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Type: Article
Published: 2019-06-28
Page range: 289–295
Abstract views: 50
PDF downloaded: 3

A new pseudoscorpion genus from western Georgia (Pseudoscorpiones: Neobisiidae: Cornuroncus n. gen.), with a key to all Neobisiinae genera

Entomology Department, Faculty of Agriculture and Natural Resources, Islamic Azad University, Arak branch, Arak, Iran.
Juan A. ZARAGOZA, Departamento de Ecología, Facultad de Ciencias, Universidad de Alicante, Alicante, Spain.
Levan MUMLADZE, Bioodiversity Research Center, Institute of Ecology and Institute of Zoology, Ilia State University, Tbilisi, Georgia.
Pseudoscorpiones Arachnida Caucasus pseudoscorpions new species taxonomy


A new genus and species of epigean pseudoscorpions belonging to the subfamily Neobisiinae Chamberlin, 1930, is described from western Georgia and its diagnostic characters are illustrated. Cornuroncus n. gen. resembles the genus Roncus L. Koch, 1873 in several characters but differs by having a short dorso-distal spine on tarsus IV, and the presence of a ventral tooth on one of the claws of tarsus IV. An identification key for all valid Neobisiinae genera is provided.



  1. Chamberlin, J.C. (1930) A synoptic classification of the false scorpions or chela-spinners, with a report on a cosmopolitan collection of the same. Part II. The Diplosphyronida (Arachnida-Chelonethida). Annals and Magazine of Natural History, Series 10, 5 (30), 585–620.

    Chamberlin, J.C. (1931) The arachnid order Chelonethida. Stanford University Publications, Biological Sciences, 7 (1), 1–284.

    Ćurčić, B.P.M. (1975) Balkanoroncus (Arachnida, Pseudoscorpiones, Neobisiidae), a new genus of pseudoscorpions based on Roncus bureschi Hadži, 1939. Glasnik Muzeja Srpske Zemlje, Beograd, (B), 30, 143–145.

    Ćurčić, B.P.M. (1987) Insulocreagris, a new genus of pseudoscorpions from the Balkan Peninsula (Pseudoscorpiones, Neobisiidae). Revue Arachnologique, 7, 47–57.

    Gardini, G. (1982) Balkanoroncus baldensis n. sp. delle Prealpi Venete (Pseudoscorpionida, Neobisiidae) (Pseudoscorpioni d’Italia XIV). Bollettino del Museo Civico di Storia Naturale, Verona, 9, 161–173.

    Gardini, G. (1998) Il genere Acanthocreagris in Italia (Pseudoscorpionida, Neobisiidae). Fragmenta Entomologica, 30, 1–73.

    Harvey, M.S. (1992) The phylogeny and classification of the Pseudoscorpionida (Chelicerata: Arachnida). Invertebrate Taxonomy, 6, 1373–1435.

    Harvey, M.S. (2013) Pseudoscorpions of the World. Version 3.0. Western Australian Museum, Perth. Available from: (accessed 17 January 2019)

    Harvey, M.S. & Volschenk, E.S. (2007) The systematics of the Gondwanan pseudoscorpion family Hyidae (Pseudoscorpiones: Neobisioidea): new data and a revised phylogenetic hypothesis. Invertebrate Systematics, 21, 365–406.

    Harvey, M.S., Ratnaweera, P.B., Udagama, P.V. & Wijesinghe, M.R. (2012) A new species of the pseudoscorpion genus Megachernes (Pseudoscorpiones: Chernetidae) associated with a threatened Sri Lankan rainforest rodent, with a review of host associations of Megachernes. Journal of Natural History, 46, 2519–2535.

    Hughes, G.B. & Moore, W. (2018) Monophyly of the subfamily Neobisiinae (Pseudoscorpiones: Neobisiidae). Journal of Arachnology, 46, 481–487

    Judson, M.L.I. (2007) A new and endangered species of the pseudoscorpion genus Lagynochthonius from a cave in Vietnam, with notes on chelal morphology and the composition of the Tyrannochthoniini (Arachnida, Chelonethi, Chthoniidae). Zootaxa, 1627 (1), 53–68.

    Murienne, J., Harvey, M.S. & Giribet, G. (2008) First molecular phylogeny of the major clades of Pseudoscorpiones (Arthropoda: Chelicerata). Molecular Phylogenetics and Evolution, 49, 170–184.

    Zaragoza, J.A. (2008) On the status of the subspecies of Roncocreagris galeonuda (Pseudoscorpiones:Neobisiidae): importance of the chelal microsetae pattern. Remarks on the genus Roncocreagris Mahnert. Revista Ibérica de Aracnología, 15, 35–46.

    Zaragoza, J.A. (2017) Revision of the Ephippiochthonius complex (Pseudoscorpiones, Chthoniidae) in the Iberian Peninsula, Balearic Islands and Macaronesia, with proposed changes to the status of the Chthonius subgenera. Zootaxa, 4246 (1), 1– 221.