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Type: Article
Published: 2020-08-13
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Some leptothecate hydroids (Cnidaria, Hydrozoa) from Hawaii, mostly from inshore and nearshore waters

Department of Natural History, Royal Ontario Museum, 100 Queen’s Park, Toronto, Ontario, Canada M5S 2C6. Research Associate, Royal British Columbia Museum, 675 Belleville Street, Victoria, British Columbia, Canada V8W 9W2.
Coelenterata Anthoathecata Hydroidolina Leptothecata marine invertebrates Medusozoa taxonomy zoological nomenclature

Abstract

This report is based on a small collection of hydroids from the Hawaiian Islands, in the central Pacific Ocean. Most of the examined material was obtained by staff of the Bernice Pauahi Bishop Museum, Honolulu, during surveys for nonindigenous marine species in shallow, sheltered, inshore or nearshore waters, and especially in harbours, bays, and lagoons. In all, 34 species of leptothecate hydroids, assigned to 14 families and 20 genera, were identified and are discussed. One of them, based on a single infertile colony with a damaged hydrotheca, was identified provisionally only to the rank of suborder. Given the limited geographic and bathymetric focus of the surveys, only four of the species, Clytia thornelyi, Halecium sibogae, Macrorhynchia balei, and M. hawaiensis, were collected at depths greater than 25 m. Seven species, Cirrholovenia tetranema Kramp, 1959, Orthopyxis crenata (Hartlaub, 1901), Clytia elongata Marktanner-Turneretscher, 1890, C. paulensis (Vanhöffen, 1910), Tridentata maldivensis (Borradaile, 1905), Monotheca flexuosa (Bale, 1894), and a hydroid identified only as Eirenida (undetermined), are recorded from Hawaii for the first time. Three others, Lytocarpia nigra (Nutting, 1905) Macrorhynchia balei (Nutting, 1905), and M. hawaiensis (Nutting, 1905), have their type localities in Hawaii, with the last of these being known to date only from the Hawaiian archipelago. Most of the species are well-known from shallow water areas across the tropical and subtropical Indo-Pacific region, and over half of them have been reported as well from warm waters in the Atlantic Ocean. Their existence in the remote islands of Hawaii is attributed to long-range dispersal by both natural and human-mediated means, including shipping.

 

References

  1. Agassiz, L. (1862) Contributions to the natural history of the United States of America. Vol. IV. Little, Brown, Boston, 380 pp.

    Alder, J. (1859) Descriptions of three new species of sertularian zoophytes. Annals and Magazine of Natural History, Series 3, 3, 353–356.

    Allman, G.J. (1872) A monograph of the gymnoblastic or tubularian hydroids. Conclusion of Part I, and Part II, containing descriptions of the genera and species of the Gymnoblastea. Vol. II. Ray Society Publication 47. Ray Society, Robert Hardwicke, Piccadilly, London, pp. 155–450.

    https://doi.org/10.5962/bhl.title.9122

    Allman, G.J. (1874) On the diagnosis of new genera and species of hydroids. Nature, 11, 179.

    Allman, G.J. (1877) Report on the Hydroida collected during the exploration of the Gulf Stream by L.F. de Pourtalès, assistant United States Coast Survey. Memoirs of the Museum of Comparative Zoölogy at Harvard College, 5 (2), 1–66.

    https://doi.org/10.5962/bhl.title.15852

    Allman, G.J. (1883) Report on the Hydroida dredged by H.M.S. Challenger during the years l873–76. Part I.—Plumularidae. Report on the Scientific Results of the Voyage of H.M.S. Challenger during the years 1873–76, Zoology, 7 (20), 1–55.

    https://doi.org/10.5962/bhl.title.11299

    Allman, G.J. (1885) Description of Australian, Cape, and other Hydroida, mostly new, from the collection of Miss H. Gatty. Journal of the Linnean Society, Zoology, 19, 132–161.

    https://doi.org/10.1111/j.1096-3642.1885.tb01994.x

    Allman, G.J. (1888) Report on the Hydroida dredged by H.M.S. Challenger during the years 1873–76. Part II.—The Tubularinae, Corymorphinae, Campanularinae, Sertularinae, and Thalamophora. Report on the Scientific Results of the Voyage of H.M.S. Challenger during the years 1873–76, Zoology, 23 (70), 1–90.

    Ansín Agís, J., Ramil, F. & Vervoort, W. (2001) Atlantic Leptolida (Hydrozoa, Cnidaria) of the families Aglaopheniidae, Halopterididae, Kirchenpaueriidae and Plumulariidae collected during the CANCAP and Mauritania-II expeditions of the National Museum of Natural History, Leiden, the Netherlands. Zoologische Verhandelingen, 333, 1–268.

    Ansín Agís, J., Vervoort W. & Ramil F. (2009) Hydroids of the family Halopterididae (Cnidaria, Hydrozoa) collected in the western Pacific by various French expeditions. Zoosystema, 31 (1), 33–61.

    https://doi.org/10.5252/z2009n1a3

    Bailey-Brock, J.H. (1989) Fouling community development on an artificial reef in Hawaiian waters. Bulletin of Marine Science, 44, 580–591.

    Bale, W.M. (1882) On the Hydroida of south-eastern Australia, with descriptions of supposed new species, and notes on the genus Aglaophenia. Journal of the Microscopical Society of Victoria, 2, 15–48.

    Bale, W.M. (1888) On some new and rare Hydroida in the Australian Museum collection. Proceedings of the Linnean Society of New South Wales, Series 2, 3, 745–799.

    https://doi.org/10.5962/bhl.title.26267

    Bale, W.M. (1894) Further notes on Australian hydroids, with descriptions of some new species. Proceedings of the Royal Society of Victoria, New Series, 6, 93–117.

    Bale, W.M. (1914) Report on the Hydroida collected in the Great Australian Bight and other localities. Part II. Zoological Results of the Fishing Experiments Carried on by F.I.S. “Endeavour”, 1909–10, 2, 164–188.

    Bedot, M. (1921) Notes systématiques sur les plumularides. 2me partie. Revue Suisse de Zoologie, 29, 1–40.

    https://doi.org/10.5962/bhl.part.84689

    Billard, A. (1912) Hydroïdes de Roscoff. Archives de Zoologie Expérimentale et Générale, 51, 459–478.

    Billard, A. (1913) Les hydroïdes de l’expédition du Siboga. I. Plumulariidae. Siboga-Expeditie, Monographie, 7a, 1–115.

    Billard, A. (1925a) Note sur quelques espèces la plupart nouvelles de synthécides et de sertularides du Siboga. Bulletin de la Société Zoologique de France, 49, 646–652. [Publication date 10 April l925, following Vervoort (1995)]

    Billard, A. (1925b) Les hydroïdes de l’expédition du Siboga. II. Synthecidae et Sertularidae. Siboga-Expeditie, Monographie, 7b, 115–232. [publication date December l925, following Vervoort (1995)]

    Billard, A. (1927a) Question de synonymie (Sertularia turbinata, S. loculosa, S. ligulata). Bulletin de la Société Zoologique de France, 51, 512–513.

    Billard, A. (1927b) Hydrozoa I. Hydrozoa benthonica (excl. Hydrocorallidae). In: Monod, Th. (Ed.), Faune des Colonies Françaises, Contribution à l’Étude de la Faune du Cameroun, 1, 467–474.

    Billard, A. (1929a) Note sur deux espèces d’halécide du genre Diplocyathus Allm. Bulletin de la Société Zoologique de France, 54, 69–71.

    Billard, A. (1929b) Note sur un genre nouveau et quelques espèces nouvelles d’Halecidae. Bulletin de la Société Zoologique de France, 54, 305–307.

    Billard, Α. (1934) Note sur quelques hydroïdes du Maroc. Bulletin de la Société Zoologique de France, 59, 227–231 + 468.

    https://doi.org/10.1080/00378941.1860.10826243

    Blackburn, M. (1938) 3. Hydrozoa. In: The Sir Joseph Banks Islands. Reports of the Expedition of the McCoy Society for field Investigation and Research. Proceedings of the Royal Society of Victoria, New Series, 50, 312–328.

    Borradaile, L.A. (1905) Hydroids. The Fauna and Geography of the Maldive and Laccadive Archipelagoes, Cambridge University Press, 2, 836–845.

    Bosc, L.A.G. (1802) Histoire naturelle des vers, contenant leur description et leurs mœurs; avec figures dessinées d’après nature. Tome 3. Guilleminet, Paris, 270 pp.

    https://doi.org/10.5962/bhl.title.64025

    Bouillon, J. (1984) Revision de la famille des Phialuciidae (Kramp, 1955) (Leptomedusae, Hydrozoa, Cnidaria), avec un essai de classification des Thecatae-Leptomedusae. Indo-Malayan Zoology, 1 (1984), 1–24.

    Bouillon, J. (1986) Nemalecium gen. nov., genre nouveau de Haleciidae (Thecatae-Leptomedusae, Hydrozoa, Cnidaria). Indo-Malayan Zoology, 3, 71–80.

    Bouillon, J., Gravili, C., Pagès, F., Gili, J.-M. & Boero, F. (2006) An introduction to Hydrozoa. Mémoires du Muséum National d’Histoire Naturelle, 194, 1–591.

    Brinckmann, A. (1965) The life cycle of the medusa Cirrholovenia tetranema Kramp 1959 (Leptomedusae, Lovenellidae) with a hydroid of the genus Cuspidella Hincks. Canadian Journal of Zoology, 43, 13–15.

    https://doi.org/10.1139/z65-002

    Broch, H. (1914) Hydrozoa benthonica. Beiträge zur Kenntnis der Meeresfauna Westafrikas, 1, 19–50.

    Bryant, P.J. & T.E. Arehart (2019) Diversity and life-cycle analysis of Pacific Ocean zooplankton by videomicroscopy and DNA barcoding: Hydrozoa. PLoS One, 14 (10), e0218848.

    https://doi.org/10.1371/journal.pone.0218848

    Busk, G. (1852) Appendix IV. An account of the Polyzoa, and sertularian zoophytes, collected in the voyage of the Rattlesnake, on the coasts of Australia and the Louisiade Archipelago, &c. In: Macgillivray, J. (Ed.), Narrative of the voyage of H.M.S. Rattlesnake, commanded by the late Captain Owen Stanley, R.N., F.R.S. &c., during the years 1846–1850. Vol. 1. T. & W. Boone, London, pp. 343–402.

    Cairns, S.D. (1978) Distichopora (Haplomerismos) anceps, a new stylasterine coral (Coelenterata: Stylasterina) from deep water off the Hawaiian Islands. Micronesica, 14, 83–87.

    Cairns, S.D. (2005) Revision of the Hawaiian Stylasteridae (Cnidaria: Hydrozoa: Athecata). Pacific Science, 59, 439–451.

    https://doi.org/10.1353/psc.2005.0034

    Cairns, S.D. (2017) New species of stylasterid (Cnidaria: Hydrozoa: Anthoathecata: Stylasteridae) from the Northwestern Hawaiian Islands. Pacific Science, 71, 77–81.

    https://doi.org/10.2984/71.1.7

    Calder, D.R. (1971) Hydroids and hydromedusae of southern Chesapeake Bay. Virginia Institute of Marine Science, Special Papers in Marine Science, 1, 125 pp.

    Calder, D.R. (1991) Shallow-water hydroids of Bermuda. The Thecatae, exclusive of Plumularioidea. Royal Ontario Museum, Life Sciences Contributions, 154, 1–140.

    Calder, D.R. (1993) Local distribution and biogeography of the hydroids (Cnidaria) of Bermuda. Caribbean Journal of Science, 29, 61–74.

    Calder, D.R. (1997) Shallow-water hydroids of Bermuda: superfamily Plumularioidea. Royal Ontario Museum, Life Sciences Contributions, 161, 1–85.

    Calder, D.R. (2000) Assemblages of hydroids (Cnidaria) from three seamounts near Bermuda in the western North Atlantic. Deep-Sea Research, Part I, 47, 1125–1139.

    https://doi.org/10.1016/S0967-0637(99)00093-X

    Calder, D.R. (2010) Some anthoathecate hydroids and limnopolyps (Cnidaria, Hydrozoa) from the Hawaiian archipelago. Zootaxa, 2590 (1), 1–91.

    https://doi.org/10.11646/zootaxa.2590.1.1

    Calder, D.R. (2012) On a collection of hydroids (Cnidaria, Hydrozoa, Hydroidolina) from the west coast of Sweden, with a checklist of species from the region. Zootaxa, 3171 (1), 1–77.

    https://doi.org/10.11646/zootaxa.3171.1.1

    Calder, D.R. (2013) Some shallow-water hydroids (Cnidaria: Hydrozoa) from the central east coast of Florida, USA. Zootaxa, 3648 (1), 1–72.

    https://doi.org/10.11646/zootaxa.3648.1.1

    Calder, D.R. (2015) George James Allman (1812–1898): pioneer in research on Cnidaria and freshwater Bryozoa. Zootaxa, 4020 (2), 201–243.

    https://doi.org/10.11646/zootaxa.4020.2.1

    Calder, D.R. (2017) Additions to the hydroids (Cnidaria, Hydrozoa) of the Bay of Fundy, northeastern North America, with a checklist of species reported from the region. Zootaxa, 4256 (1), 1–86.

    https://doi.org/10.11646/zootaxa.4256.1.1

    Calder, D.R. (2019) On a collection of hydroids (Cnidaria, Hydrozoa) from the southwest coast of Florida, USA. Zootaxa, 4689 (1), 1–141.

    https://doi.org/10.11646/zootaxa.4689.1.1

    Calder, D.R., Carlton, J.T., Larson, K., Mallinson, J.J., Choong, H.H.C., Keith, I. & Ruiz, G.M. (2019) Hydroids (Cnidaria, Hydrozoa) from marine fouling assemblages in the Galápagos Islands, Ecuador. Aquatic Invasions, 14, 21–58.

    https://doi.org/10.3391/ai.2019.14.1.02

    Calder, D.R. & Choong, H.H.C. (2018) Names of hydroids (Cnidaria, Hydrozoa) established by Charles McLean Fraser (1872–1946), excluding those from Allan Hancock Expeditions. Zootaxa, 4487 (1), 1–83.

    https://doi.org/10.11646/zootaxa.4487.1.1

    Calder, D.R., Mallinson, J.J., Collins, K. & Hickman, C.P. (2003) Additions to the hydroids (Cnidaria) of the Galápagos, with a list of species reported from the islands. Journal of Natural History, 37, 1173–1218.

    https://doi.org/10.1080/00222930110116039

    Calder, D.R., Vervoort, W. & Hochberg, F.G. (2009) Lectotype designations of new species of hydroids (Cnidaria, Hydrozoa), described by C.M. Fraser, from Allan Hancock Pacific and Caribbean Sea expeditions. Zoologische Mededelingen, 83, 919–1058.

    Carlton, J.T. & Eldredge, L.G. (2009) Marine bioinvasions of Hawai‘i. The introduced and cryptogenic marine and estuarine animals and plants of the Hawaiian Archipelago. Bishop Museum Bulletin in Cultural and Environmental Studies, 4, 1–202.

    Carlton, J.T. & Eldredge, L.G. (2015) Update and revisions of The Marine bioinvasions of Hawai‘i: the introduced and cryptogenic marine and estuarine animals and plants of the Hawaiian Archipelago. Bishop Museum Bulletin in Zoology, 9, 25–47.

    Chakraborty, O. & Raghunathan, C. (2020) Notes on seven aglaopheniids (Cnidaria: Hydrozoa: Aglaopheniidae) from Andaman and Nicobar islands with three new records to India. Zootaxa, 4790 (2), 291–317.

    https://doi.org/10.11646/zootaxa.4790.2.6

    Choong, H.H.C., Calder, D.R., Chapman, J.W., Miller, J.A., Geller, J.B. & Carlton, J.T. (2018) Hydroids (Cnidaria: Hydrozoa: Leptothecata and Limnomedusae) on 2011 Japanese tsunami marine debris landing in North America and Hawai‘i, with revisory notes on Hydrodendron Hincks, 1874 and a diagnosis of Plumaleciidae, new family. Aquatic Invasions, 13, 43–70.

    https://doi.org/10.3391/ai.2018.13.1.05

    Chu, G.W.T.C. & Cutress, C.E. (1954) Human dermatitis caused by marine organisms in Hawaii. Proceedings of the Hawaiian Academy of Science, 29, 9. [abstract]

    Ciamician, J. (1880) Ueber Lafoea parasitica n. sp. Zeitschrift für Wissenschaftliche Zoologie, 33, 673–676.

    Clark, S.F. (1875) Descriptions of new and rare species of hydroids from the New England coast. Transactions of the Connecticut Academy of Arts and Sciences, 3, 58–66.

    Clark, S.F. (1877) Report on the hydroids collected on the coast of Alaska and the Aleutian Islands, by W. H. Dall, U.S. Coast Survey, and party, from 1871 to 1874 inclusive. Proceedings of the Academy of Natural Sciences of Philadelphia, 1876, 28, 209–238.

    Cockerell, T.D.A. (1911) The nomenclature of the hydromedusae. Proceedings of the Biological Society of Washington, 24, 77–86.

    Coles, S.L., Bolick, H., Hauk, B. & Montgomery, A. (2009) Ten year resurveys of the biodiversity of marine communities and introduced species in Pearl Harbor, Honolulu Harbor, and Ke‘ehi Lagoon, O‘ahu, Hawai‘i. Bishop Museum Technical Report, 48, 1–204.

    Coles, S.L., DeFelice, R.C. & Eldredge, L.G. (1999) Nonindigenous marine species introductions in the harbors of the south and west shores of Oahu, Hawaii. Final report prepared for the David and Lucile Packard Foundation. Bishop Museum Technical Report, 15, 1–212.

    Coles, S.L., DeFelice, R.C. & Eldredge, L.G. (2002a) Nonindigenous marine species in Kane‘ohe Bay, Hawai‘i. Bishop Museum Technical Report, 24, 1–353.

    Coles, S.L., DeFelice, R.C. & Eldredge, L.G. (2002b) Nonindigenous marine species at Waikîkî and Hawai‘i Kai, O‘ahu, Hawai‘i. Bishop Museum Technical Report, 25, 1–245.

    Coles, S.L., Kandel, F.L.M., Reath, P.A., Longenecker, K. & Eldredge, L.G. (2006) Rapid assessment of nonindigenous marine species on coral reefs in the main Hawaiian Islands. Pacific Science, 60, 483–507.

    https://doi.org/10.1353/psc.2006.0026

    Coles, S.L., Reath, P.R., Longenecker, K., Bolick, H. & Eldredge, L.G. (2004) Assessment of nonindigenous marine species in harbors and on nearby coral reefs on Kaua‘i, Moloka‘i, Maui, and Hawai‘i. Bishop Museum Technical Report, 29, 1–187.

    Cooke, W.J. (1975) Shallow water hydroids from Eniwetak Atoll, Marshall Islands. Micronesica, 11, 85–108.

    Cooke, W.J. (1977) Order Hydroida. In: Devaney, D.M. & Eldredge, L.G. (Eds.), Reef and shore fauna of Hawaii. Section 1: Protozoa through Ctenophora. Bernice P. Bishop Museum, Special Publication, 64 (1), 71–104.

    Cornelius, P.F.S. (1975) The hydroid species of Obelia (Coelenterata, Hydrozoa: Campanulariidae), with notes on the medusa stage. Bulletin of the British Museum (Natural History), Zoology, 28, 249–293.

    Cornelius, P.F.S. (1992a) Medusa loss in leptolid Hydrozoa (Cnidaria), hydroid rafting, and abbreviated life-cycles among their remote-island faunae: an interim review. Scientia Marina, 56, 245–261.

    Cornelius, P.F.S. (1992b) The Azores hydroid fauna and its origin, with discussion of rafting and medusa suppression. Arquipélago. Life and Earth Sciences, 10, 75–99.

    Cornelius, P.F.S. (1995a) North-west European thecate hydroids and their medusae. Part 1. Introduction, Laodiceidae to Haleciidae. Synopses of the British Fauna, New Series, 50, 1–347.

    Cornelius, P.F.S. (1995b) North-west European thecate hydroids and their medusae. Part 2. Sertulariidae to Campanulariidae. Synopses of the British Fauna, New Series, 50, 386 pp.

    Cunha, A.F., Collins, A.G. & Marques, A.C. (2017) Phylogenetic relationships of Proboscoida Broch, 1910 (Cnidaria, Hydrozoa): are traditional morphological diagnostic characters relevant for the delimitation of lineages at the species, genus, and family levels? Molecular Phylogenetics and Evolution, 106, 118–135.

    https://doi.org/10.1016/j.ympev.2016.09.012

    Cunha, A.F., Collins, A.G. & Marques, A.C. (2020) When morphometry meets taxonomy: morphological variation and species boundaries in Proboscoida (Cnidaria: Hydrozoa). Zoological Journal of the Linnean Society, zlz166. [published online]

    https://doi.org/10.1093/zoolinnean/zlz166

    Cunha, A.F., Genzano, G.N. & Marques, A.C. (2015) Reassessment of morphological diagnostic characters and species boundaries requires taxonomical changes for the genus Orthopyxis L. Agassiz, 1862 (Campanulariidae, Hydrozoa) and some related campanulariids. PLoS ONE, 10 (2), e0117553.

    https://doi.org/10.1371/journal.pone.0117553

    De Oreo, G.A. (1946) Dermatitis venenata resulting from contact with marine animals (hydroids). Archives of Dermatology and Syphilology, 54, 637–649.

    https://doi.org/10.1001/archderm.1946.01510410003001

    Di Camillo, C.G., Bavestrello, G., Valisano, L. & Puce, S. (2008) Spatial and temporal distribution in a tropical hydroid assemblage. Journal of the Marine Biological Association of the United Kingdom, 88, 1589–1599.

    https://doi.org/10.1017/S0025315408002981

    Di Camillo, C.G., Puce, S. & Bavestrello, G. (2009) Macrorhynchia species (Cnidaria: Hydrozoa) from the Bunaken Marine Park (North Sulawesi, Indonesia) with a description of two new species. Italian Journal of Zoology, 76, 208–228.

    https://doi.org/10.1080/11250000802182018

    Edmondson, C.H. (1946) Reef and shore fauna of Hawaii. Revised edition. Bernice P. Bishop Museum Special Publication 22. Bishop Museum Press, Honolulu, 381 pp.

    Fernandez, M.O. & Marques, A.C. (2018) Combining bathymetry, latitude, and phylogeny to understand the distribution of deep Atlantic hydroids (Cnidaria). Deep-Sea Research, Part I, 133, 39–48.

    https://doi.org/10.1016/j.dsr.2018.01.008

    Fleming, J. (1828) A history of British animals, exhibiting the descriptive characters and systematical arrangement of the genera and species of quadrupeds, birds, reptiles, fishes, Mollusca, and Radiata of the United Kingdom. Bell & Bradfute, Edinburgh, 565 pp.

    https://doi.org/10.5962/bhl.title.12859

    Fraser, C.M. (1912) Some hydroids of Beaufort, North Carolina. Bulletin of the United States Bureau of Fisheries, 30, 339–387.

    Fraser, C.M. (1937) New species of hydroids from the Puerto Rican region. Smithsonian Miscellaneous Collections, 91 (28), 1–7.

    Fraser, C.M. (1938a) Hydroids of the 1934 Allan Hancock Pacific Expedition. Allan Hancock Pacific Expeditions, 4 (1), 1–105.

    Fraser, C.M. (1938b) Hydroids of the 1932, 1933, 1935, and 1938 Allan Hancock Pacific Expeditions. Allan Hancock Pacific Expeditions, 4 (3), 129–153.

    Fraser, C.M. (1944) Hydroids of the Atlantic coast of North America. University of Toronto Press, Toronto, 451 pp.

    Fraser, C.M. (1948) Hydroids of the Allan Hancock Pacific Expeditions since March, 1938. Allan Hancock Pacific Expeditions, 4 (5), 179–343.

    Fuhrmann, O. (1939) Sur Craspedacusta sowerbyi Lank. et un nouveau coelentéré d’eau douce, Calpasoma dactyloptera, n.g., n.sp. (note préliminaire). Revue Suisse de Zoologie, 46, 363–368.

    Galea, H.R. (2006) On two new species of Halopteris Allman, 1877 (Cnidaria: Hydrozoa) from Chile. Zootaxa, 1165 (1), 57–68.

    https://doi.org/10.11646/zootaxa.1165.1.4

    Galea, H.R. (2008) On a collection of shallow-water hydroids (Cnidaria: Hydrozoa) from Guadeloupe and Les Saintes, French Lesser Antilles. Zootaxa, 1878 (1), 1–54.

    https://doi.org/10.11646/zootaxa.1878.1.1

    Galea, H.R. (2010) Additional shallow-water thecate hydroids (Cnidaria: Hydrozoa) from Guadeloupe and Les Saintes, French Lesser Antilles. Zootaxa, 2570 (1), 1–40.

    https://doi.org/10.11646/zootaxa.2570.1.1

    Galea, H.R. (2016) Notes on some sertulariid hydroids (Cnidaria: Hydrozoa) from the tropical western Pacific, with descriptions of nine new species. European Journal of Taxonomy, 218, 1–52.

    https://doi.org/10.5852/ejt.2016.218

    Galea, H.R. (2018) A swimming medusoid gonophore in the life cycle of Ventromma halecioides (Alder, 1859) (Hydrozoa: Leptothecata: Kirchenpaueriidae). Revue Suisse de Zoologie, 125, 137–147.

    Galea, H.R. (2020) Aglaopheniid hydroids (Cnidaria: Hydrozoa: Aglaopheniidae) from off New Caledonia collected during KANACONO and KANADEEP expeditions of the French Tropical Deep-Sea Benthos Program. European Journal of Taxonomy, 615, 1–47.

    https://doi.org/10.5852/ejt.2020.615

    Galea, H.R., Di Camillo, C.G., Maggioni, D., Montano, S. & Schuchert, P. (2018) A reassessment of Halopteris polymorpha (Billard, 1913 (Cnidaria: Hydrozoa), with descriptions of three new species. Revue Suisse de Zoologie, 125, 21–59.

    Galea, H.R. & Ferry, R. (2015) Notes on some hydroids (Cnidaria) from Martinique, with descriptions of five new species. Revue Suisse de Zoologie, 122, 213–246.

    Galea, H.R., Ferry, R. & Bertot, J.-M. (2012) Medusoids in the life cycle of Dentitheca dendritica (Nutting, 1900) and Nemalecium gracile sp. nov. (Cnidaria: Hydrozoa). Zootaxa, 3527, 43–54.

    https://doi.org/10.11646/zootaxa.3527.1.3

    Gibbons, M.J. & Ryland, J.S. (1989) Intertidal and shallow water hydroids from Fiji. I. Athecata to Sertulariidae. Memoirs of the Queensland Museum, 27, 377–432.

    Godwin, L.S. (2003) Hull fouling of marine vessels as a pathway for marine species invasions to the Hawaiian Islands. Biofouling, 19, 123–131.

    https://doi.org/10.1080/0892701031000061750

    Govindarajan, A.F., Boero, F. & Halanych, K.M. (2006) Phylogenetic analysis with multiple markers indicates repeated loss of the adult medusa stage in Campanulariidae (Hydrozoa, Cnidaria). Molecular Phylogenetics and Evolution, 38, 820–834.

    https://doi.org/10.1016/j.ympev.2005.11.012

    Gravier-Bonnet, N. (1979) Hydraires semi-profonds de Madagascar, (Coelenterata Hydrozoa), étude systématique et écologique. Zoologische Verhandelingen, 169, 1–76.

    Gravier-Bonnet, N. & Bourmaud, C.A.-F. (2012) Hydroids (Cnidaria, Hydrozoa) of Baa Atoll (Indian Ocean, Maldives Archipelago). In: Biodiversity, resources, and conservation of Baa Atoll (Republic of Maldives). A UNESCO Man and Biosphere Reserve. Atoll Research Bulletin, 594, 85–123.

    Gravier-Bonnet, N. & Migotto, A.E. (2000) Gonangium development and medusoid of Nemalecium lighti (Hargitt, 1924) (Cnidaria: Hydrozoa, Haleciidae). Scientia Marina, 64 (Suplemento 1), 207–213.

    https://doi.org/10.3989/scimar.2000.64s1207

    Gravili, C., De Vito, D., Di Camillo, C.G., Martell, L., Piraino, S. & Boero, F. (2015) The non-siphonophoran Hydrozoa (Cnidaria) of Salento, Italy with notes on their life-cycles: an illustrated guide. Zootaxa, 3908 (1), 1–187.

    https://doi.org/10.11646/zootaxa.3908.1.1

    Gray, J.E. (1848) List of the specimens of British animals in the collection of the British Museum. Part I. Centroniae or radiated animals. British Museum, London, 173 pp.

    Grovhoug, J.G. (1979) Marine environmental assessment at three sites in Pearl Harbor, Oahu, August–October 1978. Naval Ocean Systems Center, San Diego, California, Technical Report, 441, 1–91.

    Grovhoug, J.G. & Rastetter, E.B. (1980) Marine fouling dynamics in Hawaiian nearshore ecosystems: a suggested technique for comparison and evaluation. In: Arito, L. (Ed.), Comunicaciones V Congreso Internacional de Corrosión Marina e Incrustaciones, Barcelona. Gráficas Orbe S.L., Madrid, pp. 249–266.

    Haeckel, E. (1879) Das System der Medusen. Erster Theil einer Monographie der Medusen. Denkschriften der Medicinisch-Naturwissenschaftlichen Gesellschaft zu Jena, 1, 1–360.

    Hargitt, C.W. (1924) Hydroids of the Philippine Islands. Philippine Journal of Science, 24, 467–507.

    Hartlaub, C. (1901) Hydroiden aus dem Stillen Ocean. Ergebnisse einer Reise nach dem Pacific (Schauinsland 1896–97). Zoologische Jahrbücher, Abtheilung für Systematik, Geographie und Biologie der Thiere, 14, 349–379.

    https://doi.org/10.5962/bhl.part.11530

    Hincks, T. (1869) A history of the British hydroid zoophytes. John van Voorst, London, 338 pp. [Dating of this two-volume work follows Williams (2018), who presented evidence that it was published in March 1869 and not 1868 as per the title-pages]

    Hincks, T. (1874) Notes on Norwegian Hydroida from deep water. Annals and Magazine of Natural History, Series 4, 13, 125–137.

    https://doi.org/10.1080/00222937408680824

    Hirohito, The Showa Emperor (1969) Some hydrozoans of the Amakusa Islands. Biological Laboratory, Imperial Household, Tokyo, 32 pp.

    Hirohito, The Showa Emperor (1974) Some hydrozoans of the Bonin Islands. Biological Laboratory, Imperial Household, Tokyo, 55 pp.

    Hirohito, The Showa Emperor (1983) Hydroids from Izu Ôshima and Niijima. Biological Laboratory, Imperial Household, Tokyo, 83 pp.

    Hirohito, The Showa Emperor (1995) The hydroids of Sagami Bay. Part II. Thecata. Biological Laboratory, Imperial Household, Tokyo, 355 pp.

    Hoover, J.P. (1998) Hawai‘i’s sea creatures. A guide to Hawai‘i’s marine invertebrates. Mutual Publishing, LLC, Honolulu, 366 pp.

    Hoover, J.P. (2006) Hawai‘i’s sea creatures. A guide to Hawai‘i’s marine invertebrates. Revised Edition. Mutual Publishing, LLC, Honolulu, 366 pp.

    Humara-Gil, K.J. & Cruz-Gómez, C. (2018) New records of benthic hydroids (Cnidaria: Hydrozoa) from the coast of Oaxaca, Mexico. Zootaxa, 4455 (3), 454–470.

    https://doi.org/10.11646/zootaxa.4455.3.3

    International Commission on Zoological Nomenclature (1998) Opinion 1886. Plumularia Lamarck, 1816 (Cnidaria, Hydrozoa): conserved by the designation of Sertularia setacea Linnaeus, 1758 as the type species. Bulletin of Zoological Nomenclature, 55, 47–48.

    Jäderholm, E. (1919) Zur Kenntnis der Hydroidenfauna Japans. Arkiv för Zoologi, 12 (9), 1–34.

    https://doi.org/10.5962/bhl.part.789

    Jarvis, F.E. (1922) The hydroids from the Chagos, Seychelles and other islands and from the coasts of British East Africa and Zanzibar. Transactions of the Linnean Society of London, Zoology, 18, 331–360.

    https://doi.org/10.1111/j.1096-3642.1922.tb00553.x

    Jickeli, C.F. (1883) Der Bau der Hydroidpolypen. II. Über den histiologischen Bau von Tubularia L., Cordylophora Allm., Cladonema Duj., Coryne Gärtn., Gemmaria M’Crady, Perigonimus Sars, Podocoryne Sars, Camponopsis Claus, Lafoëa Lam., Campanularia Lam., Obelia Pér., Anisocola Kirchenp., Isocola Kirchenp., Kirchenpaueria Jick. Morphologisches Jahrbuch, 8, 580–680.

    Johnston, G. (1833) Illustrations in British zoology. 15. Plumularia Catharina. Magazine of Natural History, and Journal of Zoology, Botany, Mineralogy, Geology, and Meteorology, 6, 497–499.

    Johnston, G. (1837) A catalogue of the zoophytes of Berwickshire. History of the Berwickshire Naturalists’ Club, 1, 107–108.

    Johnston, G. (1838) A history of the British zoophytes. W.H. Lizars, Edinburgh, 341 pp.

    https://doi.org/10.5962/bhl.title.110844

    Kirchenpauer, G.H. (1864) Neue Sertulariden aus verschiedenen Hamburgischen Sammlungen, nebst allgemeinen Bemerkungen über Lamouroux’s Gattung Dynamena. Verhandlungen der Kaiserlichen Leopoldino-Carolinischen Deutschen Akademie der Naturforscher, 31 (3), 1–16.

    Kirchenpauer, G.H. (1872) Ueber die Hydroidenfamilie Plumularidae, einzelne Gruppen derselben und ihre Fruchtbehälter. I. Aglaophenia Lx. Abhandlungen aus dem Gebiete der Naturwissenschaften herausgegeben von dem Naturwissenschaftlichen Verein in Hamburg, 5 (3), 1–52.

    Kirkendale, L & Calder, D.R. (2003) Hydroids (Cnidaria: Hydrozoa) from Guam and the Commonwealth of the Northern Marianas Islands (CNMI). Micronesica, 35–36, 159–188.

    Kramp, P.L. (1959) Some new and little-known Indo-Pacific medusae. Videnskabelige Meddelelser fra Dansk Naturhistorisk Forening, 121, 223–259.

    Lamarck, J.B.P.A. de (1816) Histoire naturelle des animaux sans vertebrès. Tome 2. Verdière, Paris, 568 pp.

    Lamouroux, J.V.F. (1812) Extrait d’un mémoire sur la classification des polypiers coralligènes non entièrement pierreux. Nouveau Bulletin des Sciences, par la Société Philomatique de Paris, 3, 181–188.

    Lamouroux, J.V.F. (1816) Histoire des polypiers coralligènes flexibles, vulgairement nommés zoophytes. F. Poisson, Caen, 560 pp.

    https://doi.org/10.5962/bhl.title.11172

    Lamouroux, J.V.F. (1824) Description des polypiers flexibles. In: Quoy, J.R.C. & Gaimard, J.P. (Eds.), Zoologie. Voyage autour du monde, entrepris par ordre du Roi, exécuté sur les corvettes de S.M. l’Uranie et la Physicienne, pendant les années 1817, 1818, 1819 et 1820; par M. Louis de Freycinet. Pillet Aîné, Paris, pp. 603–643.

    Lankester, E.R. (1880) On a new jelly-fish of the order Trachomedusae, living in fresh water. Nature, 22, 147–148.

    https://doi.org/10.1038/022147a0

    Leclère, L., Schuchert, P., Cruaud, C., Couloux, A. & Manuel, M. (2009) Molecular phylogenetics of Thecata (Hydrozoa, Cnidaria) reveals long-term maintenance of life history traits despite high frequency of recent character changes. Systematic Biology, 58, 509–526.

    https://doi.org/10.1093/sysbio/syp044

    Leclère, L., Schuchert, P. & Manuel, M. (2007) Phylogeny of the Plumularioidea (Hydrozoa, Leptothecata): evolution of colonial organization and life cycle. Zoologica Scripta, 36, 371–394.

    https://doi.org/10.1111/j.1463-6409.2007.00283.x

    Leloup, E. (1930) Coelentérés hydropolypes. In: Résultats scientifiques du voyage aux Indes orientales néerlandaises de LL. AA. RR. le Prince et la Princesse Léopold de Belgique. Mémoires du Musée Royale d’Histoire Naturelle de Belgique, Hors Série, 2, 1–18.

    Leloup, E. (1935) Hydropolypes calyptoblastiques et siphonophores récoltés au cours de la croisière (1934–1935) du navire-école belge “Mercator”. Bulletin du Musée Royal d’Histoire Naturelle de Belgique, 11 (34), 1–6.

    Lindner, A., Govindarajan, A.F. & Migotto, A.E. (2011) Cryptic species, life cycles, and the phylogeny of Clytia (Cnidaria: Hydrozoa: Campanulariidae). Zootaxa, 2980 (1), 23–36.

    https://doi.org/10.11646/zootaxa.2980.1.2

    Linnaeus, C. (1758) Systema naturæ per regna tria naturæ, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Editio decima, reformata. Laurentii Salvii, Holmiae, 823 pp.

    https://doi.org/10.5962/bhl.title.542

    Linnaeus, C. (1767) Systema naturae per regna tria naturæ, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Tomus I. Pars II. Editio duodecima, reformata. Laurentii Salvii, Holmiae, pp. 533–1317.

    https://doi.org/10.5962/bhl.title.156765

    Mammen, T.A. (1965) On a collection of hydroids from south India. II. Suborder Thecata (excluding family Plumulariidae). Journal of the Marine Biological Association of India, 7, 1–57.

    Marktanner-Turneretscher, G. (1890) Die Hydroiden des k. k. naturhistorischen Hofmuseums. Annalen des Kaiserlich-Königlichen Naturhistorischen Hofmuseums, 5, 195–286.

    Maronna, M.M., Miranda, T.P., Peña Cantero, Á.L., Barbeitos, M.S. & Marques, A.C. (2016) Towards a phylogenetic classification of Leptothecata (Cnidaria, Hydrozoa). Scientific Reports, 6, 18075. [published online]

    https://doi.org/10.1038/srep18075

    Marques, A.C., Haddad Jr., V. & Migotto, A.E. (2002) Envenomation by a benthic Hydrozoa (Cnidaria): the case of Nemalecium lighti (Haleciidae). Toxicon, 40, 213–215.

    https://doi.org/10.1016/S0041-0101(01)00196-9

    McCrady, J. (1859) Gymnopthalmata of Charleston Harbor. Proceedings of the Elliott Society of Natural History, 1, 103–221.

    Medel, M.D. & Vervoort, W. (1995) Plumularian hydroids (Cnidaria: Hydrozoa) from the Strait of Gibraltar and nearby areas. Zoologische Verhandelingen, 300, 1–72.

    Medel, M.D. & Vervoort, W. (1998) Atlantic Thyroscyphidae and Sertulariidae (Hydrozoa, Cnidaria) collected during the CANCAP and Mauritania-II expeditions of the National Museum of Natural History, Leiden, The Netherlands. Zoologische Verhandelingen, 320, 1–85.

    Medel, M.D. & Vervoort, W. (2000) Atlantic Haleciidae and Campanulariidae (Hydrozoa, Cnidaria) collected during the CANCAP and Mauritania-II expeditions of the National Museum of Natural History, Leiden, The Netherlands. Zoologische Verhandelingen, 330, 1–68.

    Mergner, H. & Wedler, E. (1977) Über die Hydroidpolypenfauna des Roten Meeres und seiner Ausgänge. “Meteor” Forschungs-Ergebnisse, Reihe D, 24, 1–32.

    Migotto, A.E. (1996) Benthic shallow-water hydroids (Cnidaria, Hydrozoa) of the coast of São Sebastião, Brazil, including a checklist of Brazilian hydroids. Zoologische Verhandelingen, 306, 1–125.

    Migotto, A.E. & Cabral, A.S. (2005) Lafoeina amirantensis (Cnidaria: Hydrozoa, Campanulinoidea), the hydroid stage of the medusa Cirrholovenia tetranema (Cnidaria: Hydrozoa, Lovenelloidea). Zootaxa, 919 (1), 1–16.

    https://doi.org/10.11646/zootaxa.919.1.1

    Millard, N.A.H. (1957) The Hydrozoa of False Bay, South Africa. Annals of the South African Museum, 43, 173–243.

    Millard, N.A.H. (1962) The Hydrozoa of the south and west coasts of South Africa. Part I. The Plumulariidae. Annals of the South African Museum, 46, 261–319.

    Millard, N.A.H. (1966) Hydroids of the Vema Seamount. Annals of the South African Museum, 48, 489–496.

    Millard, N.A.H. (1975) Monograph on the Hydroida of southern Africa. Annals of the South African Museum, 68, 1–513.

    Millard, N.A.H. & Bouillon, J. (1973) Hydroids from the Seychelles (Coelenterata). Annales du Musée Royal de l’Afrique Centrale, Série In-8°, Sciences Zoologiques, 206, 1–106.

    Morri, C. & Boero, F. (1986) Catalogue of main marine fouling organisms. Volume 7. Hydroids. Office d’Etudes Marines et Atmosphériques, ODEMA, Brussels, 91 pp.

    Moura, C.J., Collins, A.G., Santos, R.S. & Lessios, H. (2019) Predominant east to west colonizations across major oceanic barriers: insights into the phylogeographic history of the hydroid superfamily Plumularioidea, suggested by a mitochondrial DNA barcoding marker. Ecology & Evolution, 9 (23), 13001–13016.

    https://doi.org/10.1002/ece3.5608

    Moura, C.J., Harris, D.J., Cunha, M.R. & Rogers, A.D. (2008) DNA barcoding reveals cyptic diversity in marine hydroids (Cnidaria, Hydrozoa) from coastal and deep-sea environments. Zoologica Scripta, 37, 93–108.

    Moura, C.J., Lessios, H., Cortés, J., Nizinski, M.S., Reed, J., Santos, R.S. & Collins, A.G. (2018) Hundreds of genetic barcodes of the species-rich hydroid superfamily Plumularioidea (Cnidaria, Medusozoa) provide a guide toward more reliable taxonomy. Scientific Reports, 8, 1–14.

    https://doi.org/10.1038/s41598-018-35528-8

    Mulder, J.F. & Trebilcock, R.E. (1911) Notes on Victorian Hydroida, with description of new species. (Continued). Geelong Naturalist, Series 2, 4, 115–124.

    Mulder, J.F. & Trebilcock, R.E. (1915) Victorian Hydroida. With description of new species. Part V. Geelong Naturalist, 6 (3), 51–59.

    Nutting, C.C. (1900) American hydroids. Part I. The Plumularidae. Smithsonian Institution, United States National Museum Special Bulletin, 4 (1), 1–285.

    Nutting, C.C. (1904) American hydroids. Part II. The Sertularidae. Smithsonian Institution, United States National Museum Special Bulletin, 4 (2), 1–325.

    Nutting, C.C. (1905) Hydroids of the Hawaiian Islands collected by the steamer Albatross in 1902. Bulletin of the United States Fish Commission for 1903, pp. 931–959. [A statement in the “Contents” section of the Bulletin records that Nutting’s article, variously dated in the literature, was “Issued December 23, 1905” (Calder 2017: 68).]

    Nutting, C.C. (1915) American hydroids. Part III. The Campanularidae and the Bonneviellidae. Smithsonian Institution, United States National Museum Special Bulletin, 4 (3), 1–126.

    Nutting, C.C. (1927) Report on the Hydroida collected by the United States fisheries steamer “Albatross” in the Philippine region, 1907-1910. Bulletin of the United States National Museum, 100, 195–242.

    Oken, L. (1815) Okens Lehrbuch der Naturgeschichte. III. Theil. Zoologie. Vol. 1. Oken, Jena, 842 pp.

    Oliveira, O.M.P., Miranda, T.P., Araujo, E.M., Ayón, P., Cedeño-Posso, C.M., Cepeda-Mercado, A.A., Córdova, P., Cunha, A.F., Genzano, G.N., Haddad, M.A., Mianzan, H.W., Migotto, A.E., Miranda, L.S., Morandini, A.C., Nagata, R.M., Nascimento, K.B., Nogueira Jr., M., Palma, S., Quiñones, J., Rodriguez, C.S., Scarabino, F., Schiariti, A., Stampar, S.N., Tronolone, V.B. & Marques, A.C. (2016) Census of Cnidaria (Medusozoa) and Ctenophora from South American marine waters. Zootaxa, 4194 (1), 1–256.

    https://doi.org/10.11646/zootaxa.4194.1.1

    Park, J.H. (1993) Marine hydroids (Cnidaria: Hydrozoa: Hydroida) from Cheju Island, Korea. Korean Journal of Systematic Zoology, 9, 261–280.

    Peña Cantero, Á.L., Sentandreu, V. & Latorre, A. (2010) Phylogenetic relationships of the endemic Antarctic benthic hydroids (Cnidaria, Hydrozoa): what does the mitochondrial 16S rRNA tell us about it? Polar Biology, 33, 41–57.

    https://doi.org/10.1007/s00300-009-0683-5

    Péron, F. & Lesueur, C.A. (1810) Tableau des caractères génériques et spécifiques de toutes les espèces de méduses connues jusqu’à ce jour. Annales du Muséum d’Histoire Naturelle, 14, 325–366.

    Pictet, C. (1893) Voyage de Mm. M. Bedot et C. Pictet dans l’Archipel Malais. Étude sur les hydraires de la Baie d’Amboine. Revue Suisse de Zoologie et Annales du Musée d’Histoire Naturelle de Genève, 1, 1–64.

    https://doi.org/10.5962/bhl.part.3743

    Postaire, B., Magalon, H., Bourmaud, C.A.-F., Gravier-Bonnet, N. & Bruggemann, J.H. (2016) Phylogenetic relationships within Aglaopheniidae (Cnidaria, Hydrozoa) reveal unexpected generic diversity. Zoologica Scripta, 45, 103–114.

    https://doi.org/10.1111/zsc.12135

    Ralph, P.M. (1957) New Zealand thecate hydroids. Part I.—Campanulariidae and Campanulinidae. Transactions of the Royal Society of New Zealand, 84, 811–854.

    Randall, J.E. (1975) Notes on the Hawaiian filefish Pseudomonacanthus garretti. Japanese Journal of Ichthyology, 21, 223–226.

    Rees, W.J. & Thursfield, S. (1965) The hydroid collections of James Ritchie. Proceedings of the Royal Society of Edinburgh, Section B (Biology), 69, 34–220.

    https://doi.org/10.1017/S0080455X00010122

    Rees, W.J. & Vervoort, W. (1987) Hydroids from the John Murray Expedition to the Indian Ocean, with revisory notes on Hydrodendron, Abietinella, Cryptolaria and Zygophylax (Cnidaria, Hydrozoa). Zoologische Verhandelingen, 237, 1–209.

    Rho, B.J. (1967) Marine hydroids from the west and south sea of Korea. Papers of the Korean Culture Research Institute, 10, 341–360.

    Rho, B.J. & Park, J.H. (1986) A systematic study on the marine hydroids in Korea. 10. The family Plumulariidae. Journal of Korean Research Institute for Better Living, Ewha Womans University, 37, 87–112.

    Richards, R. (2000) Honolulu: centre of trans-Pacific trade: shipping arrivals and departures 1820 to 1840. Pacific Manuscripts Bureau, Canberra; Hawaiian Historical Society, Honolulu, 326 pp. [not seen]

    Ritchie, J. (1911) Hydrozoa (hydroid zoophytes and Stylasterina) of the “Thetis” Expedition. Australian Museum, Sydney, Memoir, 4, 805–869.

    https://doi.org/10.3853/j.0067-1967.4.1911.1512

    Ronowicz, M., Boissin, E., Postaire, B., Bourmaud, C.A.-F., Gravier-Bonnet, N. & Schuchert, P. (2017) Modern alongside traditional taxonomy—integrative systematics of the genera Gymnangium Hincks, 1874 and Taxella Allman, 1874 (Hydrozoa, Aglaopheniidae). PLoS ONE, 12 (4), e0174244.

    https://doi.org/10.1371/journal.pone.0174244

    Ryland, J.S. & Gibbons, M.J. (1991) Intertidal and shallow water hydroids from Fiji. II. Plumulariidae and Aglaopheniidae. Memoirs of the Queensland Museum, 30, 525–560.

    Sars, M. (1850) Beretning om en i Sommeren 1849 foretagen zoologisk Reise i Lofoten og Finmarken. Nyt Magazin for Naturvidenskaberne, 6, 121–211.

    Schuchert, P. (1997) Review of the family Halopterididae (Hydrozoa, Cnidaria). Zoologische Verhandelingen, 309, 1–162.

    Schuchert, P. (2003) Hydroids (Cnidaria, Hydrozoa) of the Danish Expedition to the Kei Islands. Steenstrupia, 27, 137–256.

    Schuchert, P. (2012) North-west European athecate hydroids and their medusae. Synopses of the British Fauna, New Series, 59, 1–364.

    Schuchert, P. (2014) High genetic diversity in the hydroid Plumularia setacea: a multitude of cryptic species or extensive population subdivision? Molecular Phylogenetics and Evolution, 76, 1–9.

    https://doi.org/10.1016/j.ympev.2014.02.020

    Schuchert, P. (2015) On some hydroids (Cnidaria, Hydrozoa) from the Okinawa Islands, Japan. Revue Suisse de Zoologie, 122, 325–370.

    Schuchert, P. (2020) World Hydrozoa Database. Accessed through: World Register of Marine Species at: https://www.marinespecies.org/aphia.php?p=taxdetails&id=117001 (accessed 2 February 2020)

    Song, X., Lyu, M., Ruthensteiner, B., Wang, J. & Gravili, C. (2019) Unexpected systematic affinities and geographic expansion of a marine alien hydroid (Cnidaria: Hydrozoa). Systematics and Biodiversity, 17 (3), 230–244.

    https://doi.org/10.1080/14772000.2019.1583690

    Stechow, E. (1907) Neue japanische Athecata und Plumularidae aus der Sammlung Dr. Doflein. Zoologischer Anzeiger, 32, 192–200.

    Stechow, E. (1909) Beiträge zur Naturgeschichte Ostasiens. Herausgegeben von Dr. F. Doflein. Hydroidpolypen der japanischen Ostküste. I. Teil: Athecata und Plumularidae. Abhandlungen der Mathematisch-Physikalischen Klasse der Königlichen Bayerischen Akademie der Wissenschaften, Supplement-Band 1 (6), 1–111.

    Stechow, E. (1913) Hydroidpolypen der japanischen Ostküste. II. Teil: Campanularidae, Halecidae, Lafoeidae, Campanulinidae und Sertularidae, nebst Ergänzungen zu den Athecata und Plumularidae. Abhandlungen der Mathematisch-Physikalischen Klasse der Königlichen Bayerischen Akademie der Wissenschaften, 3 (Supplement-Band 2), 1–162.

    https://doi.org/10.5962/bhl.title.11621

    Stechow, E. (1914) Zur Kenntnis neuer oder seltener Hydroidpolypen, meist Campanulariden, aus Amerika und Norwegen. Zoologischer Anzeiger, 45, 120–136.

    Stechow, E. (1919) Zur Kenntnis der Hydroidenfauna des Mittelmeeres, Amerikas und anderer Gebiete, nebst Angaben über einige Kirchenpauer’sche Typen von Plumulariden. Zoologische Jahrbücher, Abteilung für Systematik, Geographie und Biologie der Tiere, 42, 1–172.

    Stechow, E. (1920) Neue Ergebnisse auf dem Gebiete der Hydroidenforschung. Sitzungsberichte der Gesellschaft für Morphologie und Physiologie in München, 31, 9–45.

    Stechow, E. (1921) Neue Genera und Species von Hydrozoen und anderen Evertebraten. Archiv für Naturgeschichte, Abteilung A, 3. Heft, 87, 248–265.

    Stechow, E. (1922) Zur Systematik der Hydrozoen, Stromatoporen, Siphonophoren, Anthozoen und Ctenophoren. Archiv für Naturgeschichte, Abteilung A, 3. Heft, 88, 141–155.

    Stechow, E. (1923) Zur Kenntnis der Hydroidenfauna des Mittelmeeres, Amerikas und anderer Gebiete. II. Teil. Zoologische Jahrbücher, Abteilung für Systematik, Ökologie und Geographie der Tiere 47, 29–270.

    Stechow, E. (1925) Hydroiden der Deutschen Tiefsee-Expedition. Wissenschaftliche Ergebnisse der Deutschen Tiefsee-Expedition auf dem Dampfer ‘’Valdivia’’ 1898–1899, 17, 383–546.

    Thornely, L.R. (1894) Supplementary report upon the hydroid zoophytes of the L.M.B.C. district. Proceedings and Transactions of the Liverpool Biological Society, 8, 140–147.

    Thornely, L.R. (1897) On the Hydrozoa. In: Green, W.S., Notes on Rockall Island and Bank, with an account of the petrology of Rockall, and of its winds, currents, etc.: with reports on the ornithology, the invertebrate fauna of the bank, and on its previous history. Transactions of the Royal Irish Academy, 31, 81.

    Thornely, L.R. (1900) The hydroid zoophytes collected by Dr. Willey in the southern seas. In: Willey, A., Zoological results based on material from New Britain, New Guinea, Loyalty Islands and elsewhere. Part IV. Cambridge University Press, Cambridge, pp. 451–457.

    Thornely, L.R. (1904) Report on the Hydroida collected by Professor Herdman, at Ceylon, in 1902. In: Herdman, W.A., Report to the Government of Ceylon on the pearl oyster fisheries of the Gulf of Manaar, with supplementary reports upon the marine biology of Ceylon, by other naturalists, Part 2, pp. 107–126.

    Thornely, L.R. (1908) Reports on the marine biology of the Sudanese Red Sea.–X. Hydroida collected by Mr. C. Crossland from October 1904 to May 1905. Journal of the Linnean Society of London, Zoology, 31, 80–85.

    https://doi.org/10.1111/j.1096-3642.1908.tb00456.x

    Thornely, L.R. (1916) Report on the Hydroida collected by Mr. James Hornell at Okhamandal in Kattiawar in 1905–06. In: Hornell, J., Report to the Government of Baroda on the marine zoology of Okhamandal in Kattiawar. Part Π. Williams & Norgate, London, pp. 147–150.

    Torrey, H.B. (1902) The Hydroida of the Pacific coast of North America, with especial reference to the species in the collection of the University of California. University of California Publications, Zoology, 1, 1–105.

    Vanhöffen, E. (1910) Die Hydroiden der Deutschen Südpolar-Expedition 1901–1903. Deutsche Südpolar-Expedition 1901– 1903, 11, Zoologie 3, 269–340.

    Vannucci Mendes, M. (1946) Hydroida Thecaphora do Brasil. Arquivos de Zoologia do Estado de São Paulo, 4, 535–597.

    Vervoort, W. (1959) The Hydroida of the tropical west coast of Africa. Atlantide Report, Scientific Results of the Danish Expedition to the Coasts of Tropical West Africa, 1945–1946, 5, 211–325.

    Vervoort, W. (1968) Report on a collection of Hydroida from the Caribbean region, including an annotated checklist of Caribbean hydroids. Zoologische Verhandelingen, 92, 1–124.

    Vervoort, W. (1993) Cnidaria, Hydrozoa, Hydroida : hydroids from the western Pacific (Philippines, Indonesia and New Caledonia). I: Sertulariidae (Part 1). In: Crosnier, A. (Ed.), Résultats des campagnes MUSORSTOM. Vol. 11. Mémoires du Muséum National d’Histoire Naturelle, 158, pp. 89–298.

    Vervoort, W. (1995) Bibliography of Leptolida (non-siphonophoran Hydrozoa, Cnidaria). Works published after 1910. Zoologische Verhandelingen, 301, 1–432.

    Vervoort, W. & Vasseur, P. (1977) Hydroids from French Polynesia with notes on distribution and ecology. Zoologische Verhandelingen, 159, 1–98.

    Vervoort, W. & Watson, J.E. (2003) The marine fauna of New Zealand: Leptothecata (Cnidaria: Hydrozoa) (thecate hydroids). National Institute of Water and Atmospheric Research Biodiversity Memoir, 119, 1–538.

    Wagner, D., Barkman, A., Spalding, H.L., Calcinai, B. & Godwin, S.L. (2016) A photographic guide to the benthic flora and fauna from mesophotic coral ecosystems in the Papahânaumokuâkea Marine National Monument. Marine Sanctuaries Conservation Series ONMS-16-04. U.S. Department of Commerce, National Oceanic and Atmospheric Administration, Office of National Marine Sanctuaries, Silver Spring, Maryland, 86 pp.

    Watson, J.E. (2000) Hydroids (Hydrozoa: Leptothecatae) from the Beagle Gulf and Darwin Harbour, northern Australia. The Beagle, Records of the Museums and Art Galleries of the Northern Territory, 16, 1–82.

    Watson, J.E. (2005) Hydroids of the Archipelago of the Recherche and Esperance, Western Australia: annotated list, redescription of species and description of new species. In: Wells, F.E., Walker, D.I. & Kendrick, G.A. (Eds.), The marine flora and fauna of Esperance, Western Australia. Western Australian Museum, Perth, pp. 495–611.

    Watson, J.E. (2011) Review of the genus Monotheca (Hydrozoa: Leptolida) from Australia with description of a new species and a note on Monothecella Stechow, 1923. Memoirs of Museum Victoria, 68, 71–91.

    https://doi.org/10.24199/j.mmv.2011.68.05

    Watson, J.E. (2012) Some hydroids (Hydrozoa: Hydroidolina) from Dampier, Western Australia: annotated list with description of two new species. Memoirs of Museum Victoria, 69, 355–363.

    https://doi.org/10.24199/j.mmv.2012.69.09

    Williams, R.B. (2018) Correction of the spurious 1868 publication date of Thomas Hincks’s A History of the British Hydroid Zoophytes to 1869, with a list of included new taxa and nomenclatural combinations, and a bibliographical description. Zoological Bibliography, 6 (2), 3–27.

    Woods Hole Oceanographic Institution (1952) Marine fouling and its prevention. United States Naval Institute, Annapolis, Maryland, 388 pp.

    Xu, Z.-Z., Huang, J.-Q., Lin, M., Guo, D.-H. & Wang, C.-G. (2014) The superclass Hydrozoa of the phylum Cnidaria in China. Vol. II. China Ocean Press, Beijing, pp. 459–945.

    Yamada, M. (1981) A guide to identification of principal fouling organisms (6) Hydroids. Marine Fouling, 3, 97–105. [in Japanese]

    https://doi.org/10.4282/sosj1979.3.97

    Yamada, M. & Kubota, S. (1987) Preliminary report on the marine hydroid fauna in Okinawa Islands. Galaxea, 6, 35–42.