Abstract
In this contribution to the study of gryllacridines or raspy crickets (Orthoptera: Gryllacrididae), a new proposal for classification of this family is provided, dividing it into two subfamilies and ten tribes that includes most of the 114 known genera to date (including the new genera described here). It describes and redefines two subfamilies: Hyperbaeninae n. subf. and Gryllacridinae n. sensu, ten tribes: Phryganogryllacridini n. trib., Capnogryllacridini n. trib., Asarcogryllacridini n. trib., Hyperbaenini n. trib., Paragryllacridini n. trib., Ametrini n. trib., Ametroidini n. trib., Gryllacridini n. sensu., Eremini n. trib. and Progryllacridini n. trib.; ten genera: Claudiagryllacris n. gen., Griffinigryllacris n. gen., Gorochovgryllacris n. gen., Ingrishgryllacris n. gen., Rentzgryllacris n. gen., Willemsegryllacris n. gen., Karnygryllacris n. gen., Brunnergryllacris n. gen., Bianigryllacris n. gen. and Hugelgryllacris n. gen.; seven genera groups: Gryllacrae n. group. (placed under Gryllacridini n. trib. comprising ten genera: Caustogryllacris, Eugryllacris, Gryllacris, Lyperogryllacris, Nesogryllacris, Ocellarnaca, Phlebogryllacris, Prosopogryllacris, Willemsegryllacris n. gen. and Xanthogryllacris), Metriogryllacrae n. group. (under Gryllacridini grouped Metriogryllacris, Homogryllacris, Pseudasarca n. stat. and Furcilarnaca), Anancistrogerae (Anancistrogera, Ancistrogera, Angustogryllacris, Aphanogryllacris, Celebogryllacris), Triaenogryllacrae n. group. (under Gryllacridinae only comprising Triaenogryllacris) the next groups under Ametrini n. trib.: Ametrae n. group. (comprising two genera: Ametrus and Pareremus), Apotrechae n. group. (including three genera: Apotrechus, Apterolarnaca and Bianigryllacris n. gen.) and Apteronomae n. group. (comprising two genera: Ametrosomus and Apteronomus). The status of Dictogryllacris reinst. stat., was restored, previously transferred by Gorochov 2003 as subgenus of Capnogryllacris and to Pseudasarca n. stat. as full genus status is proposed. One genera, one subgenus and one fossil species are synonymized: Xiphogryllacris n. syn. (under Hyalogryllacris) and †Gryllacris brevippennis n. syn. (under †Macrelcana ungeri). The subgenus Glolarnaca n. stat. (included under Zalarnaca), is considered as full genus in Gryllacridinae (Gryllacridini). 57 new combinations are proposed, with particular emphasis on Niphetogryllacris, placing 36 of the 43 existing species of the genus: Pissodogryllacris tesellata n. comb., Willemsegryllacris barnesi n. comb. (previously placed under Gryllacris), Afrogryllacris nigripceps n. comb. (previously placed under in Barombogryllacris), Anancistrogera nigroscutata n. comb., A. cornualis n. comb. and A. genualis n. comb. (included by Karny in Caustogryllacris and by Gorochov in the subgenus Pseudolarnaca, here transfered to Anancistrogera), Hugelgryllacris tchancha n. comb. (before in Psilogryllacris), Diaphanogryllacris annandalei n. comb., D. barkudensis n. comb., D. dravida n. comb., D. gravelyi n. comb., Claudiagryllacris finoti n. comb. C. stigmata n. comb., C. lemur n. comb., C. fryeri n. comb., Stictogrtllacris pungens n. comb., S. madagassa n. comb., S. genufuscata n. comb., S. vosseleri n. comb., S. difficilis n. comb., S. paulani n. comb., Stictogryllacris signoreti n. comb., S. indecisa n. comb., S. conspersa n. comb., S. pittarellii n. comb., S. ametroides n. comb., S. jacobi n. comb., S. kilimandjarica n. comb., S. meruensis n. comb.; S. neglecta n. comb.; S. submutica n. comb., Griffinigryllacris reunionis n. comb., G. adelungi n. comb., G. mauritiana n. comb., Karnygryllacris occipitalis n. comb., K. atriceps n. comb., K. brevipennis n. comb., K. humilis n. comb., K. scurra n. comb., K. triocellata n. comb., K. pittarellii n. comb., K. grylloides n. comb., Brunnergryllacris testaceus n. comb. and B. eximia n. comb (previously placed under Niphetogryllacris), Rentzgryllacris sechellensis n. comb. (before in Prosopogryllacris and the three subspecies are treated as full species), Gorochovgryllacris navicula n. comb. (transferred from Brachybaenus), Bianigryllacris trilobus n. comb., B. bilobus n. comb., B. digitatus n. comb., B. fallax n. comb., B. nigrigeniculatus n. comb., B. parvospinus n. comb., B. quadratus n. comb. and B. transversus n. comb. (previously placed under Apotrechus), Hyalogryllacris orthoxipha n. comb. (previously placed under Xiphogryllacris), Afroneanias glauningi n. comb., A. sphinix n. comb. and Ingrishgryllacris brevifalcatus n. comb. (previously placed under Ametroides and Glomeremus respectively). A key to the identification of the subfamilies and tribes is provided, plus historical background of the major taxonomic works on the group and the few contributions on ecology, morphology, and behavior is also done. It also discusses the status of fossil taxa thought to belong to the family. A preliminary analysis of the zoogeography of the family from the parameters of richness, diversity and distribution patterns of different groups studied, and also, some final comments on what has been achieved in this contribution are give, what is missing to the study of gryllacridines to the future and some problems that still affect the taxonomy to genera and species level. Finally, a check list in which all family taxa are included with the new classification is provided, with some comments on their distribution and taxonomic status if necessary.
References
Ander, K. (1934) Entomologiske Meddelelser, 18 (5), 485–490.
Ander, K. (1939) Vergleichend-anatomische und phylogenetische Studien über die Ensifera (Saltatoria). Opuscula Entomologica, 2 (Supplement), 1–306.
Bey-Bienko, G.Y. (1957) Results of Chinese-Soviet zoological-botanical expeditions to south-western China 1955–1956: Tettigonioidea (Orthoptera) of Yunnan. Entomologicheskoe Obozrenie, 36, 401–417.
Bey-Bienko, G.Y. (1962) Results of the Chinese-Soviet zoological-botanical expeditions to south-western China 1955-1957. New or less known Tettigonioidea (Orthoptera) from Szechuan and Yunnan. Trudy Zoologitscheskogo Instituta, Akademiia Nauk SSSR, Leningrad [= Proceedings of the Zoological Institute, USSR Academy of Sciences, Leningrad], 30, 110–138.
Bian, X. & Shi, F.M. (2015) A newly recorded genus Zalarnaca Gorochov (Orthoptera: Stenopelmatoidea: Gryllacridinae) from China. Far Eastern Entomologist, 302, 10–13.
Bian, X. & Shi, F.M. (2016a) Review of the genus Eugryllacris Karny, 1937 (Orthoptera: Gryllacridinae) from China. Zootaxa, 4066 (4), 438–450.
https://doi.org/10.11646/zootaxa.4066.4.5
Bian, X. & Shi, F.M. (2016b) Review of the genus Neolarnaca Gorochov, 2005 (Orthoptera: Gryllacrididae) from China. Zoological Systematics, 41 (2), 227–230.
https://doi.org/10.11865/zs.201621
Bian, X.; Shi, F.M. & Guo, L. (2013a) Review of the genus Ocellarnaca Gorochov, 2004 (Orthoptera: Gryllacrididae: Gryllacridinae) of China. Journal of Orthoptera Research, 22 (1), 57–66.
https://doi.org/10.1665/034.022.0109
Bian, X.; Shi, F.M. & Guo, L. (2013b) Review of the genus Furcilarnaca Gorochov, 2004 (Orthoptera: Gryllacrididae, Gryllacridinae) from China. Far Eastern Entomologist, 268, 1–8.
Bian, X., Guo, L. & Shi, F.M. (2015) A new record of Larnaca Walker, 1869 (Orthoptera: Gryllacrididae: Gryllacridinae) from China with description of a new species. Zootaxa, 3994, 292–294.
https://doi.org/10.11646/zootaxa.3994.2.9
Bian, X., Lu, R.S. & Shi, F.M. (2016) Review of the genus Apterolarnaca Gorochov (Orthoptera: Gryllacrididae) with proposal of a new synonym and description two new species. Zootaxa, 4079 (3), 357–362.
https://doi.org/10.11646/zootaxa.4079.3.3
Blanchard, E. (1845) Histoire naturelle des insectes. Colèoptères, Orthoptères, Thysanoptères, Névroptères, Lépidoptères, Hémiptères, Aphaniptères, Strepsiptères, Diptères, Anoplures et Thysanures. Histoire des insectes : traitant de leurs moeurs et de leurs métamorphoses en général et comprenant une nouvelle classification fondée sur leurs rapports naturels, 2, 1–524.
Bland, R.G. & Rentz, D.C.F. (1991a) External morphology and abundance of mouthpart sensilla in Australian Gryllacrididae, Stenopelmatidae and Tettigoniidae. Journal of Morphology, 207 (3), 315–325.
https://doi.org/10.1002/jmor.1052070308
Bland, R.G. & Rentz, D.C.F. (1991b) Studies in Australian Gryllacrididae: the proventriculus as a taxonomic character. Invertebrate Taxonomy, 5, 443–455.
https://doi.org/10.1071/IT9910443
Bland, R.G. & Rentz, D.C.F. (1994) External morphology and abundance of antennal sensilla in Australian Gryllacrididae. Journal of Morphology, 222 (1), 11–18.
https://doi.org/10.1002/jmor.1052220103
Bolívar, I. (1895) Mission scientifique de M. Ch. Alluaud aux îles Séchelles. Orthoptères. Annales de la Société Entomologique de France, 64, 369–385.
Bolívar, I. (1912) Orthoptera. Acrydidae. Phasgonuridae. Gryllidae (of the Seychelles). The Transactions of the Linnean Society of London, 15 (2), 263–292.
https://doi.org/10.1111/j.1096-3642.1912.tb00101.x
Brunner von Wattenwyl, C. (1888) Monographie der Stenopelmatiden und Gryllacriden. Verhandlungen der Kaiserlich-Königlichen Zoologisch-Botanischen Gesellschaft in Wien, 38, 247–394.
Brunner von Wattenwyl, C. (1898) Orthopteren des Malayischen Archipels gesammelt von Prof. Dr. W. Kükenthal in den Jahren 1893 and 1894. Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft, 24 (2), 193–288.
Burmeister, H. (1838) Kaukerfe, Gymnognatha (Erste Hälfte: Vulgo Orthoptera). Handbuch der Entomologie, 22 (I–VIII), 397–756.
Cadena-Castañeda, O.J. & Cortés-Torres, C. (2013) New species and taxonomic rearrangements of Anostostomatid Crickets (Orthoptera: Stenopelmatoidea: Anostostomatidae) from the Neotropics. Zootaxa, 3664 (3), 335–348.
https://doi.org/10.11646/zootaxa.3664.3.4
Cadena-Castañeda, O.J., Rodríguez, W.D. & Navarrete-Heredia, J.L. (2019) Orthoptera (Ensifera & Caelifera) collected using carrion traps in a Quercus forest in Jalisco, Mexico, with description of a new genus and a new species (Trigonidiidae: Nemobiinae). Zootaxa, 4550 (3), 401–415.
https://doi.org/10.11646/zootaxa.4550.3.7
Charpentier, T. (1843) Über einige fossile Insekten aus Radoboj in Croatien. Verhandlungen der Kaiserlich Leopoldinisch-Carolinischen Deutschen Akademie der Naturforscher, 12 (1), 399–410.
Chopard, L. (1937) Description de trois Gryllacrides nouveaux (Orth.). Bulletin de la Société Entomologique de France, 42, 77–80.
Chopard, L. (1952) Notes sur les Orthopteroides de Madagascar IV. Faune de la forêt de mousses du Tsaratanana. Mémoires de l'Institut Scientifique de Madagascar, Série E Entomologie, 1 (2), 463–516.
Chopard, L. (1954) La Réserve naturelle intégrale du Mt. Nimba, III. Orthoptères Ensifères. Mémoires de l'Institut Français d'Afrique Noire, 40 (2), 25–97.
Chopard, L. (1958) Les Orthopteroides des Comores. Mémoires de l'Institut Scientifique de Madagascar, Série E Entomologie, 10, 3–40.
Cockerell, T.D.A. (1909) Description of Tertiary insects VII. American Journal of Science. 4 (28), 283–286.
https://doi.org/10.2475/ajs.s4-28.165.283
Deichmüller, L. (1886) Die Insecten aus dem lithographischen Schiefer in Dresdener Museum. Mitteilungen aus dem Königlichen Mineralogisch-Geologischen und Prähistorischen Museum in Dresden, 7, 1–88.
Drury, D. (1770) Illustrations of Natural History, wherein are exhibited upwards of 240 figures of Exotic Insects. Printed for the author and sold by B. White, London, 70 pp.
Du, B.J., Bian, X. & Shi, F.M. (2016a) Review of the genus Diaphanogryllacris (Orthoptera: Gryllacrididae: Gryllacridinae) with description two new species from China. Zootaxa, 4079 (4), 448–456.
https://doi.org/10.11646/zootaxa.4079.4.4
Du, B.J., Bian, X. & Shi, F.M. (2016b) Notes on the genus Homogryllacris Liu, 2007 (Orthoptera: Gryllacrididae: Gryllacridinae) with description of two new species from China. Zootaxa, 4111 (2), 187–193.
https://doi.org/10.11646/zootaxa.4111.2.7
Cigliano, M.M., Braun, H., Eades, D.C. & Otte, D. (2019) Orthoptera Species File. Version 5.0/5.0. Available from: http://Orthoptera.SpeciesFile.org (accessed 20 January 2019).
Field, L.H. & Bailey, W.J. (1997) Sound production in primitive Orthoptera from Western Australia: sounds used in defence and social communication in Ametrus sp. and Hadrogryllacris sp. (Gryllacrididae: Orthoptera). Journal of Natural History, 31, 1127–1141.
https://doi.org/10.1080/00222939700770591
Gerstaecker, A. (1860) Ueber die Locustinen-Gattung Gryllacris. Archiv für Naturgeschichte, 26, 245–278.
Giglio-Tos, E. (1907) Ortotteri Africani. Parte I Acridoidea. Bollettino dei Musei di Zoologia ed Anatomia Comparata della R. Università di Torino, 22 (554), 1–35.
Gorochov, A.V. (1995) System and evolution of the suborder Ensifera (Orthoptera). Part 1 [in Russian]. TrudyvZoologicheskogo Instituta, 260 (1), 1–224.
Gorochov, A.V. (2001) The higher classification, phylogeny and evolution of the superfamily Stenopelmatoidea. In: Field, L.H. (Ed.), The biology of wetas, king crickets and their allies. CABI Publishing, Wallingford, pp. 3–33.
https://doi.org/10.1079/9780851994086.0003
Gorochov, A.V. (2004) A contribution to the fauna of Stenopelmatoidea (Orthoptera) of Indochina and some other territories. V. Entomological Review [English translation of Entomologicheskoe Obozrenie], 84 (8), 900–921.
Gorochov, A.V. (2005) Contribution to the fauna and systematics of the Stenopelmatoidea (Orthoptera) of Indo-China and some other territories. VI. Entomologicheskoe Obozrenie, 84 (4), 806–825.
Gorochov, A.V. (2007) Contribution to the fauna and systematics of the Stenopelmatoidea (Orthoptera) of Indochina and some other territories. VII. Entomologicheskoe Obozrenie, 86 (4), 845–862.
https://doi.org/10.1134/S0013873807090151
Gorochov, A.V. (2008) Contributions to the fauna and systematics of the Stenopelmatoidea (Orthoptera) of Indochina and some other territories: VIII. Entomological Review [English translation of Entomologicheskoe Obozrenie], 88 (4), 406–420.
https://doi.org/10.1134/S0013873808040040
Gorochov, A.V. (2010) New and little-known orthopteroid insects (Polyneuroptera) from fossil resins: Communication 4. Paleontologicheskii Zhurnal, 6, 656–671.
https://doi.org/10.1134/S0031030110060080
Gorochov, A.V. & Woznessenskij, A.J. (2000) Species of Diaphanogryllacris from Indochina and adjacent territories (Orthoptera: Stenopelmatidae: Gryllacridinae). Zoosystematica Rossica, 8 (2), 257–265.
Gorochov, A.V., Dawwrueng, P. & Artchawakom, T. (2015a) Study of Gryllacridinae (Orthoptera: Stenopelmatidae) from Thailand and adjacent countries: the genera Ultragryllacris gen. nov. and Capnogryllacris. Zootaxa, 4021 (4), 565–577.
https://doi.org/10.11646/zootaxa.4021.4.6
Gorochov, A.V., Dawwrueng, P. & Artchawakom, T. (2015b) Study of Gryllacridinae (Orthoptera: Stenopelmatidae) from Thailand and adjacent countries: the genera Diaphanogryllacris and Gryllacris. Zoosystematica Rossica, 24 (2), 219–235.
Gorochov, A.V. & Cadena-Castañeda, O.J. (2016) New and little known Stenopelmatoidea (Orthoptera: Ensifera) from America. Zoosystematica Rossica, 25 (1), 98–143.
Green, O.R. & Graeme, W.R. (2003) A winged weta, Pterapotrechus (Orthoptera: Gryllacrididae), established in New Zealand. New Zealand Entomologist, 26, 75–77.
https://doi.org/10.1080/00779962.2003.9722111
Griffini, A. (1908a) Le specie africane del genere Gryllacris Serv. Studio monografico, 1908, 1–56.
Griffini, A. (1908b) Intorno a quattro Grillacridi dell’America meridionale. Zoologischer Anzeiger, 33 (2–3), 60–67.
Griffini, A. (1909a) Il sottogenere Papuogryllacris. Zoologische Jahrbücher. Abteilung für Systematik, Geographie und Biologie der Tiere, 28 (2), 139–154.
Griffini, A. (1909b) Intorno ad una nuova Gryllacris di Madagascar. Wiener Entomologische Zeitung, 28 (11), 66–69.
Griffini, A. (1909c) Studi sui Grillacridi del Museo di Oxford. Parte Ia, specie etiopiche, indo-malesi ed australiane. Atti della Società Italiana di Scienze Naturali e del Museo Civico di Storia Naturale di Milano, 47, 300–338.
Griffini, A. (1911a) Nota sopra alcuni Stenopelmatidi e Gryllacridi del Museo de Sarawak. Bollettino dei Musei di Zoologia ed Anatomia Comparata della R. Università di Torino, 26 (636), 1–19.
Griffini, A. (1911b) Note intorno ad alcuni Grillacridi e Stenopelmatidi del Muséum d'histoire naturelle de Genève. Revue Suisse de Zoologie, 19, 461–500.
Griffini, A. (1911c) Notes sur quelques Gryllacrides du Musée Zoologique de l'Académie Impériale des Sciences à St. Pétersbourg. Annuaire du Musée Zoologique de l'Académie Impériale des Sciences de St.-Pétersbourg, 16, 65–85.
Griffini, A. (1911d) Le species africane del genere Neanias Brunner. Zoologische Jahrbücher. Abteilung für Systematik, Geographie und Biologie der Tiere, 31, 461–468.
Griffini, A. (1912) Un nuovo genere di Grillacridi dell’Africa Orientale. Atti della Società Italiana di Scienze Naturali e del Museo Civico di Storia Naturale di Milano, 50, 43–51.
Griffini, A. (1913) Note sopra Grillacridi australiani, indo-malesi ed etiopici [del Museum d' Histoire naturelle di Parigi]. Atti della Società Italiana di Scienze Naturali e del Museo Civico di Storia Naturale di Milano, 51, 217–246.
Griffini, A. (1914a) Studi sui Grillacridi dell'Indian Museum di Calcutta. Atti della Società Italiana di Scienze Naturali e del Museo Civico di Storia Naturale di Milano, 52, 201–274.
Griffini, A. (1914b) I Grillacridi del Tonkino. Studio monografico. Zoologische Jahrbücher. Abteilung für Systematik, Geographie und Biologie der Tiere, 38, 79–108.
Guo, L. & Shi, F.M. (2012) Notes on the genus Apotrechus (Orthoptera: Gryllacrididae: Gryllacridinae) from China. Zootaxa, 3177, 52–58.
Guo, L. & Shi, F.M. (2011) Descriptions of new species of the genus Woznessenskia (Orthoptera: Gryllacrididae: Gryllacridinae) from China. Zootaxa, 2784 (1), 62–68.
https://doi.org/10.11646/zootaxa.2784.1.6
Hale, R.J. & Rentz, D.C.F. (2001) The Gryllacrididae: An overview of the world fauna with emphasis on Australian examples. In: Field, L.H. (Ed.), The biology of wetas, king crickets and their allies. CABI Publishing, Wallingford, pp. 1–540.
https://doi.org/10.1079/9780851994086.0095
Handlirsch, A.P.J. (1907) Die fossilen Insekten und die Phylogenie der rezenten Formen. Ein Handbuch für Paläontologen und Zoologen, 1 (5), 641–800.
Heer, O. (1849) Zweiter Theil: Heuschrecken, Florfliegen, Aderflügler, Schmetterlinge und Fliegen. Die Insektenfauna der Tertiärgebilde von Oeningen und von Radoboj in Croatien, 1849, 1–131.
Hugel, S., Claire, M., Jacques, F., Ben H.W., Gauvin-Bialecki, A., Pailler, T., Chase, M.W. & Strasberg, D. (2010) Glomeremus species from the Mascarene islands (Orthoptera, Gryllacrididae) with the description of the pollinator of an endemic orchid from the island of Réunion. Zootaxa, 2545, 58–68.
https://doi.org/10.11646/zootaxa.2545.1.6
Ichikawa, A. (2001) Japanese Stenopelmatidae (Orthoptera) with some notes on some species. Tettigonia: Memoirs of the Orthopterological Society of Japan, 3, 69–74.
Ingrisch, S. (2018) New taxa and records of Gryllacrididae (Orthoptera, Stenopelmatoidea) from South East Asia and New Guinea with a key to the genera. Zootaxa, 4510 (1), 1–278.
https://doi.org/10.11646/zootaxa.4510.1.1
Jost, M.C. & Shaw. K.L. (2006) Phylogeny of Ensifera (Hexapoda: Orthoptera) using three ribosomal loci, with implications for the evolution of acoustic communication. Molecular Phylogenetics and Evolution, 38, 510–530.
https://doi.org/10.1016/j.ympev.2005.10.004
Karny, H.H. (1910) Orthoptera (s. str.). In Schultze, L. Zoologische und anthropologische Ergebnisse einer Forschungsreise im westlichen und zentralen Südafrika ausgefürhrt in den Jahren 1903-1905. Denkschriften der Medizinisch-Naturwissenschaftlichen Gesellschaft zu Jena, 4, 35–90.
Karny, H.H. (1923) On Malaysian katydids (Gryllacridae and Tettigoniidae) from the Raffles Museum, Singapore. Journal of the Malaysian Branch of the Royal Asiatic Society, 1, 117–193.
Karny, H.H. (1924) Beiträge zur malayischen Orthopterenfauna VI. Die Gryllacriden des Buitenzorger Museums. Treubia, 5 (1–3), 19–105.
Karny, H.H. (1927). Revision der Gryllacriden des Zoologischen Institutes in Halle a.S., sowie einiger Tettigoniiden-Typen von Burmeister und Giebel. Zeitschrift für die gesammten Naturwissenschaften, 88, 1–14.
Karny, H.H. (1928a) Revision der Gryllacriden der Stuttgarter Naturaliensammlung. Zoologische Jahrbücher. Abteilung für Systematik, Geographie und Biologie der Tiere, 55, 501–534.
Karny, H.H. (1928b) Revision der Gryllacriden des ungarischen National-Museums. Annales Historico-Naturales Musei Nationalis Hungarici, 25, 215–260.
Karny, H.H. (1928c) Revisione dei Grillacridi dei Musei di Genova e Torino e della collezione Griffini. Memorie della Società Entomologica Italiana, 7, 5–153.
Karny, H. H. (1928d) Gryllacriden aus verschiedenen deutschen und österreichischen Sammlungen. Stettiner Entomologische Zeitung, 89, 247–312.
Karny, H.H. (1929a) Revision der Gryllacriden des Naturhistorischen Museums in Wien einschlieBlich der Collection Brunner v. Wattenwyl. 1. Hälfte. Annalen des Naturhistorischen Museums in Wien, 43, 3–186.
Karny, H.H. (1929b) Ueber einige Gryllacrinen des Madrider Naturwissenschaftlichen Museums (Orth. Tettig.). Eos, Revista española de Entomología, 5 (1), 33–80.
Karny, H.H. (1929c) Ueber einige Gryllacrinen des Madrider Naturwissenschaftlichen Museums (Orth. Tettig.). Eos, Revista española de Entomología, 5 (1), 33–80.
Karny, H.H. (1930a) Revision der Gryllacriden des Naturhistorischen Museums in Wien einschließlich der Collection Brunner v. Wattenwyl. 2. Hälfte. Annalen des Naturhistorischen Museums in Wien, 44, 45–198.
Karny, H.H. (1930b) On the geographical distribution of the Pacific Gryllacrids. Proceedings of the fourth Pacific Science Congress, 3, 157–172.
Karny, H.H. (1931a) Über die Gryllacrinen des Senckenbergischen Museums in Frankfurt a. M. Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft, 42 (5), 383–436.
Karny, H.H. (1931b) On the geographical distribution of the Indo-African and Mediterranean Gryllacrids. Transactions of the Royal Society of South Africa, 19, 99–105.
https://doi.org/10.1080/00359193109518828
Karny, H.H. (1932) Die Gryllacrididen des Pariser Museums und der Collection L. Chopard. Eos, Revista española de Entomología, 8, 235–309.
Karny, H.H. (1932b) Über zwei angebliche Gryllacris Arten aus dem Miocan von Radoboj. Jahrbuch der Geologischen Bundesanstalt, 82 (1–2), 65–69.
Karny, H.H. (1935) Die Gryllacrididen des Pariser Museums und der Collection L. Chopard. Eos, Revista española de Entomología, 10 (3–4), 293–393.
Karny, H.H. (1937) Orthoptera Fam. Gryllacrididae, Subfamiliae Omnes. Genera Insectorum, 206, 1–317.
Karsch, F.A.F. (1891) Übersicht der von Dr. Paul Preuss auf der Barombi-Station in Kamerun gesammelten Locustodeen. Berliner Entomologische Zeitschrift, 36 (2), 317–346.
Krenn, H., Fournel. J., Bauder, J. & Hugel, S. (2016) Mouthparts and nectar feeding of the flower visiting cricket Glomeremus orchidophilus (Gryllacrididae). Arthropod Structure & Development, 45 (3), 221–229.
https://doi.org/10.1016/j.asd.2016.03.002
Li M., Fang, Y., Liu, X. & Li, K. (2014a) Taxonomic revision of the genus Phryganogryllacris (Orthoptera: Gryllacrididae) from China. Zoological Systematics, 39 (4), 507–519.
https://doi.org/10.11865/zs.20140404
Li, M., Fang, Y., Liu, X. & Li, K. (2014b) Review of the species of the genus Marthogryllacris (Orthoptera, Gryllacrididae, Gryllacridinae). Zootaxa, 3889 (2), 277–288.
https://doi.org/10.11646/zootaxa.3889.2.6
Li, M., Fang, Y., Liu, X. & Li, K. (2014c) A taxonomic study on the species of the genus Ocellarnaca (Orthoptera, Gryllacrididae, Gryllacridinae). Zootaxa, 3835 (1), 127–139.
https:// doi.org/10.11646/zootaxa.3835.1.8
Li, M., Sun, M.L., Liu, X. & Li, K. (2015) A taxonomic study on the species of the genus Furcilarnaca (Orthoptera, Gryllacrididae, Gryllacridinae). Zootaxa, 4039 (3), 418–430.
https://doi.org/10.11646/zootaxa.4039.3.2
Li, M., Liu, X. & Li, K. (2015) Review of the genus Apotrechus in China (Orthoptera, Gryllacrididae, Gryllacridinae). ZooKeys, 482, 143–155.
https://doi.org/10.3897/zookeys.482.8713
Liu, X. (2007) A new genus of the subfamily Gryllacrinae from China (Orthoptera: Stenopelmatidae: Gryllacridae). Scientific Research Monthly, 6 (7), 1–2.
Liu, X. & Yin, H. (2002) Two new species of Gryllacridinae from China (Orthoptera: Stenopelmatidae). Zoological Research, 23 (5), 417–418.
Liu, X. & Bi, W. (2008) Two new species of the genus Apotrechus (Orthoptera: Stenopelmatidae: Gryllacrinae). Entomotaxonomia, 30 (1), 11–15.
Liu, X., Bi, W. & Zhang, F. (2010) Orthoptera: Stenopelmatoidea. Xu & Ye (Eds.). Insects of Fengyangshan National Nature Rreserve, 53–68.
Morton, S.R. & Rentz, D.C.F. 1983. Ecology and taxonomy of fossorial, granivorous gryllacridids (Orthoptera: Gryllacrididae) from arid central Australia. Australian Journal of Zoology, 31 (4), 557–579.
https://doi.org/10.1071/ZO9830557
Olivier, G.A. (1789) Histoire Naturelle. Insectes. Encyclopédie méthodique. Histoire naturelle. Entomologie, ou histoire naturelle des crustacés, des arachnides et des insectes, 4, 12–26.
Otte, D. (2000) Gryllacrididae, Stenopelmatidae, Cooloolidae, Schizodactylidae, Anastostomatidae and Rhaphidophoridae. Orthoptera Species File. No. 8. Orthopterists Society, Philadelphia, 97 pp.
Piza Jr., S.T. (1975) Duas novas espécies de Gryllacris do Brasil (Orthopera, Gryllacridae). Revista de Agricultura (Piracicaba), 50 (3–4), 93–94.
Ragge, D.R. (1955) Wing-Venation of the Orthoptera Saltatoria with notes on Dictyopteran Wing-Venation. Printed by order of the Trustees of the British Museum, London, 159 pp.
Rehn, J.A.G. (1903) A contribution to the knowledge of the Orthoptera of Mexico and Central America. Transactions of the American Entomological Society, 29 (1), 1–34.
Rentz, D.C.F. (1997) The world's most unusual gryllacridid (Orthoptera: Gryllacrididae). Journal of Orthoptera Research, 6, 57–68.
https://doi.org/10.2307/3503536
Rentz, D.C.F. & John, B. (1990) Studies in Australian Gryllacrididae: taxonomy, biology, ecology and cytology. Invertebrate Taxonomy, 3 (8), 1053–1210.
https://doi.org/10.1071/IT9891053
Rentz, D.C.F., Su, Y. & Béthoux, O. (2018) A new raspy cricket from the northern Australian rainforests (Orthoptera: Gryllacrididae). Zootaxa, 4514 (1), 41–52.
https://doi.org/10.11646/zootaxa.4514.1.3
Scudder, S.H. (1890) Pre-Tertiary insects of North America. Fossil insects of North America, 1, 1–455.
Serville, J.G.A. (1831) Revue méthodique des insectes de l'ordre des Orthoptères. Annales des Sciences Naturelles, 22 (86), 134–167.
Sharov, A.G. (1968) Filogniya orthopteroidnykh nasekomykh [1971 English translation: Phylogeny of the Orthopteroidea]. Trudy Paleontologicheskogo Instituta, Akademiia Nauk SSSR [= Transactions of the Institute of Paleontology, USSR Academy of Sciences], 118, 1–216.
Shi, F.M., Guo, L. & Bian, X. (2012) Remarks on the genus Homogryllacris Liu, 2007 (Orthoptera: Gryllacrididae: Gryllacridinae). Zootaxa, 3414, 58–68.
Shi, F.M., Bian, X. & Guo, L. (2016) First record of the genus Dialarnaca Gorochov, 2005 from China, with description of two new species (Orthoptera, Gryllacrididae, Gryllacridinae). ZooKeys, 558, 1–8.
https://doi.org/10.3897/zookeys.558.6165
Song, H., Amédégnato, C., Cigliano, M.M., Desutter-Grandcolas, L., Heads, S., Huang, Y., Otte, D. & Whiting, M.F. (2015) 300 million years of diversification: elucidating the patterns of orthopteran evolution based on comprehensive taxon and gene sampling. Cladistics, 31 (6), 621–651.
https://doi.org/10.1111/cla.12116
Strauss, J. & Lakes-Harlan, Reinhard. (2008) Neuroanatomy and Physiology of the Complex Tibial Organ of an Atympanate Ensiferan, Ametrus tibialis (Brunner von Wattenwyl, 1888) (Gryllacrididae, Orthoptera) and Evolutionary Implications. Brain Behavior and Evolution, 71 (3), 167–180.
https://doi.org/10.1159/000114405
Tan, M.K. & Wahab, R.B.A. (2018) Notes on the taxonomy of poorly known Gryllacrididae (Stenopelmatoidea) from Brunei Darussalam, Borneo. Zootaxa, 4462 (4), 579–591.
https://doi.org/10.11646/zootaxa.4462.4.9
Tepper, J.G.O. (1892) The Gryllacrididae and Stenopelmatidae of Australia and Polynesia. Transactions of the Royal Society of South Australia, 15, 137–178.
Uhler, P.R. (1864) Orthopterological Contributions. Proceedings of the Entomological Society of Philadelphia, 2, 543–555.
Uvarov, B.P. & Popov, G.B. (1957) The saltatorial Orthoptera of Socotra. Zoological Journal of the Linnean Society, 43, 359–389.
https://doi.org/10.1111/j.1096-3642.1957.tb01558.x
Walker, A.A., Weisman, S., Jeffrey, S., Church, D., Merritt, J., Mudie, S.T. & Sutherland, T.D. (2012) Silk from crickets: a new twist on spinning. PLOS ONE, 7 (2), e30408.
https://doi.org/10.1371/journal.pone.0030408
Walker, F. (1869) Catalogue of the Specimens of Dermaptera Saltatoria in the Collection of the British Museum, 1, 1–224.
Willemse, C. (1953) On a collection of Orthoptera from the Solomon Islands, chiefly from the Institut Royal des Sciences naturelles de Belgique. Bulletin de l'Institut Royal des Sciences Naturelles de Belgique, 29 (7), 1–27.
Zeuner, F.E. (1939) Fossil Orthoptera, Ensifera. Printed by order of the Trustees of the British museum, London, 321 pp.