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Type: Article
Published: 2018-08-14
Page range: 53–84
Abstract views: 76
PDF downloaded: 3

The millipede genus Epanerchodus Attems, 1901 in continental China, with descriptions of seven new cavernicolous species (Diplopoda, Polydesmida, Polydesmidae)

Department of Entomology, College of Agriculture, South China Agricultural University, 483 Wushanlu, Guangzhou 510642, China
Institute for Problems of Ecology and Evolution, Russian Academy of Sciences, Leninsky pr. 33, Moscow 119071, Russia
Myriapoda Epanerchodus new species troglobite key southern China


Seven new species of Epanerchodus Attems, 1901 are described from caves in southern China: Epanerchodus latus n. sp., Epanerchodus tujiaphilus n. sp., Epanerchodus jiangxiensis n. sp., Epanerchodus coniger n. sp., Epanerchodus parvus n. sp., Epanerchodus gladiatus n. sp. and Epanerchodus chutou n. sp. The first three species are from Chongqing, Hunan and Jiangxi, respectively; the last four are from Guizhou. Additional records, illustrations and descriptive notes are given for the following troglobitic species: Epanerchodus lipsae Golovatch & Geoffroy, 2014, from Sichuan; Epanerchodus soror Geoffroy & Golovatch, 2004, from Yunnan; Epanerchodus stylotarseus Chen & Zhang, 1990, from Guizhou; Epanerchodus varius Geoffroy & Golovatch, 2004, from Hubei, Sichuan and Chongqing; and Epanerchodus draco Geoffroy & Golovatch, 2004, from Yunnan and Guizhou. New records of E. varius from Chongqing and E. draco from Guizhou are documented. A key and distribution maps are provided to all 25 species of Epanerchodus currently known to occur in mainland China.



  1. Attems, C.G. (1901) Neue Polydesmiden des Hamburger Museums. Mitteilungen aus dem Naturhistorischen Museum in Hamburg, Jahrgang 18, 83–107.

    Chen, J.X. & Zhang, C.Z. (1990) A cave-dwelling new species of the diplopod genus Epanerchodus from Guizhou Province (Polydesmida: Polydesmidae). Acta Zootaxonomica Sinica, 15, 406–409. [in Chinese, English summary]

    Enghoff, H., Golovatch, S.I., Short, M., Stoev, P. & Wesener, T. (2015) Diplopoda—taxonomic overview. In: Minelli, A. (Ed.) Treatise on Zoology—Anatomy, Taxonomy, Biology. The Myriapoda, 2 (16), pp. 363–453.

    Geoffroy, J.-J. & Golovatch, S.I. (2004) Some polydesmidan millipedes from caves in southern China (Diplopoda: Polydesmida), with descriptions of four new species. Arthropoda Selecta, 13 (1–2), 19–28.

    Golovatch, S.I. (1991) The millipede family Polydesmidae in Southeast Asia, with notes on phylogeny (Diplopoda: Polydesmida). Steenstrupia, 17 (4), 141–159.

    Golovatch, S.I. (2014a) Review of the millipede genus Epanerchodus Attems, 1901 in continental China, with descriptions of new species (Diplopoda: Polydesmidae). Zootaxa, 3760 (2), 275–288.

    Golovatch, S.I. (2014b) Two new and one little-known species of the millipede genus Epanerchodus Attems, 1901 from southern China (Diplopoda, Polydesmida, Polydesmidae). Fragmenta Faunistica, 56 (2), 157–166. [for 2013]

    Golovatch, S.I. (2015a) Two new species of the millipede order Polydesmida from southern China (Diplopoda). Zoologicheskii zhurnal, 64 (9), 1023–1028.

    Golovatch, S.I. (2015b) Cave Diplopoda of southern China with reference to millipede diversity in Southeast Asia. ZooKeys, 510, 79–94.

    Golovatch, S.I. & Geoffroy, J.-J. (2014) On some new or poorly-known species of the millipede family Polydesmidae from southern China (Diplopoda: Polydesmida). Russian Entomological Journal, 23 (2), 91–105.

    Golovatch, S.I., Geoffroy, J.-J. & Mauriès, J.-P. (2007) Several new or poorly known cavernicolous millipedes (Diplopoda) from southern China. Arthropoda Selecta, 15 (2), 81–89. [for 2006]

    Golovatch, S.I., Liu, W.X., Li, Y.B. & Geoffroy, J.-J. (2012) One new and two little-known species of the millipede family Polydesmidae from southern China (Diplopoda: Polydesmida). Arthropoda Selecta, 21 (2), 131–136.

    Jiang, X.K., Guo, X. Chen, H.M. and Xie, Z.C. (2018) Four new species of the Glyphiulus javanicus group from southern China (Diplopoda, Spirostreptida, Cambalopsidae). ZooKeys, 741, 155–179.

    Jiang, X.K., Lv, J.C., Guo, X., Yu, Z.G. & Chen, H.M. (2017) Two new species of the millipede genus Glyphiulus Gervais, 1847 from Southwest China (Diplopoda: Spirostreptida: Cambalopsidae). Zootaxa, 4323 (2), 197–208.

    Likhitrakarn, N., Golovatch, S.I., Srisonchai, R. & Panha, S. (2017) A new species of Trichopeltis Pocock, 1894 from southern China, with a checklist and a distribution map of Trichopeltis species (Diplopoda, Polydesmida, Cryptodesmidae). ZooKeys, 725, 132–137.

    Liu, W.X., Golovatch, S.I. & Tian, M.Y. (2017b) Three new cavernicolous species of the millipede genus Trichopeltis Pocock, 1894 (Diplopoda: Polydesmida: Cryptodesmidae) from southern China. ZooKeys, 710, 1–14.

    Liu, W.X., Golovatch, S.I., Wesener, T. & Tian, M.Y. (2017a) Convergent Evolution of Unique Morphological Adaptations to a Subterranean Environment in Cave Millipedes (Diplopoda). PLOS ONE, 12 (2), e0170717.

    Miyosi, Y. (1956) Beiträge zur Kenntnis japanischer Myriopoden 15. Aufsatz: Über eine neue Gattung von Polydesmidae. Dobutsugaku Zasshi (Zoological Magazine), 65 (1–12), 10–12.