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Type: Article
Published: 2019-12-17
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An overview of the Ephydridae (Diptera) of Saudi Arabia

Centre for Environmental Research and Studies, Jazan University, P.O. Box 2095, Jazan, Kingdom of Saudi Arabia.
Centre for Environmental Research and Studies, Jazan University, P.O. Box 2095, Jazan, Kingdom of Saudi Arabia.
Centre for Environmental Research and Studies, Jazan University, P.O. Box 2095, Jazan, Kingdom of Saudi Arabia.
Institute of Biology, Opole University, ul. Oleska 22, 45-052 Opole, Poland.
Diptera Ephydridae faunistics new records shore flies Saudi Arabia

Abstract

Despite the species richness of Ephydridae world-wide (2000 species) and its prominent environmental roles as a minor pest and as a food for wildlife, only 13 species have been recorded from Saudi Arabia. Between 2012 and 2016, a biodiversity study of Diptera was conducted at Jazan, Asir, and Najran in south-western Saudi Arabia, at 22 sites, was performed mainly using Malaise traps and sweep nets. In this study, 43 known species of Ephydridae were identified, 37 of them for the first time from southwestern Saudi Arabia and 16 from Arabian Peninsula. This brings the total number of Ephydridae species in Saudi Arabia to 49 (including previous records). There were a further four species, which could not be identified authoritatively to species level. A list of all species of Ephydridae recorded from Saudi Arabia is appended. Images of thirty-two species are presented. The species recorded in this study are more Palaearctic in origin as compared to other regions. Biological information (where known) and world-wide distribution are included. This study has added new records to the Saudi Arabian checklist of Ephydridae, which will provide the basis for systematic studies and fauna analyses of future work on the family. The need for further field and laboratory work and surveillance is highlighted.

 

References

  1. Abdul-Rassoul, M.S. (1976) Checklist of Iraq Natural History Museum Insects Collection. Publication 30. University of Baghdad, Baghdad, 41 pp.

    Abu-Zoherah, R., Al-Taher, K. & Tilkian, S. (1993) List of Insects Recorded from Saudi Arabia. Ministry of Agriculture and Water, National Agriculture and Water Research Centre, Riyadh, 394 pp.

    Adams, C.F. (1905) Diptera africana, I. The Kansas University Science Bulletin, 3, 149–208.

    https://doi.org/10.5962/bhl.part.8100

    Al-Ahmadi, A.Z. & Salem, M.M. (1999) Entomofauna of Saudi Arabia, Part 1: Checklist of Insects. Academic Publishing & Press, King Saud University, Riyadh, 240 pp.

    Al-Houty, W. (1989) Insect Fauna of Kuwait. Fahad Al-Marzouk, Printing & Publishing Establishment, Kuwait, 189 pp.

    Bächli, G. (1990) Type specimens of Drosophilidae (Diptera) described by Linnaeus, Fallén, Wahlberg and Zetterstedt. Revue suisse de zoologie, 97 (4), 843–849.

    https://doi.org/10.5962/bhl.part.79764

    Balachowsky, A. & Mesnil, L. (1935) Les Insectes Nuisibles aux Plantes Cultivees, Vol. I. Etablissements Busson, Paris, 1137 pp.

    Becker, T. (1896) Dipterologische Studien IV. Ephydridae. Berliner Entomologische Zeitschrift, 41, 91–276.

    https://doi.org/10.1002/mmnd.48018960505

    Becker, T. (1902) Die Meigen’schen Typen der sogen. Muscidae acalypterae (Muscaria holometopa) in Paris und Wien. Zeitschrift für systematische Hymenopterologie und Dipterologie, 2, 209–256 + 289–349.

    Becker, T. (1903) Aegyptische Dipteren Gesammelt und Beschrieben (Fortsetzung und Schluss). Mitteilungen aus dem Zoologischen Museum in Berlin, 2 (1), 67–195.

    Becker, T. (1905) Ephydridae. In: Becker, T., Bezzi, M., Kertesz, K. & Stein, P. (Eds.), Katalog der paläarktischen Dipteren. Vol. 4. G. Wessele’nyi in Hodmezovasarhely, Budapest, pp. 185–215.

    Becker, T. (1907) Die Ergebnisse meiner dipterologischen fruhjahrsreise nach Algier und Tunis. 1906. (concl.). Zeitschrift für Systematische Hymenopterologie und Dipterologie, 7, 369–407.

    https://doi.org/10.5962/bhl.title.9280

    Becker, T. (1910) Dipteren aus Südarabien und von der Insel Sokotra. Denkschrift der Akademie der Wissenschaften, Wien, 71, 131–160.

    https://doi.org/10.5962/bhl.title.9285

    Becker, T. (1922) Wissenschaftliche Ergebnisse der mit unterstutzung der Akademie der Wissenschaften in Wien aus der Erbschaft Treitl von F. Werner unternommen zoologischen Expedition nach dem anglo-agyptischen Sudan (Kordofan) 1914. VI. Diptera. Denkschriften der Akademie der Wissenschaften in Wien Mathematisch-Naturwissenschaftliche Klasse, 98 (1923), 57–82.

    Becker, T. (1924) H. Sauters Formosa Ausbeute. Ephydridae (Diptera). Entomologische Mitteilungen, 13, 89–93.

    Becker, T. (1926) 56a Ephydridae und 56b Canaceidae. In: Lindner, E. (Ed.), Die Fliegen der Palaearktischen Region. Band VI. 1. E. Schweizerbart’sche Verlag, Stuttgart, pp. 1–115.

    Becker, T. & Stein, P. (1912) Persische Dipteren von den Expeditionen des Herrn N. Zarudny 1898 und 1901. Annuaire du Musée Zoologique de l’Académie Impériale des Sciences de St.-Pétersbourg, 18, 503–654.

    Bergenstamm, J. von (1864) Metamorphose von Discomyza incurva Fall. Verhandlungen der Zoologisch-botanischen Gesellschaft in Wien, 14, 713–716.

    Beschovski, V. (1966) Allotrichoma valkanovi n. sp.—eine neue Art von Diptera (Ephydridae) aus Bulgarien. Comptes rendus de l’Académie bulgare des Sciences, 19, 851–853.

    Beyer, A. (1939) Morphologische, ökologische und physiologische Studien an den Larven der Fliegen: Ephydra riparia Fallèn, E. micans Haliday und Caenia fumosa Stenhammar. Kieler Meeresforschungen, 3, 265–320.

    Bezzi, M. (1895) Eine neue Art der Dipterengattung Psilopa Fallèn. Wiener Entomologische Zeitung, 14, 137–138.

    Bezzi, M. (1908) Ditteri Eritrei raccolti dal Dot. Adrani e Prof. Tellini. Parte Seconda (1). Diptera Cyclorrhapha. Bullettino della Società Entomologica Italiana, 39, 3–199.

    Brocher, F. (1910) Observations biologiques sur quelques Diptères et Hymenopteres dits “aquatiques”. Annales de Biologie lacustre, 4, 170–186.

    Canzoneri, S. (1986) Nuovi dati sugli Ephydridae (Diptera) della Sierra Leone, Accademia Nazionale dei Lincei. Problemi Attuali di Scienza e di Cultura, Sezione: Missioni ed Esplorazioni, X, 260, 67–75.

    Canzoneri, S. (1987) Sugli Ephydridae e Canacidae del Sudan (Diptera, Cyclorrhapha). Bollettino del Museo civico di Storia naturale di Venezia, 37 (1986), 79–97.

    Canzoneri, S. (1988) Una nuova specie di Allotrichoma dell’Etiopia (Diptera, Ephydridae). Società Veneziana di Scienze Naturali, Lavori, 13, 3–4.

    Canzoneri, S. & Meneghini, D. (1969) Sugli Ephydridae e Canaceidae della fauna etiopica. Bolletino del museo Civico di Storia Naturale di Venezia, 19, 101–185.

    Canzoneri, S. & Meneghini, D. (1983) Ephydridae e Canaceidae. In: Fauna d’Italia. Vol. XX. Edizioni Calderini, Bologna, xi + 337 pp.

    Clausen, P.J. (1977) A revision of the Nearctic, Neotropical, and Palearctic species of the genus Ochthera, including one Ethiopian species, and with one new species from India. Transactions of the American Entomological Society, 103, 451–529.

    Clausen, P.J. (1979) Lectotype designation of Ochthera schembrii Rondani (Diptera: Ephydridae) and resulting synonomies. Entomological News, 90, 141–142.

    Coe, R.L. (1945) 56. Ephydridae. In: Kloet, G.S. & Hincks, W.D. (Eds.), A Check List of British Insects, London, 1945, pp. 395–397.

    Cogan, B.H. (1968) A revision of the Ethiopian species of the tribe Notiphiline (Diptera: Ephydridae). Bulletin of the British Museum (Natural History), Entomology, 21 (6) 279–365.

    Cogan, B.H. (1980) 71. Family Ephydridae. In: Crosskey, R.W. (Ed.), Catalogue of the Diptera of the Afrotropical Region. British Museum (Natural History), London, pp. 655–669. [1437 pp.]

    Cogan, B.H. (1984) Family Ephydridae. In: Soós, A. & Papp, L. (Eds.), Catalogue of Palaearctic Diptera, Vol.10. Elsevier, Budapest, pp. 126–176. [402 pp.]

    Cogan, B.H. & Wirth, W.W. (1977) Family Ephydridae. In: Delfinado, M.D. & Hardy, D.E. (Eds.), A Catalogue of the Diptera of the Oriental Region. Vol. III. Suborder Cyclorrhapha (Excluding Division Aschiza). University Press of Hawaii, Honolulu, pp. 321–339.

    Collin, J.E. (1949) Results of the Armstrong College Expedition to Siwa Oasis (Libyan Desert), 1935, under the leadership of Prof. J. Omer-Cooper. Diptera Empididae, Dolichopodidae, Aschiza and Acalypterae. Bulletin de la Société Foudad I-er d’Entomologie, 33, 175–225.

    Coquillett, D.W. (1900a) New genera and species of Ephydridae. The Canadian Entomologist, 32 (2), 33–36.

    https://doi.org/10.4039/Ent3233-2

    Coquillett, D.W. (1900b) Report on a collection of dipterous insect from Puerto Rico. Proceedings of the United States National Museum, 22, 249–270.

    https://doi.org/10.5479/si.00963801.22-1198.249

    Coquillett, D.W. (1910) The type-species of the North American genera of Diptera. Proceedings of the United States National Museum, 37, 499–647.

    https://doi.org/10.5479/si.00963801.37-1719.499

    Cox, R.P.Jr. & Kadlec, J.A. (1995) Dynamics of potential waterfowl foods in Great Salt Lake marshes during summer. Wetlands, 15 (1), 1–8.

    https://doi.org/10.1007/BF03160674

    Cresson, E.T. (1916a) Studies in the American Ephydridae (Diptera) I: A revision of the species of the genus Paralimna. Transactions of the American Entomological Society, 42, 101–124.

    Cresson, E.T. (1916b) Description of new genera and species of the dipterous family Ephydridae. III. Entomological News, 27 (4), 147–152.

    Cresson, E.T. (1917) Description of new genera and species of the dipterous family Ephydridae IV. Entomological News, 28 (8), 340–341.

    Cresson, E.T. (1925) Studies in the dipterous family Ephydridae, excluding the North and South American faunas. Transactions of the American Entomological Society, 51, 227–258.

    Cresson, E.T. (1929) Studies in the dipterous family Ephydridae. Paper II. Transactions of the American Entomological Society, 55, 165–195.

    Cresson, E.T. (1930) Studies in the dipterous family Ephydridae III. Transactions of the American Entomological Society, 56, 93–131.

    Cresson, E.T. (1932) Studies in the dipterous family Ephydridae, paper IV. Transactions of the American Entomological Society, 58, 1–34.

    Cresson, E.T. (1933) New African species of the dipterous genus Paralimna (Ephydridae). Annals and Magazine of Natural History, Series 10, 11 (61), 24–33.

    https://doi.org/10.1080/00222933308673621

    Cresson, E.T. (1936) Description and notes on genera and species of the dipterous family Ephydridae. II. Transactions of the American Entomological Society, 62, 257–270.

    Cresson, E.T. (1939) Description of a new genus and ten new species of Ephydridae, with a discussion of the species of the genus Discomyza (Diptera). Notulae Natulae, The Academy of Natural Sciences of Philadelphia, 21, 1–12.

    Cresson, E.T. (1943a) Description of new genera and species of the dipterous family Ephydridae. Paper XIII. Notulae Naturae, 121, 1–4.

    Cresson, E.T. (1943b) The species of the tribe Ilytheini (Diptera: Ephydridae: Notiphilinae). Transactions of the American Entomological Society, 69, 1–16.

    Cresson, E.T. (1946) A systematic annotated arrangement of the genera and species of the Ethiopian Ephydridae (Diptera). I. The subfamily Psilopinae. Transactions of the American Entomological Society, 72, 241–264.

    Cresson, E.T. (1947) A systematic annotated arrangement of the genera and species of the Ethiopian Ephydridae (Diptera). II. The subfamily Notiphilinae. Transactions of the American Entomological Society, 73, 105–124.

    Curtis, J. (1832) British entomology, being illustrations and descriptions of the genera of insects found in Great Britain and Ireland: containing coloured figures from nature of the most rare and beautiful species, and in many instances of the plants upon which they are found. Vol. 9. Printed for the author, London, 50 pls. [pls. 384–433, 1823–1840]

    https://doi.org/10.5962/bhl.title.8148

    Curtis, J. (1837) A guide to an arrangement of British insects: being a catalogue of all the named species hitherto discovered in Great Britain and Ireland, Ed. 2. Printed for the author, published and sold by J. Pigot, London, 294 pp. [1837]

    https://doi.org/10.5962/bhl.title.46860

    Czerny, L. (1909) Spanische Dipteren. III Cyclorrhapha Schizophora Holometopa. Verhandlungen der Zoologisch-botanischen Gesellschaft in Wien, 59, 247–290.

    Dabbour, A.I. (1979a) Short note on dipterous flies in western and central regions of Saudi Arabia. Journal of Agriculture Research, Riyadh University, 4, 81–83.

    Dabbour, A.I. (1979b) Note on dipterous flies in western and central regions of Saudi Arabia. Journal of Faculty of Science, Riyadh University, 10, 117–119.

    Dabbour, A.I. & El-Dawy, M. (1981) Morphological and classification studies of some dipterous flies in the Kingdom of Saudi Arabia. Agriculture Research Center Bulletin. No. (10). Faculty of Agriculture, Riyadh University, Riyadh, 92 pp. [in Arabic]

    Dahl, R.G. (1959) Studies on Scandinavian Ephydridae (Diptera Brachycera). Opuscula Entomologica Supplementum, 15, 1–224.

    Dawah, H.A. & Abdullah, M.A. (2006) The Ephydridae (Diptera: Brachycera: Muscomorpha) of south-western Saudi Arabia. Fauna of Arabia, 21, 383–394.

    Deeming, J.C. (1995) Diptera (true flies) from the Kenfig National Nature Reserve, Glamorgan. National Museum of Wales, Entomology Series No. 4, 1–113.

    Deeming, J.C. (2002) Some species of the genus Hydrellia Robineau-Desvoidy (Diptera: Ephydridae) in Africa and Arabia with special reference to species of economic importance. Mitteilungen aus dem Museum fur Naturkunde in Berlin, Deutsche Entomologische Zeitschrift, 49, 47–59.

    https://doi.org/10.1002/mmnd.20020490105

    de Meijere, J.C.H.de. (1924) Studien über südostasiatische Dipteren XVI. Tijdschrift voor Entomologie, 67, 197–224.

    Deonier, D.L. (1971) A systematic and ecological study of Nearctic Hydrellia (Diptera: Ephydridae). Smithsonian Contributions to Zoology, 68, 1–147.

    https://doi.org/10.5479/si.00810282.68

    Dikow, T. & Mathis, W.N. (2002) A revision and phylogenetic study of Actocetor Becker (Diptera: Ephydridae). Proceedings of the Entomological Society of Washington, 104 (2), 249–290.

    Duda, O. (1942) Neue oder ungenügend bekannte Zweiflügler der paläarktischen Region aus meiner Sammlung 2: Fortsetzung. Deutsche Entomologische Zeitschrift, 1942 (1–4), 1–39.

    https://doi.org/10.1002/mmnd.48019420102

    Duponchel, A. [as (D)] (1845) Hydrellie. In: d’Orbigny C.V.D. (Ed.), Dictionnaire universel d’histoire naturelle résumant et complétant tous les faits présentés par les encyclopédies. Vol. 6. Bureau de principal des éditeurs, Paris, pp. 1–753.

    Edmiston, J.F. & Mathis, W.N. (2005) A revision of the New World species of shore-fly genus Nostima Coquillett (Diptera: Ephydridae). Smithsonian Contributions to Zoology, 623, 1–108.

    https://doi.org/10.5479/si.00810282.623

    El-Hawagry, M.S., Abdel-Dayem, M.S., Elgharbawy, A.A. & Al Dhafer, H.M. (2016) A preliminary account of the fly fauna in Jabal Shada al-A’la Nature Reserve, Saudi Arabia, with new records and biogeographical remarks (Diptera, Insecta). Zookeys, 636, 107–139.

    https://doi.org/10.3897/zookeys.636.9905

    El-Hawagry, M.S., Abdel-Dayem, M.S., El-Sonbati, S.A. & Al Dhafer, H.M. (2017) A preliminary account of the fly fauna in Garf Raydah Nature Reserve, Kingdom of Saudi Arabia, with new records and biogeographical remarks (Diptera: Insecta). Journal of Natural History, 51 (25–26), 1499–1530.

    https://doi.org/10.1080/00222933.2017.1347299

    El-Hawagry, M.S., Abdel-Dayem, M.S. & Al Dhafer, H.M. (2018a) A contribution to the knowledge of fly fauna in the Kingdom of Saudi Arabia: new country records and an account of flies identified from Rawdhats, Riyadh Region, with biogeographical remarks (Insecta: Diptera). Journal of Natural History, 52 (21–22), 1377–1393.

    https://doi.org/10.1080/00222933.2018.1456575

    El-Hawagry, M.S., Al Dhafer, H.M. & Abdel-Dayem, M.S. (2019) On the fly fauna of the central region of the Kingdom of Saudi Arabia: new country records from Riyadh Region, with a list of associated fly species and zoogeographical remarks (Insecta: Diptera). Journal of Natural History, 53 (1–2), 17–43.

    https://doi.org/10.1080/00222933.2019.1568601

    El-Hawagry, M.S., Zatwarnicki, T. & Ebrahim, A.M. (2018b) Catalogue of the Egyptian Ephydroidea (Diptera: Schizophora: Acalyptratae). Zootaxa, 4444 (3), 201–246.

    https://doi.org/10.11646/zootaxa.4444.3.1

    Enderlein, G. (1922) Eine neue Ephydridae aus dem Banat. Zoologischer Anzeiger, 55, 129–130.

    Enderlein, G. (1934) Heringiinae, eine neue minirende Chloropiden Unterfamilie. Zoologischer Anzeiger, 105, 191–195.

    Fallèn, D.F. (1810) Specimen Entomologicum novam Diptera disponendi methodum exhibens. Litteris Berlingianis, Lundae, 26 pp.

    Fallén, C.F. (1813) Beskrifning Öfver någr i Sverige Funna Vattenflugor (Hydromyzidae). Kungliga Svenska Vetenskaps Akademiens Hindlinger, 3, 1813, 240–257.

    Fallén, C.F. (1823) Hydromyzides Sveciae. Berlingianis, Lundae (Lund), 12 pp.

    Fan C., Qi, G., Li, M., Yang, R. & Xia, P. (1983) Two new species of the genus Hydrellia on rice, wheat or barley (Diptera: Ephydridae). Entomotaxonomia, 5 (1), 7–12.

    Ferrar, P. (1987) A guide to the breeding habits and immature stages of Diptera Cyclorrhapha. Entomonograph 8. Part 1 & 2. E.J. Brill/Scandinavian Science Press, Leiden and Copenhagen, 478 pp. [Lyneborg, L. (Ed.)]

    Foote, B.A. (1995) Biology of shore flies. Annual Review of Entomology, 40, 417–442.

    https://doi.org/10.1146/annurev.en.40.010195.002221

    Foote, B.A. & Eastin, W.C. (1974) Biology and immature stages of Discocerina obscurella (Diptera: Ephydridae). Proceedings of the Entomological Society Washington, 76 (4), 401–408.

    Franz, H. (1989) Die Nordost-Alpen im Spiegel Ihrer Landtierwelt. Band VI/2. Diptera Cyclorapha. Universitätsverlag Wagner, Innsbruck, 445 pp.

    Frey, R. (1958) Zur Kenntnis der Brachycera p. p. der Kapverdischen Inseln. Commentationes Biologicae, 18 (4), 1–61.

    Gatt, P. & Ebejer, M.J. (2003) The Ephydridae (Diptera: Brachycera, Muscomorpha) of the Maltese Islands. Studia Dipterologica, 10, 199–214.

    Giordani Soika, A.G. (1956a) Contributions a l’etude de la fauna entomologique du Ruanda-Urundi (Mission P. Basilewsky 1953) CVII Diptera Ephydridae. Annales du Musée Royal du Congo Belge, Tervuren, Belgique, Series 8, Sciences Zoologiques, 51, 490–505.

    Giordani Soika, A.G. (1956b) Diagnosi preliminari di nuovi Ephydridae e Canaceidae della Regione Etiopica e del Madagascar (Diptera). Bollettino del Museo civico di Storia naturale di Venezia, 9, 123–130.

    Giordani Soika, A.G. (1956c) Viaggi di A. Giordani Soika nel Sahara IX. Contibuto allo studio del popolamento del Sahara Diptera Ephydridae. Bollettino del Museo civico di Storia naturale di Venezia, 9, 95–114.

    Greathead, D.J. (1963) A review of the insect enemies of Acridoidea (Orthoptera). Transactions of the Royal Entomological Society of London, 114, 437–517.

    https://doi.org/10.1111/j.1365-2311.1963.tb01073.x

    Grigarick, A.A. (1959) Bionomic of the rice leaf miner, Hydrellia griseola (Fallén), in California (Diptera: Ephydridae). Hilgardia, 29, 1–80.

    https://doi.org/10.3733/hilg.v29n01p001

    Haliday, A.H. (1833) Catalogue of Diptera occurring about Holywood in Downshire. The Entomologist Magazine, 1 (2), 147–180.

    Haliday, A.H. (1839) Remarks on the generic distribution of the British Hydromyzidae. Annals of Natural History, 3, 217–224 + 401–411.

    https://doi.org/10.1080/03745483909443252

    Haliday, A.H. (1855) Descriptions of insects figured, and references to plates illustrating the notes on Kerry insects. Dublin University Zoological Associations Proceedings for June 2nd 1855. Natural History Review, 2, 115–124.

    Harten, van.A. & Wagener, B. (1994) Terrestrial arthropods of the Republic of Yemen. A check-list. Yemeni-German Plant Protection, Sana’a, Yemen, ii + 147pp.

    Hendel, F. (1913) H. Sauter’s Formosa-Ausbeute. Acalyptrate Musciden (Diptera). II. Supplementa Entomologica, 2, 77–112.

    Hendel, F. (1917) Beiträge zur Kenntniss der acalyptraten Musciden. Deutsche Entomologische Zeitschrift, 1917 (6), 33–47.

    Hendel, F. (1931) Neue aegyptische Dipteren aus der Gruppe der acalyptraten Musciden, gesammelt van Prof. Efflatoun Bey. Bulletin of the Entomological Society of Egypt, New Series, 15, 59–73.

    Hendel, F. (1933) Neue Acalyptratae Musciden aus der Paläarktischen Region (Dipt.). Deutsche Entomologische Zeitschrift, 1933, 39–56.

    https://doi.org/10.1002/mmnd.48019330103

    Hennig, W. (1941) Verzeichnis der Dipteren von Formosa. Entomologische Beihefte aus Berlin-Dahlem, 8, 1–239.

    Hering, E.M. (1950) Beitrag zur Kenntnis der Gattung Hydrellia (Diptera: Ephydridae). Archiv für Hydrobiologie, 43, 234–240.

    Hesler, L.S. (1995) Bibliography on Hydrellia griseola Fallén (Diptera: Ephydridae) and review of its biology and pest status. Insecta Mundi, 9 (1–2), 25–35.

    Hollmann-Schirrmacher, V. (1998) Phylogeny of the subfamily Ilytheinae (Diptera: Ephydridae) with special reference to the genus Philygria. Studia Dipterologica, Supplement 5, 1–144.

    Katzenberger, J., Langbehn, T. & Zacharias, D. (2013) Erstnachweis von Hydrellia tarsata (Diptera: Ephydridae) fur Bremen in Bluten von Stratiotes aloides. Abhandlungen des Naturwissenschaftlichen Vereins zu Bremen, 47, 193–195.

    Keiper, J.B., Jiannino, J., Sanford, M. & Walton, W.E. (2001) Biology and immature stages of Typopsilopa nigra (Williston) (Diptera: Ephydridae), a secondary consumer of damaged stems of wetland monocots. Proceedings of the Entomological Society of Washington, 103, 89–97.

    Keiper, J.B. & Walton, W.E. (2000) Biology and mature stages of Brachydeutera sturtevanti (Diptera: Ephydridae), a hyponeustic generalist. Annals of Entomological Society of America, 93 (3), 468–475.

    https://doi.org/10.1603/0013-8746(2000)093[0468:BAISOB]2.0.CO;2

    Keiper, J.B., Walton, W.E. & Foote, B.A. (2002) Biology and ecology of higher Diptera from freshwater wetlands. Annual Revue of Entomology, 47, 207–232.

    https://doi.org/10.1146/annurev.ento.47.091201.145159

    Krivosheina, M. & Zatwarnicki, T. (1997) Some notes on the Old world Allotrichoma (Diptera: Ephydridae) with description of three new species. Polskie Pismo Entomologiczne, 66, 391–310.

    Lamb, C.G. (1912) The Percy Sladen Trust expedition to the Indian ocean in 1905, under the leadership of Mr J. Stanley Gardiner, M.A. Vol. 4. No. XlX. Diptera: Lonchaeidae, Sapromyzidae, Ephydridae, Chloropidae, Agromyzidae. Transactions of Linnean Society, London, 2 (Zoology), 15, 303–348.

    https://doi.org/10.1111/j.1096-3642.1912.tb00104.x

    Latreille, P.A. (1802) Histoire naturelle, generale et particuliere, des Crustacés et des Insectes, Tome troisième. In: Sonnini, C.S. (Ed.), Histoire naturelle par Buffon, Paris, 1802, pp. 1–468.

    Latreille, P.A. (1804) Tableu methodique des Insectes. In: Tableaux methodiques d’histoire naturelle, Paris, 1804, pp. 129–200.

    Latreille, P.A. (1810) Considérations générales sur l’orde naturel des animaux composant les classes des Crustaces, des Arachnides et des Insectes. Chez F. Schoell, Paris, 444 pp.

    https://doi.org/10.5962/bhl.title.39620

    Laurence, B.R. (1952) Observations on Hydrellia (Hydropota) griseola (Fallèn) (Diptera, Ephydridae). The Entomologist’s Monthly Magazine, 88, 31–33.

    Lilljeborg, W. (1861) Om en flugas (Hydrellia griseola) harjningar å kornfälten i Östra Skåne, Blekinge och Södra dalarna af Kalmar län, under sommaren 1860. Tidskrift för Landtmanna och Kommunalekonomien, 1861, 205–215.

    Loew, H. (1847) Dipterologisches. Stettiner Entomologische Zeitung, 8 (12), 368–376.

    Loew, H. (1856) Neue Beiträge zur Kenntniss der Dipteren. Virter Beitrag, E. S. Mittler & Sohn, Berlin, 57 pp.

    Loew, H. (1860) Neue Beiträge zur Kenntnis der Dipteren. Siebenter Beitrag. Programm Konigligen Realschule zu Meseritz, 1860, 1–46.

    Loew, H. (1862a) Monographs of the Diptera of North America. Part 1. Smithsonian Miscellaneous Collections, 6 (141), 1–221.

    Loew, H. (1862b) Bidrag till kannedomen om Afrikas Diptera. Ofversigt af Kongl Vetenskaps-Akademiens Forhandlinger Stockholm, 19, 3–14.

    Macquart, M.J. (1835) Histoire Naturelle des Insectes. Diptères, Tome deuxième. In: Roret, N.E. (Ed.), Diptera. Vol. 2. Collection des Suites a Buffon, Paris, pp. 1–703, 12 pls.

    https://doi.org/10.5962/bhl.title.14274

    Macquart, M.J. (1843) Diptères exotiques nouveaux ou peu connus. Mémoires de la Société Royale des Sciences, de I’ Agriculture et des Arts á Lille, 1842, 162–460.

    Macquart, M.J. (1851) Diptères exotiques nouveaux ou peu connus. Suite du 4e supplément publié dansles mémoires de 1849 (concl.). Mémoires de la Société royale des Sciences, de l’Agriculture et des Arts de Lille, 1850, 134–294.

    Marshall, S.A. (2012) Flies the Natural History & Diversity of Diptera. Firefly Books Ltd, Ontario, 616 pp.

    Mathis, W.N. (1978) New synonymy in Ephydridae (Diptera). Entomological News, 89 (2–3), 105–106.

    Mathis, W.N. (1984) A revision of the genus Asmeringa Becker (Diptera: Ephydridae), Israel Journal of Entomology, 17 (1983), 67–79.

    Mathis, W.N. (1986) A revision of the subgenus Eremotrichoma Soika of the shore fly genus Allotrichoma Becker (Diptera, Ephydridae). Israel Journal of Entomology, 19 (1985), 127–139.

    Mathis, W.N. (1991) Studies of Gymnomyzinae (Diptera: Ephydridae), III: A Revision of the Shore Fly Subgenus Pseudohecamede Hendel of the Genus Allotrichoma Becker. Smithsonian Contributions to Zoology, 522, i–iii + 1–28.

    https://doi.org/10.5479/si.00810282.522

    Mathis, W.N. (1993) Studies of Gymnomyzinae (Diptera: Ephydridae), IV: A revision of the shore-fly genus Hecamede Haliday. Smithsonian Contributions to Zoology, 541, 1–46.

    https://doi.org/10.5479/si.00810282.541

    Mathis, W. (2010) Ephydridae (Shore Flies). In: Brown, B.V., Borkent, A., Cumming, J.M., Wood, D.M., Woodley, N.E. & Zumbado, M.A. (Eds.), Manual of Central American Diptera. Vol. 1. NRC Research Press, Ottawa, pp. 1211–1233.

    Mathis, W.N. & Deeming, J.C. (1987) A revision of the shore fly genus Elephantinosoma Becker (Diptera: Ephydridae). Proceedings of the Entomological Society of Washington, 89 (3), 562–570.

    Mathis, W.N. & Ghorpadé, K.D. (1985) Studies of Parydrinae (Diptera: Ephydridae), I: A review of the genus Brachydeutera Loew from the Oriental, Australian, and Oceanian Region. Smithsonian Contributions to Zoology, 406, i–iii + 1–25.

    https://doi.org/10.5479/si.00810282.406

    Mathis, W.N. & Steiner, Jr.W.E. (1986) An adventive species of Brachydeutera Loew in North America (Diptera: Ephydridae). Journal of the New York Entomological Society, 94 (1), 56–61.

    Mathis, W.N. & Winkler, I.S. (2003) A review of the neotropical species of Brachydeutera Loew (Diptera: Ephydridae). Proceedings of the Entomological Society of Washington, 105 (2), 407–424.

    Mathis, W.N. & Zatwarnicki, T. (1990) A review of the western palearctic species of Athyroglossa (Diptera: Ephydridae). Transactions of the American Entomological Society, 116 (1), 103–133.

    Mathis, W.N. & Zatwarnicki, T. (1993) Revision of the shore-fly genus Chlorichaeta Becker (Diptera: Ephydridae). Tijdschrift voor Entomologie, 136 (1), 55–70.

    Mathis, W.N. & Zatwarnicki, T. (1995) A world catalog of the shore flies (Diptera: Ephydridae). Memoirs on Entomology, International. Vol. 4.Associated Publishers, Grainesville, Florida, 423 pp.

    Mathis, W.N. & Zatwarnicki, T. (1998) 3.49. Family Ephydridae. In: Papp, L. & Darvas. B. (Eds.), Contributions to a Manual of Palaearctic Diptera (with special reference to flies of economic importance). Vol. 3. Higher Brachycera. Science Herald, Budapest, pp. 537–570.

    Mathis, W.N. & Zatwarnicki, T. (2002) A Phylogenetic Study of the Tribe Dryxini Zatwarnicki (Diptera: Ephydridae). Smithsonian Contributions to Zoology, 617, i–iv + 1–101, 154 figs, 2 tabs.

    https://doi.org/10.5479/si.00810282.617

    Mathis, W.N. & Zatwarnicki, T. (2003) Shore flies of the Republic of Seychelles (Diptera: Ephydridae). Annales Zoologici, Warszawa, 53 (4), 585–650.

    Mathis, W.N. & Zatwarnicki, T. (2010) New species and other taxonomic modifications for shore flies from the Delmarva States (Diptera: Ephydridae). Proceedings of the Entomological Society of Washington, 112 (1), 97–128.

    https://doi.org/10.4289/0013-8797-112.1.97

    Mathis, W.N., Zatwarnicki, T., Stuke, J.H. & Deeming, J.C. (2017) A conspectus on shore-flies from the United Arab Emirates. In: van Harten, A. (Ed.), Arthropod Fauna of the United Arab Emirates. Vol. 6. Dar Al-Ummah Printing, Publishing, Distribution & Advertising, Abu Dhabi, pp. 63661. [754 pp.]

    Meigen, J.W. (1830) Systematische Beschreibung der bekannten europaischen zweiflügeligen Insekten. Vol. 6. Schulz, Hamm, 6, xi + 401 pp.

    Meigen, J.W. (1838) Systematische Beschreibungen der bekannten europaischen zweiflügeligen Insekten. 7 (Supplementary Volume). Bei Friedrich Wilhelm Forstmann, Gedrukt bei Beaufort Sohn, Aachen, XII + 434 pp.

    Merritt, R.W., Schlinger, E.I. & Webb, D.W. (1996) Aquatic Diptera. Part Two. Pupae and adults of aquatic Diptera. In: Merritt, R.W. & Cummins, K.W. (Eds.), An Introduction to the Aquatic Insects of North America. 3rd Edition. Kendall-Hunt Publ. Co., Dubuque, Iowa, pp. 515–554.

    Miyagi, I. (1977) Ephydridae (Insecta: Diptera). Fauna Japonica, 1977, 1–113 pp., 49 pls.

    Müller, G.W. (1922) Insektenlarven an Wurzeln von Wasserpflanzen. Mittheilungen aus dem Naturwissenschaftlichen Verein für Neu-Vorpommern und Rügen in Greifswald, 48/49, 30–50.

    Nasr, S.A., Isa, A.L., Kira, T. & Al-Tantawy, A.M. (1970) Effect of the blood worms (Chironomus sp.) and the rice fly (Ephydra macellaria Egger) on rice (Diptera: Chironomidae and Ephydridae). Bulletin of the Entomological Society of Egypt, 54, 203–211.

    Nye, I.W.B. (1958) The external morphology of some of the Dipterous larvae living in the Graminae of Britain. The Transactions of the Entomological Society of London, 110, 411–487.

    https://doi.org/10.1111/j.1365-2311.1958.tb00380.x

    Papp, L. (1974) New or little known species of Ephydridae and Sphaeroceridae from Hungary (Diptera: Acalyptratae). Acta Zoologica Hungarici, 20 (3–4), 403–409.

    Papp, L. (1979) A contribution to the revision of the Palearctic Ephydridae (Diptera). Folia Entomologica Hungarica, Series Nova, 32 (1), 97–104.

    Risbec, J. (1951) Les insectes nuisibles au riz de Camargue au debut de son developpment. Revue de Pathologie Végétale et d’Entomologie Agricole de France, 30, 211–227.

    Robineau-Desvoidy, J.B. (1830) Essai sur les Myodaires. Mémoires présentés par divers savans à l’Académie royale des science de l’Institut de France, et imprimés par son ordre: Sciences Mathématiques et Physiques, Paris, 2, 1–813.

    Rondani, C. (1847) Nota septima ad inserviendum diperologiae italicae. Nova species generis Ochtherae Latr., descripta a Camillo Rondani. Annales de Societè Entomologique France, 2 (5), Bull, XIX–XXXI.

    Rondani, C. (1856) Dipterologiae Italicae prodomus. Vol 1. Genera Italica ordinis dipterorum ordinatim disposita et distincta et in familias et stirpes aggregata, Parmae, 1856, 1–228.

    Scheiring, J.F. & Foote, B.A. (1973) Habitat Distribution of the Shore Flies of Northeastern Ohio (Diptera: Ephydridae). The Ohio Journal of Science, 73 (3), 152–166, 10 figs.

    Schiner, J.R. (1864) Catalogus systematicus Dipterorum Europae. impensis Societatis zoologico-botanicae editus, Vindobonae, 115 pp.

    https://doi.org/10.5962/bhl.title.39977

    Séguy, E. (1934) Diptères (Brachyceres) (Muscidae Acalypterae et Scatophagidae). Faune de France. Vol. 28. Lechevalier et fils, Paris, 832 pp.

    Siepel, H., Bink, F.A., Broekhuizen, S., Stumpel, A.H.P. & van Wingerden, W.K.R.E. (1993) De internationale betekenis van Nederland voor de fauna. 1. De terrestrische fauna. IBN-Rapport 012. IBN-DLO, Wageningen, 234 pp.

    Simpson, K.W. (1975) Biology and immature stages of three species of Nearctic Ochthera (Diptera: Ephydridae). Proceedings of the Entomological Society of Washington, 77, 129–155.

    Steinly, B.A. (2004) Primary consumer and detritivore communities (Diptera: Ephydridae) in newly restored and constructed wetlands. Proceedings of the Entomological Society of Washington, 106 (2), 460–471.

    Stenhammar, C. (1844) Forsok till Gruppering och Revision af de Svenska Ephydrinae. Kungliga Svenska Vetenskapsakademiens Hindlinger, 1843 (3), 75–272.

    https://doi.org/10.5962/bhl.title.16236

    Steyskal, G.C. (1968) Notes and descriptions of Egyptian acalyptrate Diptera (Ephydridae, Milichiidae, Otitidae). Bulletin of the Entomological Society of Egypt, 50 (1966), 109–125.

    Steyskal, G.C. & El-Bialy, S. (1967) A list of Egyptian Diptera with a bibliography and key to families. Technical Bulletin of the Ministry of Agriculture, UAR, 3, 1–87.

    Stower, W.J., Popov, G.B. & Greathead, D.J. (1958) Oviposition behaviour and egg mortality of the desert locust (Schistocera gregaria Forskål) on the coast of Eritrea. Anti-locust Bulletin, 30, 1–33.

    Stuke, J.-H. (2011) Neue Uferfliegen-Funde (Diptera: Ephydridae) von Madeira-Archipel, mit der Beschreibung einer neuen Art. Studia Dipterologica, 17 (1/2), 211–221. [2010]

    Stuke, J.-H. (2012) Funde von Ephydridae (Diptera) aus Jordanien, mit der Beschreibung von fünf neuen Arten. Studia dipterologica, 18 (1–2), 197–224.

    Venkatesh, M.G. (1976) Some observations on the biology of Brachydeutera longipes Hendel (Diptera: Ephydridae) from West Bengal. Journal of the Bombay Natural History Society, 57, 345–346.

    Walker, F. (1856) Insecta Britannica. Diptera. Vol. 3. John Edward Taylor, London, XXIV + 352 pp.

    Walker, D.H. & Pittaway, A.R. (1987) Insects of Eastern Arabia. Textbook, published by Macmillan Publishers Ltd., New York, 175 pp.

    Waltl, J. (1837) Neue Gattungen von Mücken bey Passau, Isis, Oken, 4, 279–283.

    Westwood, J.O. (1840) Order XIII. Diptera Aristotle (Antilata Fabricius. Halteriptera Clairv.). pp. 125–154. In: An introduction to the modern classification of insects; Founded on the natural habits and corresponding organisation of the different families. Synopsis of the genera of British Insects. Longman, Orme, Brown, Green and Longmans, London, 158 pp.

    Order XIII. Diptera Aristotle (Antilata Fabricius. Halteriptera Clairv.). In: An introduction to the modern classification of insects; Founded on the natural habits and corresponding organisation of the different families. Synopsis of the genera of British Insects. Longman, Orme, Brown, Green and Longmans, London, pp. 125–154. [158 pp]

    Wiedemann, C.R.W. (1824) Munus rectoris in Academia Christiana Albertina aditurus Analecta entomologica ex Museo Regio Havniensi maxime congresta profert iconibusque illustrat. Kiliae, eregio typoguapheo scholarum,1824, Kiliae, 60 pp.

    https://doi.org/10.5962/bhl.title.77322

    Wiedemann, C.R.W. (1830) Aüssereuropäische zweiflügelige Insekten (concl.). Vol. 2. Schulz, Hamm, xii + 684 pp.

    Willcocks, F.C. (1925) The insect and related pests of Egypt. II. Insects and mites feeding on graminous crops and products in the field, granary and mill. Sultanic Agricultural Society, Technical Section, Procaccia, Lencioni and Co., Cairo, VIII + 418 pp.

    Wirth, W.W. (1964a) A revision of the shore flies of the genus Brachydeutera Loew (Diptera: Ephydridae). Annals of the Entomological Society of America, 57 (1), 3–12.

    https://doi.org/10.1093/aesa/57.1.3

    Wirth, W.W. (1964b) A new species and new records of African Ephydridae (Dipt.). Ergebnisse der Forschungsreise Lindner 1958/59. Nr. 18. Stuttgarter Beiträge zur Naturkunde, 133, 1–3.

    Wirth, W.W. (1975) A revision of the brine flies of the genus Ephydra of the Old World (Diptera: Ephydridae). Entomologica Scandinavica, 6, 11–44.

    https://doi.org/10.1163/187631275X00325

    Zangheri, S. (1956) Un dittero minatore del riso nel Basso Ferrarese (Hydrellia griseola Fallèn, Dipt., Ephydridae). Bullettino della Società Entomologica Italiana, 86, 12–16.

    Zatwarnicki, T. (1987) New synonyms and records of Palearctic Scatophila (Diptera, Ephydridae). Polskie Pismo Entomologiczne, 57 (2), 277–298.

    Zatwarnicki, T. (1988) Materials to the knowledge of the genus Hydrellia Robineau-Desvoidy (Ephydridae: Diptera). Polskie Pismo Entomologiczne, 58, 587–634.

    Zatwarnicki, T. (1997) Diptera Ephydridae, shore flies. In: Nilsson, A. (Ed.), The aquatic insects of north Europe. Vol 2. Apollo Books, Stenstrup, pp. 383–399.

    Zatwarnicki, T. (2002) A revision of the genus Orasiopa Zatwarnicki et Mathis (Diptera: Ephydridae). Annales Zoologici, Warszawa, 52 (2), 297–317.

    Zatwarnicki, T. (2018) Solving the puzzle of taxonomic position of the petroleum fly by resurrection of Diasemocera Bezzi from Psilopa Fallén (Diptera: Ephydridae) with proposed specific and generic synonymies. Annales Zoologici, 68 (3), 527–552.

    https://doi.org/10.3161/00034541ANZ2018.68.3.012

    Zatwarnicki, T., Cielniak, M. & Pochrząst, K. (2016a) A revised phylogeny of the tribe Discocerinini (Diptera: Ephydridae) with an emphasis on structures of the proboscis. Annales Zoologici, 66 (1), 1–34.

    https://doi.org/10.3161/00034541ANZ2016.66.1.001

    Zatwarnicki, T., Cielniak, M. & Pochrząst, K. (2016b) A Revision of the Old World species of the genus Leptopsilopa Cresson (Diptera: Ephydridae). Zootaxa, 4093 (3), 343–362.

    https://doi.org/10.11646/zootaxa.4093.3.3

    Zatwarnicki, T. & Mathis, W.N. (2001) A Generic Classification of the tribe Discocerinini (Diptera: Ephydridae). Annales Zoologici, 51 (1), 5–51.

    Zatwarnicki, T. & Mathis, W.N. (2007) A revision of the Palearctic species of the shore-fly genus Discomyza Meigen (Diptera: Ephydridae). Insect Systematics and Evolution, 38, 241–266.

    https://doi.org/10.1163/187631207788754448

    Zetterstedt, J.W. (1838) Insecta Lapponica. Voss, Lipsiae, VI + 1140 pp.

    Zetterstedt, J.W. (1846) Diptera Scandinaviae. Disposita et Descripta. Tomus Quintus, Lundae, pp. 1739–2162.