

    Chevrolat, L.A.A. (1848) Rhyzodes. In: d’Orbigny, C.V.D. (Ed.), Dictionnaire universel d’histoire naturelle résumant et complétant tous les faits présentés par les encyclopédies, les anciens dictionnaires scientifiques, les oeuvres complètes de Buffon, et les meilleurs traités spéciaux sur les diverses branches des sciences naturelles; donnant la description des êtres et des divers phénomènes de la nature, l’étymologie et la définition des noms scientifiques, les principales applications des corps organiques et inorganiques, à l’agriculture, à la médecine, aux arts industriels, etc.; dirigé par M. Charles d’Orbigny, et enrichi d’un magnifique atlas de planches gravées sur acier. Tome onzième. MM. Renard, Martinet et Cie., Paris, p. 133.

    Chevrolat, L.A.A. (1873) Descriptions de Rhysodides nouveaux et énumération des espèces décrites. Annales de la Société Entomologique de France, 3 (5), 207–218.

    Chûjô, M. & Lee, C.-E. (1993) Records of some subcortical Coleoptera new to the fauna of Korea (Insecta). Esakia, 33, 123–124.

    Costa, C., Vanin, S.A. & Casari-Chen, S.A. (1988) 6. Rhysodidae. In: Larvas de Coleoptera do Brasil. Museu de Zoologica, Universidade de São Paulo, São Paulo, pp. 71–72, pl. 5.

    https://doi.org/10.5962/bhl.title.100233

    Deuve, T. (1988) Etude phylogénétique des Coléoptères Adephaga: redéfinition de la famille des Harpalidae, sensu novo, et position systématique des Pseudomorphinae et Brachinidae. Bulletin de la Société Entomologique de France, 92 (5–6), 161–182.

    Deuve, T. (1993) L’abdomen et les genitalia des femelles de Coléoptères Adephaga. Mémoires du Muséum National d’Histoire Naturelle, 155, 1–184.

    Erichson, W.F. (1845) Rhysodides. In: Naturgeschichte der Insecten Deutschlands. Erste Abtheilung. Coleoptera. Dritter Band. Lieferung 2. Nicolai, Berlin, pp. 297–301.

    Erwin, T.L. (1985) The taxon pulse: a general pattern of lineage radiation and extinction among carabid beetles. In: Ball, G.E. (Ed.), Taxonomy, Phylogeny and Zoogeography of Beetles and Ants. W. Junk, Dordrecht, pp. 437–472.

    Erwin, T.L. (1991) The ground-beetles of Central America (Carabidae), Part II: Notiophilini, Loricerini, and Carabini. Smithsonian Contributions to Zoology, 501, 1–30.

    https://doi.org/10.5479/si.00810282.501

    Fairmaire, L. (1895) Descriptions de quelques Coléoptères de Madagascar. Annales de la Société Entomologique de Belgique, 39, 8–40.

    https://doi.org/10.5962/bhl.part.2025

    Fauvel, C.A.A. (1889) Rectifications au Catalogus Coleopterorum Europae et Caucasi (Suite). Revue d’Entomologie, 8, 175–203.

    Forsyth, D.J. (1972) The structure of the pygidial defence glands of Carabidae (Coleoptera). The Transactions of the Zoological Society of London, 32 (4), 249–309.

    https://doi.org/10.1111/j.1096-3642.1972.tb00029.x

    Ganglbauer, L. (1891) VII. Familie. Rhysodidae. In: Die Käfer von Mitteleuropa. Die Käfer der österreichisch-ungarischen Monarchie, Deutschlands, der Schweiz, sowie des französischen und italienischen Alpengebietes. Erster Band. Familienreihe Caraboidea. Carl Gerold’s Sohn, Wien, pp. 530–534.

    https://doi.org/10.5962/bhl.title.8764

    Germar, E.F. (1822) Augusti Ahrensii Fauna Insectorum Europae. fasc. sextus [6]. Car. Aug. Kümmel, Halae (Halle), [1] + [25] pp., 25 pls.

    Grandi, G. (1972) Comparative morphology and ethology of insects with a specialized diet, Rhysodes germari Ganglb. Bolletino dell’Istituto di Entomologia della Università degli Studi di Bologna, 30, 31–47.

    Grouvelle, A. (1903) Synopsis des Rhysodides et descriptions d’espèces nouvelles. Revue d’Entomologie, 22, 85–148.

    Grouvelle, A. (1914) H. Sauter’s Formosa-Ausbeute. Rhysodidae, Nitidulidae, Ostomidae, Colydiidae, Passandridae, Cucujidae, Cryptophagidae, Diphyllidae, Lathridiidae, Mycetophagidae, Dermestidae. Archiv für Naturgeschichte, 79 (11), 33–34.

    Hincks, W.D. (1950) Coleopterorum Catalogus, Supplementa, Paris 1, Rhysodidae (Editio seconda). W. Junk, The Hague, 18 pp.

    Hovorka, O. (2018) A new Yamatosa species from China (Coleoptera: Carabidae: Rhysodini) and new distributional records of Omoglymmius sakuraii (Nakane, 1973). Studies and Reports, Taxonomical Series, 14 (2), 269–273.

    Hua, L.-Z. (2002) (V) Superfamily Rhysodoidea. Family Rhysodidae. In: List of Chinese Insects. Vol. II. Zhongshan (Sun Yat-sen) University Press, Guangzhou, p. 42.

    Huber, C., Marggi, W. & Löbl, I. (2017) Family Rhysodidae Laporte, 1840. In: Löbl, I. & Löbl, D. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 1. Archostemata—Myxophaga—Adephaga. Revised and Updated Edition. Brill, Leiden and Boston, pp. 29–31.

    https://doi.org/10.1163/9789004330290

    Jacquelin du Val, P.N.C. (1858) s.n. In: Manuel entomologique. Généra des coléoptères d’Europe comprenant leur classification en familles naturelles, la description de tous les genres, des tableaux synoptiques destinés à faciliter l’étude, le catalogue de toutes les espèces, de nombreux dessins au trait de charactères et plus de treize cents types représentant un ou plusieurs insectes de chaque genre dessinés et peints d’après nature avec le plus grand soin par M. Jules Migneaux. Tome deuxième. A. Deyrolle, Paris, p. 187.

    https://doi.org/10.5962/bhl.title.35866

    Kirby, W.F. (1830) The characters of Clinidium, a new genus of insects in the order of Coleoptera, with a description of Clinidium guildingii. The Zoological Journal, 5, 6–10.

    Kiriakoff, S.G. (1967) Lepidoptera. Familia Notodontidae. Pars secunda Genera Palaearctica. In: Wytsman, P. (Ed.), Genera Insectorum. Fascicle 217B. Kraainem. L. Desmet-Verteneuil, Bruxelles, pp. 1–238., 8 pls.

    Kryzhanovskij, O.L. (1983) 1. Fam. Rhysodidae. In: The beetles of the suborder Adephaga: families Rhysodidae, Trachypachidae, Carabidae (introduction and a review of the USSR fauna). Fauna SSSR. Coleoptera. Vol. 1. No. 2. Nauka, Leningrad, pp. 17–39.

    Kryzhanovskij, O.L., Belousov, I.A., Kabak, I.I., Kataev, B.M., Makarov, K.V. & Shilenkov, V.G. (1995) A Checklist of the Ground-Beetles of Russia and Adjacent Lands (Insecta, Coleoptera, Carabidae). Pensoft Publishers, Sofia-Moscow, 271 pp.

    Lacordaire J.T. (1854) Famille XIX. Rhysodides. In: Histoire naturelle des insectes. Genera des Coléoptères ou exposé méthodique et critique de tous les genres proposés jusqu’ici dans cet ordre d’insectes. Tome deuxième contenant les familles des paussides, staphyliniens, psélaphiens, scydménides, silphales, sphériens, trichoptérygiens, scaphidiles, histériens, phalacrides, nitidulaires, trogositaires, colydiens, rhysodides, cucujipes, cryptophagides, lathridiens, mycétaphagides, thorictides, dermestins, byrrhiens, géoryssins, parnides, hétérocérides. Roret, Paris, p. 385–389.

    https://doi.org/10.5962/bhl.title.67686

    Latreille, P.A. (1829) Le règne animal distribué d’après son organisation, pour servir de base à l’histoire naturelle des animaux et d’introduction à l’anatomie comparée. Par M. le Baron Cuvier. Avec figures, dessinées d’après nature. Nouvelle édition, revue et augmentée. Tome IV. Crustacés, arachnides et partie des insectes. Déterville, Paris, p. 487.

    Lawrence, J.F. & Britton, E.B. (1994) Australian Beetles. Melbourne University Press, Melbourne, 192 pp.

    https://doi.org/10.1071/9780643100695

    Lawrence, J.F. & Newton, A.F. (1995) Families and subfamilies of Coleoptera (with selected genera, notes, references and data on family-group names). In: Pakaluk, J. & Ślipiński, S.A. (Eds.), Biology, Phylogeny, and Classification of the Coleoptera: Papers Celebrating the 80th Birthday of Roy A. Crowson. Muzeum i Instytut Zoologii PAN, Warszawa, pp. 779–1006.

    Lawrence, J.F. & Ślipiński, A. (2013) Australian Beetles. Morphology, Classification and Keys. Volume 1. CSIRO Publishing, Collingwood, Victoria, viii + 561 pp., 98 pls.

    https://doi.org/10.1071/9780643097292

    LeConte, J.L. (1875) Notes on the Rhysodidae of the United States. Transactions of the American Entomological Society, 5, 162–168.

    https://doi.org/10.2307/25076302

    LeConte, J.L. & Horn, G.H. (1883) Fam. XXIV.—Rhyssodidae. In: Classification of the Coleoptera of North America. Prepared for the Smithsonian Institution. Smithsonian Miscellaneous Collection [No] 507. The Smithsonian Institution, Washington, D.C., p. 130.

    https://doi.org/10.5962/bhl.title.41105

    Lewis, G. (1888) On the Family Rhysodidae. The Annals and Magazine of Natural History, 2 (6), 76–85.

    https://doi.org/10.1080/00222938809460879

    Li, J.-K. (1992) Rhysodidae. In: The Coleoptera Fauna of Northeast China. Jilin Education Publishing House, Changchun, p. 2.

    Li, Z.-Y., Li, X.-X., Song, N., Tang, H.-J. & Yin, X.-M. (2020) The mitochondrial genome of Amara aulica (Coleoptera, Carabidae, Harpalinae) and insights into the phylogeny of ground beetles. Genes, 11 (2), E181.

    https://doi.org/10.3390/genes11020181

    Liebherr, J.K. & Will, K.W. (1998) Inferring phylogenetic relationships within Carabidae (Insecta, Coleoptera) from characters of the female reproductive tract. In: Ball, G.E., Casale, A. & Vigna Taglianti, A. (Eds.), Phylogeny and Classification of Caraboidea (Coleoptera: Adephaga). Proceedings of a Symposium, Florence, Italy, 28 August, 1996. XX International Congress of Entomology. Museo Regionale di Scienze Naturali, Torino, pp. 101–170.

    López-López, A. & Vogler, A.P. (2017) The mitogenome phylogeny of Adephaga (Coleoptera). Molecular Phylogenetics and Evolution, 114, 166–174.

    https://doi.org/10.1016/j.ympev.2017.06.009

    Lorenz, W. (2005) Subfam. Rhysodinae Laporte de Castelnau 1840. In: Systematic list of extant ground beetles of the world (Insecta Coleoptera “Geadephaga”: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae). Second Edition. Wolfgang Lorenz, Tutzing, pp. 156–159.

    Maddison, D.R., Baker, M.D. & Ober, K.A. (1999) Phylogeny of carabid beetles as inferred from 18S ribosomal DNA (Coleoptera: Carabidae). Systematic Entomology, 24 (2), 103–138.

    https://doi.org/10.1046/j.1365-3113.1999.00088.x

    Maddison, D.R., Moore, W., Baker, M.D., Ellis, T.M., Ober, K.A., Cannone, J.J. & Gutell, R.R. (2009) Monophyly of terrestrial adephagan beetles as indicated by three nuclear genes (Coleoptera: Carabidae and Trachypachidae). Zoologica Scripta, 38 (1), 43–62.

    https://doi.org/10.1111/j.1463-6409.2008.00359.x

    Makarov, K.V. (2008) Larval chaetotaxy in the genus Rhysodes Dalman, 1823 and the position of Rhysodidae within Adephaga (Coleoptera). In: Penev, L., Erwin, T.L. & Assmann, T. (Eds.), Back to the Roots and Back to the Future: towards a New Synthesis amongst Taxonomic, Ecological and Biogeographical Approaches in Carabidology. Proceedings of the XIII European carabidologists meeting, Blagoevgrad, 2024 August 2007. Pensoft, Sofia-Moscow, pp. 101–123.

    McKenna, D.D., Wild, A.L., Kanda, K., Bellamy, C.L., Beutel, R.G., Caterino, M.S., Farnum, C.W., Hawks, D.C., Ivie, M.A., Jameson, M.L., Leschen, R.A.B., Marvaldi, A.E., McHugh, J.V., Newton, A.F., Robertson, J.A., Thayer, M.K., Whiting, M.F., Lawrence, J.F., Ślipiński, A., Maddison, D.R. & Farrell, B.D. (2015) The beetle tree of life reveals that Coleoptera survived end-Permian mass extinction to diversify during the Cretaceous terrestrial revolution. Systematic Entomology, 40, 835–880.

    https://doi.org/10.1111/syen.12132

    Miwa, Y. (1931) Rhysodidae. In: A Systematic Catalogue of Formosan Coleoptera. Reprinted from the Report No. 55. Department of Agriculture, Government Research Institute, Taihoku, Formosa. Contribution No. 32. Entomological Laboratory, Taihoku Imperial University, Taipei, p. 21.

    Miwa, Y. (1934) On a new species of Rhysodidae from Formosa. Transactions of the Natural History Society of Formosa, 24 (133), 256–257.

    Nagel, P. (1979) The classification of Carabidae. In: den Boer, P.J., Thiele, H.-U. & Weber, F. (Eds.), On the Evolution of Behaviour in Carabid Beetles. Report of a symposium, held at the fieldstation Rees-Grietherbusch of the Zoological Institute of the University of Cologne 1013 September 1978. Veenman & Zonen, Wageningen, pp. 7–14.

    Nakane, T. (1960) On the Rhysodidae of Japan (Coleoptera). Akitu, 9 (3–4), 69–71.

    Nakane, T. (1973) The beetles of Japan, new series (1). Cupedidae and Rhysodidae. The Nature and Insects, 8 (9), 2–5.

    Nakane, T. (1978) New or little-known Coleoptera from Japan and its adjacent regions, XXIX. Reports of the Faculty of Science, Kagoshima University (Earth Sciences and Biology), 11, 129–134.

    Newman, E. (1838) A descriptive list of the species of Rhysodes. The Magazine of Natural History, 2 (2), 663–667.

    Reichardt, H. (1977) A synopsis of the genera of Neotropical Carabidae (Insecta: Coleoptera). Quaestiones Entomologicae, 13 (4), 346–493.

    Reitter, E. (1882) Bestimmungs-Tabelle der europäischen Coleopteren. VI. Enthaltend die Familien: Colydiidae, Rhysodidae, Trogositidae. Verhandlungen des Naturforschenden Vereins in Brünn, 20, 113–149.

    Reitter, E. (1908) VII. Familie. Rhyssodidae. In: Fauna Germanica. Die Käfer des Deutschen Reiches. I. Band. Schriften des Deutschen Lehrervereins für Naturkunde 22. K.G. Lutz, Stuttgart, pp. 237–238, pl. 40.

    Reitter, E. (1922) Fam. Rhysodidae. In: Bestimmungs-Tabellen der europäischen Coleopteren. VI. Heft. Enthaltend die Familien: Colydiidae, Rhysodidae, Ostomidae. 2. Gänzlich umgearbeitete und auf die palaearktische Fauna ausgedehnte Auflage. E. Reitter, Troppau, pp. 54–57.

    Regenfuss, H. (1975) Die Antennen-Putzeinrichtung der Adephaga (Coleoptera), parallele evolutive Vervollkommnung einer komplexen Struktur. Zeitschrift fur Zoologische Systematik und Evolutionsforschung, 13 (4), 278–299.

    https://doi.org/10.1111/j.1439-0469.1975.tb00510.x

    Seidlitz, G.C.M. von (1888) 24. Fam. Rhysodidae. In: Fauna Transsylvanica. Die Käfer (Coleoptera) Siebenbürgens. Mit 1 Tafel. Lieferungen 1–2. Hartung, Königsberg, p. 54.

    Sundukov, Y.N. (2013) Family Rhysodidae Laporte, 1840. In: An Annotated Catalogue of the Ground Beetles (Coleoptera: Caraboidea) of Sikhote-Alin. Dalnauka, Vladivostok, p. 74.

    Sundukov, Y.N. & Makarov, K.V. (2014) Review of the family Rhysodidae (Coleoptera Adephaga) of Kuril Islands, Russia. Far Eastern Entomologist, 273, 18–20.

    Thomson, C.G. (1863) Familia Rhyssodidae. In: Skandinaviens Coleoptera, synoptiskt bearbetade. Tom. V. Lundbergska Boktryckeriet, Lund, pp. 110–112.

    Wang, C.-B. (2016) Rhyzodiastes (Temoana) xii sp. nov. (Coleoptera: Rhysodidae: Clinidiini), a new species from Hainan Island, China. Zootaxa, 4126 (2), 287–294.

    https://doi.org/10.11646/zootaxa.4126.2.9

    Wang, C.-B., Růžička, J. & Liu, B. (2017) Omoglymmius (s. str.) wukong sp. n., a new species from Xizang, China (Coleoptera, Rhysodidae, Omoglymmiini). ZooKeys, 706, 95–107.

    https://doi.org/10.3897/zookeys.706.14655

    Westwood, J.O. (1835) On the affinities of the genus Clinidium of Kirby. The Zoological Journal, 5, 213–237.

    Winkler, A. (1924) Rhysodidae. In: Catalogus Coleopterorum Regionis Palaearcticae. Albert Winkler, Wien, pp. 239–240.

    Yahiro, K. (1996) Comparative morphology of the alimentary canal and reproductive organs of the terrestrial Caraboidea (Coleoptera: Adephaga) Part 1. Japanese Journal of Entomology, 64 (3), 536–550.

    Zhang, W.-W. (2014) Rhysodidae. In: Insect Genealogy. Chongqing University Press, Chongqing, p. 172.