Skip to main content Skip to main navigation menu Skip to site footer
Published: 2020-04-30

Recent family-group names (2013–2019) in the scale insects (Hemiptera: Sternorrhyncha: Coccomorpha)

Department of Life Sciences, The Natural History Museum, London, SW7 5BD, U.K.
Hemiptera nominal genus new ranks


A list is provided of scale insect (Hemiptera: Sternorrhyncha: Coccomorpha) family-group names published between 2013 and the end of 2019. This is a supplement to two previous lists by Williams, covering names published from 1814–1968, and those published from 1969–2012. As in the previous lists, each entry is headed by the nominal genus on which each family‑group name is based followed by any new ranks.



  1. Choi, J. & Lee, S. (2020) Molecular phylogeny of the family Coccidae (Hemiptera, Coccomorpha) with a discussion of their waxy ovisacs. Systematic Entomology, 45 (2), 396–414. [Note: this work was published online on 4December 2019]

    Danzig, E.M. & Gavrilov-Zimin, I.A. (2014) Palaearctic mealybugs (Homoptera: Coccinea: Pseudococcidae). Part1. Subfamily Phenacoccinae. Fauna of Russia and Neighbouring Countries. New Series, 418, 1–678.

    Fallén, C.F. (1814) Specimen Novam Hemiptera Disponendi Methodum Exibens. litteris Berlingianis, Lundae, 26 pp.

    Foldi, I. (2005) Ground pearls: a generic revision of the Margarodidae sensu stricto (Hemiptera: Sternorrhyncha: Coccoidea). Annales de la Société entomologique de France, New Series, 41, 81–125.

    García Morales, M., Denno, B.D., Miller, D.R., Miller, G.L., Ben-Dov, Y. & Hardy, N.B. (2020) ScaleNet: a literature-based model of scale insect biology and systematics. Available from: (accessed 12 February 2020)

    Gavrilov-Zimin, I.A. (2018) Ontonogenesis, morphology and higher classification of archaeococcids [Homoptera: Coccinea: Orthezioidea]. Zoosystematica Rossica, Supplementum 2, 1–260.

    Gullan, P.J. & Cook, L.G. (2007) Phylogeny and higher classification of the scale insects (Hemiptera: Sternorrhyncha: Coccoidea). Zootaxa, 1668 (1), 413–425.

    Hodgson, C.J. (2014) Phenacoleachia, Steingelia, Pityococcus and Puto – neococoids or archaeococcoids? An intuitive phylogenetic discussion based on adult male characters. Acta Zoologica Bulgarica, Supplement 6, 41–50.

    Hughes-Schrader, S. (1948) Cytology of coccids (Coccoidea-Homoptera). Advances in Genetics, 2, 127–203.

    Normark, B.B. (2003) The evolution of alternative genetic systems in insects. Annual Review of Entomology, 48, 397–423.

    Normark, B.B., Okusu, A., Morse, G.E., Peterson, D.A., Itioka, T. & Schneider, S.A. (2019) Phylogeny and classification of armored scale insects (Hemiptera: Coccomorpha: Diaspididae). Zootaxa, 4616 (1), 1–98.

    Vea, I.M. & Grimaldi, D.A. (2015) Diverse new scale insects (Hemiptera: Coccoidea) in amber from the Cretaceous and Eocene with a phylogenetic framework for fossil Coccoidea. American Museum Novitates, 3823, 1–80.

    Williams, D.J. (1969) The family-group names of the scale insects (Hemiptera: Coccoidea). Bulletin of the British Museum (Natural History), Entomology, 23, 315–341.

    Williams, D.J. (2013) Family-group names in the scale insects (Hemiptera: Sternorrhyncha: Coccoidea)—a supplement. Zootaxa, 3616 (4), 325–344.

How to Cite

WILLIAMS, D. (2020). Recent family-group names (2013–2019) in the scale insects (Hemiptera: Sternorrhyncha: Coccomorpha). Zootaxa, 4768(1), 143–145.