Abstract
The known cicada fauna of Ecuador is identified. Durangona exechopyga n. sp., Calyria chaetoacontia n. sp., Calyria xiphion n. sp., Carineta coronida n. sp., and Carineta tiarata n. sp. are described as new. The subtribe Plautillina Distant, 1905h rev. stat. is formed for the genera Plautilla Stål, 1865 and Onoralna Boulard, 1996 within the Zammarini Distant, 1905b. Zammara erna Schmidt, 1919 n. syn., Orellana brunneipennis Goding, 1925 n. syn. and Orellana pulla Goding, 1925 n. syn. are reported to be junior synonyms of Zammara intricata Walker, 1850 and Coata facialis var. Jacobi 1907a n. syn. is shown to be unavailable and a junior synonym of Coata facialis Distant 1906b. The first records of Fidicinoides besti Boulard & Martinelli, 1996, Fidicinoides brunnea Boulard & Martinelli, 1996, Fidicinoides descampsi Boulard & Martinelli, 1996, Fidicinoides ptychodiropeda Sanborn 2020b, Ariasa bilaqueata (Uhler, 1903), Guyalna bicolor (Olivier, 1790), Guyalna dyticamazona Sanborn, 2020b, Calyria fenestrata (Fabricius, 1803), Taphura boulardi Sanborn, 2011a, Carineta lichiana Boulard, 1986a, Carineta peruviana Distant, 1905c, Carineta producta Walker, 1858b, Carineta rufescens (Fabricius, 1803), Herrera concolor Sanborn, 2019b and Toulgoetalna tavakiliani Boulard, 1982 are provided. The new records represent the first records for Ariasa Distant 1905d, Calyria Stål, 1862, the tribe Parnisini Distant, 1905e, and Toulgoetalna Boulard, 1982, in Ecuador. The record for Calyria fenestrata is the first known specific locality for the species. An additional first record of Fidicinoides besti is provided for Colombia to expand the known distribution of this species. Previous records of Zammara calochroma Walker, 1858a, Dorisiana semilata (Walker, 1850), Carineta bilineosa Walker, 1858b, Carineta fasciculata (Germar, 1821), and Selymbria stigmatica (Germar, 1834) are considered to be misidentifications of Dorisiana metcalfi Sanborn & Heath, 2014 (= Cicada viridis Olivier, 1790), C. maculosa Torres, 1848, C. pilifera Walker, 1858a, and S. ecuadorensis Sanborn, 2019a, respectively, so that Z. calochroma, D. semilata, C. bilineosa, C. fasciculata, and S. stigmatica are removed from the cicada fauna of Ecuador. In addition, Chilecicada occidentis (Walker, 1850) is removed from the Ecuadorian cicada fauna as the species is restricted to Chile. The currently known Ecuadorian cicada fauna is comprised of 98 species from 24 genera, nine tribes and three subfamilies.
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Sanborn, A.F. (2006a) New records for the cicada fauna from four Central American countries (Hemiptera: Cicadoidea: Cicadidae). Florida Entomologist, 89, 75–79.
https://doi.org/10.1653/0015-4040(2006)89[75:NRFTCF]2.0.CO;2
Sanborn, A.F. (2006b) New records of cicadas from Mexico (Hemiptera: Cicadoidea: Cicadidae). Southwestern Naturalist, 51, 255–257.
https://doi.org/10.1894/0038-4909(2006)51[255:NROCFM]2.0.CO;2
Sanborn, A.F. (2007a) New species, new records and checklist of cicadas from Mexico (Hemiptera: Cicadoidea: Cicadidae). Zootaxa, 1651 (1), 1–42.
https://doi.org/10.11646/zootaxa.1651.1.1
Sanborn, A.F. (2007b) Additions to the cicada fauna of Venezuela with the description of a new species and checklist of the Venezuelan cicada fauna (Hemiptera: Cicadoidea: Cicadidae). Zootaxa, 1503 (1), 21–32.
https://doi.org/10.11646/zootaxa.1503.1.3
Sanborn, A.F. (2008) New records of Brazilian cicadas including the description of a new species (Hemiptera: Cicadoidea, Cicadidae). Neotropical Entomology, 37, 685–690.
https://doi.org/10.1590/s1519-566x2008000600010
Sanborn, A.F. (2009a) Checklist, new species and key to the cicadas of Cuba (Hemiptera: Cicadoidea: Cicadidae). Mitteilungen aus dem Museum für Naturkunde in Berlin – Deutsche Entomologische Zeitschrift, 59, 85–92.
https://doi.org/10.1002/mmnd.200900007
Sanborn, A.F. (2009b) A new species of Cicadetta (Hemiptera: Cicadomorpha: Cicadidae) from Hispaniola. Caribbean Journal of Science, 45, 1–7.
https://doi.org/10.18475/cjos.v45i1.a2
Sanborn, A.F. (2010a) Two new species and new records of cicadas from Central America (Hemiptera: Cicadoidea: Cicadidae). Studies on Neotropical Fauna and Environment, 45, 67–76.
https://doi.org/10.1080/01650521003778814
Sanborn, A.F. (2010b) The cicadas of Colombia including new records and the description of a new species (Hemiptera: Cicadidae). Journal of Natural History, 44, 1577–1607.
https://doi.org/10.1080/00222931003760046
Sanborn, A.F. (2011a) Checklist of the cicadas of French Guiana including new records and the description of nine new species (Insecta, Hemiptera, Cicadoidea, Cicadidae). Zoosystema, 33, 377–418.
https://doi.org/10.5252/z2011n3a6
Sanborn, A.F. (2011b) Checklist of the cicadas (Insecta: Hemiptera: Cicadidae) of Paraguay including new records for six species. Check List, 7, 465–467.
https://doi.org/10.15560/7.4.465
Sanborn, A.F. (2013) Catalogue of the Cicadoidea (Hemiptera: Cicadoidea). With contributions to the bibliography by Martin H. Villet. Elsevier/Academic Press, San Diego, 1001 pp.
https://doi.org/10.1016/b978-0-12-416647-9.00001-2
Sanborn, A.F. (2014a) Checklist of the cicadas (Insecta: Hemiptera: Cicadidae) of Costa Rica including new records for fourteen species. Check List, 10, 246–252.
https://doi.org/10.15560/10.2.246
Sanborn, A.F. (2014b) A new genus and new tribe of cicada from South America (Hemiptera: Cicadoidea: Cicadidae) with a note on the taxonomic position of Ahomana Distant, 1905. Proceedings of the Entomological Society of Washington, 116, 339–348.
https://doi.org/10.4289/0013-8797.116.3.339
Sanborn, A.F. (2016a) The cicada genus Guyalna Boulard & Martinelli, 1996 (Hemiptera: Cicadidae: Cicadinae: Fidicinini): generic description, twelve new combinations, and a key to species. Zootaxa, 4105, 430–454.
https://doi.org/10.11646/zootaxa.4105.5.2
Sanborn, A.F. (2016b) Five new species, a new genus and a new record of cicadas from French Guiana with four new combinations and three new synonymies (Insecta, Hemiptera, Cicadoidea, Cicadidae). Zoosystema, 38, 177–199.
https://doi.org/10.5252/z2016n2a2
Sanborn, A.F. (2016c) Generic redescription, two new species, and a key to the species of the cicada genus Ariasa Distant, 1905 with the description of a new genus (Hemiptera: Cicadidae: Cicadinae: Fidicinini). Zootaxa, 4137 (4), 501–519.
https://doi.org/10.11646/zootaxa.4137.4.3
Sanborn, A.F. (2017a) Generic redescription, seven new species and a key to the Taphura Stål, 1862 (Hemiptera: Cicadidae: Cicadettinae: Taphurini). Zootaxa, 4324 (3), 451–481.
https://doi.org/10.11646/zootaxa.4324.3.3
Sanborn, A.F. (2017b) A new species of Carineta Amyot & Audinet-Serville, 1843 (Hemiptera: Cicadoidea: Cicadidae) from Martinique. Proceedings of the Entomological Society of Washington, 119, 629–636.
https://doi.org/10.4289/0013-8797.119.4.629
Sanborn, A.F. (2018a) The cicada genus Procollina Metcalf, 1952 (Hemiptera: Cicadidae): redescription including fourteen new species, with a key to the species of the subtribe Dazina Kato, 1932 rev. stat., the description of the Aragualnini n. tribe, and one new combination. Zootaxa, 4389 (1), 1–65.
https://doi.org/10.11646/zootaxa.4389.1.1
Sanborn, A.F. (2018b) Two new genera and two new species of cicadas from Central America (Hemiptera: Cicadidae: Cicadinae). Zootaxa, 4418 (2), 148–160.
https://doi.org/10.11646/zootaxa.4418.2.4
Sanborn, A.F. (2018c) The cicadas (Hemiptera: Cicadidae) of Panama including the description of six new species, three new combinations, one new synonymy, and nine new records. Zootaxa, 4493 (1), 1–69.
https://doi.org/10.11646/zootaxa.4493.1.1
Sanborn, A.F. (2019a) The cicada genus Selymbria Stå1, 1861 (Hemiptera: Cicadidae: Tibicininae: Selymbrini): redescription including ten new species and a key to the genus. Zootaxa, 4614 (3), 401–448.
https://doi.org/10.11646/zootaxa.4614.3.1
Sanborn, A.F. (2019b) The cicadas (Hemiptera: Cicadidae) of Bolivia including the descriptions of fifteen new species, the resurrection of one genus and two species, seven new combinations, six new synonymies, and twenty-eight new records. Zootaxa, 4655 (1), 1–104.
https://doi.org/10.11646/zootaxa.4655.1.1
Sanborn, A.F. (2019c) Two new species of Neotropical cicadas (Hemiptera: Cicadidae: Cicadettinae: Carinetini) from Bolivia. Journal of Insect Biodiversity, 13, 27–35.
https://doi.org/10.12976/jib/2019.13.2.2
Sanborn, A.F. (2020a) Eight new species and three new records of Neotropical cicadas (Hemiptera: Cicadidae) from Venezuela. Journal of Insect Biodiversity, 16, 6–37.
https://doi.org/10.12976/jib/2020.16.1.2
Sanborn, A.F. (2020b) The cicadas (Hemiptera: Cicadidae) of Peru including the descriptions of twenty-four new species, three new synonymies, and thirty-seven new records. Zootaxa, 4785 (1), 1–129.
https://doi.org/10.11646/zootaxa.4785.1.1
Sanborn, A.F. (2020c) Two new species of Neotropical cicadas (Hemiptera: Cicadidae: Cicadettinae) from southeastern Brazil. Journal of Insect Biodiversity, 19, 14–23.
https://doi.org/10.12976/jib/2020.19.1.3
Sanborn, A.F. (2020d) A new species of Herrera Distant, 1905 (Hemiptera: Cicadoidea: Cicadidae: Cicadettinae: Carinetini) from Panama with new records for two additional species. Proceedings of the Entomological Society of Washington, 122. [in press]
Sanborn, A.F. (2020e) The cicadas (Hemiptera: Cicadidae) of Trinidad and Tobago including the descriptions of three new species and seven new records, with new records for several additional Caribbean Islands. Zootaxa, 4838 (4), 535–565.
https://doi.org/10.11646/zootaxa.4838.4.7
Sanborn, A.F. & Heath, M.S. (2012) Catalogue of the cicadas (Hemiptera: Cicadoidea: Cicadidae) of continental North America North of Mexico. Thomas Say Publications in Entomology: Monographs. Entomological Society of America, Lanham, Maryland, 227 pp.
https://doi.org/10.3390/d5020166
Sanborn, A.F. & Heath, M.S. (2014) The cicadas of Argentina with new records, a new genus and fifteen new species (Hemiptera: Cicadoidea: Cicadidae). Zootaxa, 3883 (1), 1–94.
https://doi.org/10.11646/zootaxa.3883.1.1
Sanborn, A.F. & Heath, M.S. (2017) Catalogue of the cicadas (Hemiptera: Cicadoidea: Cicadidae) of continental North America North of Mexico, Second Edition. Thomas Say Publications in Entomology. Monographs. Entomological Society of America, Lanham, Maryland, 302 pp.
https://doi.org/10.3390/d5020166
Sanborn, A.F., Heath, M.S., Heath, J.E. & Noriega, F.G. (1995) Diurnal activity, temperature responses and endothermy in three South American cicadas (Homoptera: Cicadidae: Dorisiana bonaerensis, Quesada gigas, and Fidicina mannifera). Journal of Thermal Biology, 20, 451–460.
https://doi.org/10.1016/0306-4565(95)00004-G
Sanborn, A.F., Heath, J.E., Philllips, P.K., Heath, M.S. & Noriega, F.G. (2011a) Thermal adaptation and diversity in tropical ecosystems: evidence from cicadas (Hemiptera, Cicadidae). PLoS ONE, 6 (e29368), 1–12.
https://doi.org/10.1371/journal.pone.0029368
Sanborn, A.F., Heath, M.S., Phillips, P.K. & Heath, J.E. (2011b) The genus Cacama Distant, 1904 (Hemiptera: Cicadidae) with the description of three new species. Zootaxa, 2897 (1), 35–50.
https://doi.org/10.11646/zootaxa.2891.1.4
Sanborn, A.F., Heath, M.S., Phillips, P.K. & Heath, J.E. (2012) The genus Pacarina Distant, 1905 (Hemiptera: Cicadidae) with the description of a new species. Journal of Natural History, 46, 923–941.
https://doi.org/10.1080/00222933.2011.651647
Sanborn, A.F., Heath, M.S., Sueur, J. & Phillips, P.K. (2005) The genus Neocicada Kato, 1932 (Hemiptera: Cicadomorpha: Cicadidae), with descriptions of three new species. Systematic Entomology, 30, 191–207.
https://doi.org/10.1111/j.1365-3113.2005.00266.x
Sanborn, A.F. & Maes, J.-M. (2012) Checklist of the cicadas (Insecta: Hemiptera: Cicadidae) of Nicaragua including new records for seventeen species. Check List, 8, 437–442.
https://doi.org/10.15560/8.3.437
Sanborn, A.F., Moore, T.E. & Young, A.M. (2008) Two new cicada species from Costa Rica (Hemiptera: Cicadomorpha: Cicadidae) with a key to Fidicinoides in Costa Rica. Zootaxa, 1846 (1)), 1–20.
https://doi.org/10.11646/zootaxa.1846.1.1
Sanborn, A.F. & Phillips, P.K. (2013) Biogeography of the cicadas (Hemiptera: Cicadidae) of North America, North of Mexico. Diversity, 5, 166–239.
https://doi.org/10.3390/d5020166
Sanborn, A.F., Phillips, P.K. & Sites, R.W. (2007) Biodiversity, biogeography, and bibliography of the cicadas of Thailand (Hemiptera: Cicadoidea: Cicadidae). Zootaxa, 1413 (1), 1–46.
https://doi.org/10.11646/zootaxa.1413.1.1
Santos, R.S. & Martinellli, N.M. (2009) Descrição de novas espécies de Fidicinoides Boulard & Martinelli (Hemiptera: Cicadidae) do Brasil. Neotropical Entomology, 38, 638–642.
https://doi.org/10.1590/s1519-566x2009000500012
Santos, R.S., Martinelli, N.M., Maccagnan, D.H.B., Sanborn, A.F. & Ribeiro, R. (2010) Description of a new cicada species associated with the coffee plant and an identification key to the species of Fidicinoides (Hemiptera: Cicadidae) from Brazil. Zootaxa, 2602, 48–56.
https://doi.org/10.11646/zootaxa.2602.1.2
Santos-Cividanes, T.M., Seguino, E., Cividanes, F.J., Martinelli, N.M., Martins, A.M. & Perdoná, M.J. (2013) First record of Dorisiana viridis (Hemiptera: Cicadidae) on Macadamia Nut (Proteales: Proteacea) in Brazil. Florida Entomologist, 96, 1221–1223.
https://doi.org/10.1653/024.096.0373
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