Skip to main content Skip to main navigation menu Skip to site footer
Type: Article
Published: 2018-07-10
Page range: 201–246
Abstract views: 193
PDF downloaded: 9

Catalogue of the Egyptian Ephydroidea (Diptera: Schizophora: Acalyptratae)

Entomology Department, Faculty of Science, Cairo University, Egypt.
Department of Biosystematics, Opole University, ul. Oleska 22, 45–052 Opole, Poland.
Classification Research Dept., Plant Protection Research Institute, Dokki, Egypt.
Diptera Ephydridae Drosophilidae Braulidae distribution Egyptian localities dates of collection new synonyms lectotype designations

Abstract

We present a catalogue of all known taxa of the superfamily Ephydroidea in Egypt. Old World synonymies, type localities, type depositories, world distributions by biogeographic realm(s) and country, Egyptian localities and dates of collection are provided. A total number of 117 species belonging to 45 genera, 18 tribes, 7 subfamilies, and representing 3 families has been catalogued. The treated families are: Ephydridae (shore flies or brine flies), Drosophilidae (vinegar flies or fruit flies) and Braulidae (bee lice). One shore fly species, Psilopa clara (Wollaston, 1858), is recorded for the first time from Egypt. Two new synonyms of shore flies are proposed, namely: Philotelma ulianai Raffone, 2011 = Ephydra (Ephydra) macellaria Egger, 1862 and Notiphila rufitarsis Macquart, 1851 = Allotrichoma biroi Cresson, 1929. Lectotypes are designated for three shore fly species, namely: Psilopa pectinata Hendel, 1931; Rhynchopsilopa nitidissima Hendel, 1931 and Notiphila rufitarsis Macquart, 1851.

 

References

  1. Bächli, G. (1982) On the type material of Palearctic species of Drosophilidae (Diptera). Beiträge zur Entomologie, 32, 289–301.

    Bächli, G., Burla, H. & Jungen, H. (1982) Drosophila subobscura in Egypt and its probable derivation. Genetica, 59, 3–7.
    https://doi.org/10.1007/BF00130809

    Becker, Th. (1896) Dipterologische Studien IV. Ephydridae. Berliner Entomologische Zeitschrift, 41 (2), 91–276.
    https://doi.org/10.1002/mmnd.48018960505

    Becker, Th. (1903) Ägyptische Dipteren gesammelt und beschrieben. Mitteilungen aus dem Zoologischen Museum in Berlin, 2 (3), 67–195.

    Becker, Th. (1907) Die Ergebnisse meiner dipterologischen Frühjahrsreise nach Algier und Tunis, 1906 [part 7]. Zeitschrift für Systematische Hymenopterologie und Dipterologie. Leipzig, 12, 369–407.

    Becker, T. (1908) Dipteren der Kanarischen Inseln. Mitteilungen aus dem Zoologischen Museum in Berlin, 4, 1–180.

    Becker, Th. (1922) Wissenschaftliche Ergebnisse der mit Unterstützung der Akademie der Wissenschaften in Wien aus der Erbschaft Treitl von F. Werner unternommenen zoologischen Expedition nach dem anglo-Ägyptischen Sudan (Kordofan) 1914. VI. Diptera. Denkschriften der Akademie der Wissenschaften in Wien Mathematisch-Naturwissenschaftliche Klasse, 98 (1923), 57–82.

    Becker, Th. (1926) 56a. Ephydridae und 56b. Canaceidae. In: Lindner, E. (Ed.), Die Fliegen der paläarktischen Region. Band VI. 1. E. Schweizerbart'sche Verlag, Stuttgart, pp. 1–115.

    Becker, Th. & Stein, P. (1912) Persische Dipteren von den Expeditionen des Herrn N. Zarudny 1898 und 1901. Annuaire du Musée Zoologique de l'Académie Impériale des Sciences de St. Pétersbourg, 18, 503–654.

    Bezzi, M. (1895) Eine neue Art der Dipterengattung Psilopa Fall. Wiener Entomologische Zeitung, 14 (4), 137–139.

    Bigot, J.M.F. (1885) Diptères nouveaux ou peu connus, 27 partie. XXXV: Famille dès Anomalocarerati (mihi). Annales de la Société Entomologique de France, 5, 15–246.

    Burla, H. & Bächli, G. (1978) Collecting Drosophila in Egypt. Bulletin of the European Drosophila Population Biology Group, 2, 7–8.

    Buzzati-Traverso, A. (1943) Morfologia, citologia e biologia di due nuove specie di Drosophila (Diptera acalyptera). Rendioconti / Istituto Lombardo di scienze e lettere, 77, 37–49.

    Canzoneri, S. (1987) Sugli Ephydridae e Canacidae del Sudan (Diptera, Cyclorrhapha). Bollettino del Museo civico di Storia naturale di Venezia, 37 (1986), 79–97.

    Canzoneri, S. (1988) Una nuova specie di Allotrichoma dell'Etiopia (Diptera, Ephydridae). Società Veneziana di Scienze Naturali, Lavori, 13, 3–4.

    Canzoneri, S. & Meneghini, D. (1969) Sugli Ephydridae e Canceidae della fauna etiopica. Bollettino del Museo civico di Storia naturale di Venezia, 19, 101–185.

    Canzoneri, S. & Meneghini, D. (1981) Su due specie di Ephydridae descritte da Becker (Diptera, Brachycera). Società Veneziana di Scienze Naturali, 6 (2), 89–90.

    Canzoneri, S. & Rampini, L. (1992) Una nuova specie di Glenanthe dell'Iran (Diptera, Ephydridae). Bollettino del Museo civico di Storia naturale di Venezia, 41 (1990), 231–233.

    Clausen, P.J. (1977) A revision of the Nearctic, Neotropical, and Palearctic species of the genus Ochthera including one Ethiopian species, and one new species from India. Transactions of the American Entomological Society, 103, 451–530.

    Clausen, P.J. (1979) Lectotype designation of Ochthera schembrii Rondani (Diptera: Ephydridae) and resulting synonymies. Entomological News, 90 (3), 141–142.

    Coe, R.L. (1943) Drosophila repleta Wollaston (Dipt., Drosophilidae) new to Britain, with notes on the species and some account of its breeding-habits. Entomologist's Monthly Magazine, 79, 204–206.

    Cogan, B.H. (1968) A revision of the Ethiopian species of the tribe Notiphilini (Diptera: Ephydridae). Bulletin of the British Museum (Natural History), Entomology, 21 (6), 281–365.

    Collin, J.E. (1930) Some new species of the dipterous genus Scatella Dsv. and the differentiation of Stictoscatella gen. nov. (Ephydridae). The Entomologist's Monthly Magazine, 66, 133–139.

    Collin, J.E. (1949) Results of the Armstrong College Expedition to Siwa Oasis (Libyan Desert), 1935, under the leadership of Prof. J. Omer-Cooper. Diptera Empididae, Dolichopodidae, Aschiza and Acalypterae. Bulletin de la Société Foudad ler d'Entomologie, 33, 175–25.

    Coquillett, D.W. (1900a) New genera and species of Ephydridae. The Canadian Entomologist, 32 (2), 33–36.
    https://doi.org/10.4039/Ent3233-2

    Coquillett, D.W. (1900b) Report on a collection of dipterous insects from Puerto Rico. Proceedings of the United States National Museum, 22, 249–270.
    https://doi.org/10.5479/si.00963801.22-1198.249

    Coquillett, D.W. (1901a) Three New Species of Diptera. Entomological News, 12, 16–18.

    Coquillett, D.W. (1901b) New Diptera from Southern Africa. Proceedings of the United States National Museum, 24, 27–32.

    Coquillett, D.W. (1910) The type-species of the North American genera of Diptera. Proceedings of the United States National Museum, 37, 499–647.
    https://doi.org/10.5479/si.00963801.37-1719.499

    Costa, A. (1846) Storia completa dell' Entomibia apum Costa (Nuovo genere d'insecli ditteri) et su i danni che arrcca alle api da miele (Istituto d'incoraggiamento Vol. VII). Atti del R. Istituto d'incoraggiamento alle scienze naturali di Napoli, 7, 291–306.

    Cresson, E.T. Jr. (1916) Studies in the American Ephydridae (Diptera). I. Revision of the species of the genus Paralimna. Transactions of the American Entomological Society, 42, 101–124.

    Cresson, E.T., Jr. (1924) Descriptions of new genera and species of the dipterous family Ephydridae. VI. Entomological News, 35 (5), 159–164.

    Cresson, E.T., Jr. (1925) Studies in the dipterous family Ephydridae, excluding the North and South American faunas. Transactions of the American Entomological Society, 51, 227–258.

    Cresson, E.T., Jr. (1929) Studies in the dipterous family Ephydridae. Paper II. Transactions of the American Entomological Society, 55, 165–195.

    Cresson, E.T. Jr. (1930) Studies in the dipterous family Ephydridae. Paper III. Transactions of the American Entomological Society, 56, 93–131.

    Cresson, E.T. Jr. (1930) Descriptions of new species of the dipterous family Ephydridae. Entomological News, 44, 65–70.

    Cresson, E.T. Jr. (1932) Studies in the dipterous family Ephydridae. Paper IV. Transactions of the American Entomological Society, 58, 1–34.

    Cresson, E.T. Jr. (1935) Descriptions of new genera and species of the dipterous family Ephydridae. Transactions of the American Entomological Society, 61, 345–372.

    Cresson, E.T. Jr. (1946) A systematic annotated arrangement of the genera and species of the Ethiopian Ephydridae (Diptera) I. The subfamily Psilopinae. Transactions of the American Entomological Society, 72, 241–264.

    Curtis, J. (1832) British Entomology; Being illustrations and descriptions of the genera of insects found in Great Britain and Ireland: containing coloured Figs. from nature of the most rare and beautiful species, and in many instances of the plants upon which they are found. E. Ellis and Company, Simpkin and Marshall, J. B. Bailliere, London, 44 pp.

    Czerny, L. (1909) Cyclorrhapha Schizophora Holometopa. Spanische Dipteren. III. Verhandlungen der kaiserlich-königlichen zoologisch-botanischen Gesellschaft in Wien, 59, 247–290.

    Deeming, J.C. (2002) Some species of the genus Hydrellia Robineau-Desvoidy (Diptera, Ephydridae) in Africa and Arabia, with special reference to species of economic importance. Mitteilungen aus dem Museum für Naturkunde in Berlin, Deutsche Entomologische Zeitschrift, 49 (2002), 47–59.
    https://doi.org/10.1002/mmnd.20020490105

    de Meijere, J.C.H. (1913) Praeda Itineris a L. F. de Beaufort in Archipelago indico facti annis 1909–1910. VI. Dipteren I. Bijdragen tot de Dierkunde, 19, 45–68.

    de Meijere, J.C.H. (1916) Studien über südostasiatische Dipteren XII. Javanische Dolichopodiden und Ephydriden. Tijdschrift voor Entomologie, 29, 225–273.

    Dikow, T. & Mathis, W.N. (2002) A revision and phylogenetic study of Actocetor Becker (Diptera: Ephydridae). Proceedings of the Entomological Society of Washington, 104 (2), 249–290.

    Dobzhansky, T. & Pavan, C. (1943) Studies on Brazilian species of Drosophila. Boletim da Faculdade de Filosofia, Ciências e Letras, Universidade de São Paulo, 36 (4), 7–72.

    Duda, O. (1921) Kritische Bemerkungen zur Gattung Scaptomyza Hardy (Dipteren). Jahresheft des Vereins für schlesische Insektenkunde zu Breslau, 13, 57–69.

    Duda, O. (1924) Beitrag zur Systematik der Drosophiliden unter besonderer Berücksichtigung der paläarktischen u. orientalischen Arten (Dipteren). Archiv für Naturgeschichte, 90A (3), 172–234.

    Duda, O. (1942) Neue oder ungenügend bekannte Zweiflügler der paläarktischen Region aus meiner Sammlung. 2. Fortsetzung. Deutsche Entomologische Zeitschrift, 1942 (1–4), 1–39.
    https://doi.org/10.1002/mmnd.194219420102

    Ebrahim, A.M., El-Hawagry, M.S., Negm, F.H. & El-Moursy, A.A. (2007) Key to tribes, genera, and species of the subfamily Gymnomyzinae (Ephydridae) from Egypt, with description of a new species. Proceedings of the Second International Conference of Economic Entomology. Entomological Society of Egypt, 1, 253–266.

    Egger, J. (1862) Dipterologische Beiträge. Fortsetzung der Beschreibungen neuer Dipteren. Verhandlungen der kaiserlich-königlichen Zoologisch-Botanischen Gesellschaft in Wien, 12, 777–784, 1233–1236.

    El-Hawagry, M. (2017) Catalogue of Egyptian Tephritoidea (Diptera: Schizophora: Acalyptratae). Zootaxa, 4299 (2), 151–190.
    https://doi.org/10.11646/zootaxa.4299.2.1

    El-Hawagry, M. & Gilbert, F. (2014) Zoogeographical affinities and faunal relationships of bee flies (Diptera: Bombyliidae) in Egypt. Zoology in the Middle East, 60 (1), 50–56.
    https://doi.org/10.1080/09397140.2014.892339

    Ellis, J.D. & Nalen, C.M.Z. (2008) Bee Louse, Bee Fly, Braulidae, Braula coeca Nitzsch (Insecta: Diptera: Braulidae). Available from: https://edis.ifas.ufl.edu/pdffiles/IN/IN84500.pdf (accessed 1 January 2018)

    El-Moursy, A.A., Negm, F.H., El-Hawagry, M.S. & Ebrahim, A.M. (2006) Review of the subfamily Ephydrinae (Diptera: Ephydridae) from Egypt, with a description of a new species. Bulletin of the Entomological Society of Egypt, 83, 71–97.

    Enderlein, G. (1934) Heringiinae, eine neue minierende Chloropiden Unterfamilie. Zoologischer Anzeiger, 105, 191–194.

    Fabricius, J.C. (1787) Mantissa insectorum sistens species nuper detectas adiectis synonymis, observationibus, descriptionibus, emendationibus. Tom. II. C.G. Proft, Hafniae (= Copenhagen), 382 pp.

    Fabricius, J.C. (1794) Entomologia systematica emendata et aucta. Secundum classes, ordines, genera, species adjectis synonimis, locis, observationibus, descriptionibus. Tom. IV. C.G. Proft, Hafniae [= Copenhagen], 472 pp.

    Fallén, C.F. (1810) Specimen entomologicum novam Diptera disponendi methodum exhibens. Berlingianis, Lundae (= Lund), 26 pp.

    Fallén, C.F. (1813) Beskrifning öfver några i Sverige funna Vattenflugor (Hydromyzides). Kongliga Vetenskaps-Academiens Handlingar, Series 3, 1813, 240–257.

    Fallén, C.F. (1820) Oscinides Sveciae. Quarum descriptionem Venia Ampl. Facult. Philos. Acad. Lund. Berlingianis, Lundae [= Lund], 12 pp.

    Fallén, C.F. (1823a) Hydromyzides Sveciae. Berlingianis, Lundae [= Lund], 12 pp.

    Fallén, C.F. (1823b) Geomyzides Sveciae. Berlingianis, Lundae, 8 pp.

    Fontdevila, A., Ruiz, A., Alonso, G. & Ocaña, J. (1981) The evolutionary history of Drosophila buzzati. I. Natural chromosomal polymorphism in colonized populations of the Old World. Evolution, 35, 148–157.

    Foote, B.A. (1995) Biology of shore flies. Annual Review of Entomology, 40, 417–442.
    https://doi.org/10.1146/annurev.en.40.010195.002221

    Frey, R. (1954) Diptera brachycera und Sciaridae von Tristan da Cunha. Results of the Norwegian Scientific Expedition to Tristan da Cunha, 1937–1938, 26, 1–55.

    Frey, R. (1958) Kanarische Diptera brachycera p.p., von Håkan Lindberg gesammelt. Societas Scientiarum Fennica. Commentationes Biologicae, 17 (4), 1–63.

    Frolova, S.L. & Astaurow, B.L. (1929) Die Chromosomengarnitur als systematisches Merkmal (eine vergleichende Untersuchung der russischen und amerikanischen Drosophila obscura Fall.). Zeitschrift für Zellforschung und Mikroskopische Anatomie, 10, 201–213.

    Ginsburg, I. (1940) Divergence and probability in taxonomy. Zoologica, New York, 25, 15–31.

    Giordani Soika, A.G. (1950) Vicarianze nella Fauna litoriparia del litorale veneto in rapporto alle caratteristiche del terreno. Studi sulle Olocenosi V. Bollettino del Museo civico di Storia naturale di Venezia, 5, 1–16.

    Giordani Soika, A.G. (1956) Viaggi di A. Giordani Soika nel Sahara IX. Contibuto allo studio del popolamento del Sahara Diptera Ephydridae. Bollettino del Museo civico di Storia naturale di Venezia, 9, 95–114.

    Godbole, N.N. & Vaidya, V.G. (1972) A new species of Drosophilidae from Poona (India): Zaprionus (Zaprionus) paravittiger sp. nov. Drosophila Information Service, 48, 135–136.

    Gordon, C. (1936) The frequency of heterozygosis in free-living populations of Drosophila melanogaster and Drosophila subobscura. Journal of Genetics, 33, 25–60.
    https://doi.org/10.1007/BF03027603

    Gupta, J.P. (1970) Description of a new species of Phorticella and Zaprionus. (Drosophilidae) from India. Proceedings of the Indian National Science. Academy, 36, 62–70.

    Hafez, M. (1947) Further additions to the dipterous fauna of dung in Egypt, with some biological observations. Bulletin de la Société Foudad I-er d'Entomologie, 31, 307–312.

    Haliday, A.H. (1833) Catalogue of Diptera Occurring about Holywood in Downshire. Entomological Magazine, 1, 147–180.

    Haliday, A.H. (1837) [New taxa]. In: Curtis, J. (Ed.), A Guide to an Arrangement of British Insects; Being a Catalogue of all the Named species Hitherto Discovered in Great Britain and Ireland. 2nd Edition, Pigot, Sherwood & Co., London, pp. 1–294.

    Haliday, A.H. (1839) Remarks on the generic distribution of the British Hydromyzidae (Diptera). Annals of Natural History, 3, 217–224, 401–411.
    https://doi.org/10.1080/03745483909443252

    Haliday, A.H. (1855) Recent works on the Diptera of northern Europe. The Natural History Review, 2, 59–64.

    Haliday, A.H. (1856) Addenda and corrigenda. In: Walker, F. (Ed.), Insecta Britannica, Diptera. Vol. 3. Reeve & Benham, London, pp. 342–346.

    Hardy, J. (1849) Note on Remedies for the Turnip-Fly amongst the Ancients, and on the Turnip-Fly of New Holland, with Notice of a New Genus and Species of Diptera. History of the Berwickshire Naturalist's Club, 2, 359–362.

    Harris, T.W. (1835) Insects. In: Hitchcock, E. (Ed.), Report on the geology, mineralogy, botany, and zoology of Massachusetts. 2nd Edition. J. S. & C. Adams, Amherst, pp. 553602.

    Hendel, F. (1913) H. Sauter's Formosa-Ausbeute. Acalyptrate Musciden (Dipt.) II. Supplementa Entomologica, 2, 77–112.

    Hendel, F. (1917) Beiträge zur Kenntnis der acalyptraten Musciden. Deutsche entomologische Zeitschrift, 1917, 33–47.

    Hendel, F. (1928) Über die minierenden europäischen Scaptomyza-Arten und ihre Biologie (Diptera). Zoologischer Anzeiger, 76, 289–302.

    Hendel, F. (1930) Die Ausbeute der deutschen Chaco-Expedition 1925/26. Diptera. XIX. Ephydridae. Konowia, 9 (2), 127–155.

    Hendel, F. (1931) Neue aegyptische Dipteren aus der Gruppe der acalyptraten Musciden, gesammelt von Prof. Efflatoun Bey. Bulletin de la Société Royale Entomologique d'Égypte, 15 (2), 59–73.

    Hollmann-Schirrmacher, V. (1998) Phylogeny of the subfamily Ilytheinae (Diptera, Ephydridae) with special reference to the genus Philygria. Studia Dipterologica, 5 (Supplement), 1–144.

    Hüttinger, E. (1980) Braula hansruttneri sp. n., eine neue Bienenlaus aus Osterreich (Braulidae, Diptera). Entomofauna, 1, 298–301.

    Hutton, F.W. (1901) Synopsis of the Diptera Brachycera of New Zealand. Transactions of the New Zealand Institute, 1900 (I), Zoology, 33, 1–95.

    Isa, A.L., Ismail, I.I. & Farrag, L.M. (1979) Certain ecological studies on the rice leafminer, Hydrellia prosternalis Deeming in Egypt (Diptera: Ephydridae). Agricultural Research Review, 57 (1), 53–62.

    Khishin, A.F. (1956) Drosophila in Egypt. Drosophila Information Service, 30, 124.

    Kirby, W. & Spence, W. (1815) An introduction to entomology or elements of the natural history of insects: with plates. Longman, Hurst, Rees, Orme, and Brown, London, 523pp.

    Knab, F. (1914) Drosophilidae with parasitic larvae. Insecutor Inscitiae Menstruus, 2 (1), 165–169.

    Krivosheina, M.G. (1989) Muchi-beregovushki roda Halmopota (Diptera, Ephydridae) Palearktiki [Palearctic Flies of the Genus Halmopota (Diptera, Ephydridae)], Vestnik Zoologii, 1989 (1), 18–24. [in Russian]

    Krivosheina, M.G. (1998) A revision of the shore-fly genus Notiphila Fallén of Palaearctic (Diptera Ephydridae). International Journal of Dipterology, 9 (1), 31–63.

    Krivosheina, M.G. & Zatwarnicki, T. (1998) Some notes on the Old World Allotrichoma (Diptera: Ephydridae) with description of three new species. Polskie Pismo Entomologiczne, 66, 291–310. [1997]

    Latreille, P.A. (1802) Histoire naturelle, générale et particulière des Crustacés et des Insectes. Tome troisième. In: Sonnini, C.S. (Ed.), Histoire naturelle par Buffon. F. Dufart, Paris , 468 pp

    Latreille, P.A. (1804) Tableau méthodique des insectes. Nouveau Dictionnaire d’Histoire Naturelle, 24, 129–200.

    Latreille, P.A. (1805) Histoire naturelle, générale et particulière, des Crustaces et des Insectes. Tome Quatorzième. Dufart, Paris, 432 pp.

    Latreille, P.A. (1810) Considérations générales sur l'ordre naturel des animaux composant les classes des crustacés, des arachnides, et des insectes; avec un tableau méthodique de leurs genres, disposés en familles. F. Schoell, Paris, 444 pp.

    Loew, H. (1847) Dipterologisches. Stettiner Entomologische Zeitung, 8 (12), 368–376.

    Loew, H. (1848) Ueber die Arten der Gattung Gymnopa. Stettiner Entomologische Zeitung, 9 (1), 13–15.

    Loew, H. (1856) Neue Beiträge zur Kenntnis der Dipteren. Fünfter Beitrag. Programm der Königlichen Realschule zu Meseritz, 1856, 1–56.

    Loew, H. (1858) Ueber Cacoxenus indagator nov. sp. und seine Verwandten. Wiener entomologische Monatschrift, 2, 213–222.

    Loew, H. (1860) Neue Beiträge zur Kenntniss der Dipteren. Siebenter Beitrag. Die Europaeischen Ephydrinidae und die bisher in Schlesien beobachteten Arten derselben. Programm der Königlichen Realschule zu Meseritz, 1860, 1–46.

    Loew, H. (1862) Monographs of the Diptera of North America. Part 1. Smithsonian Institution, Smithsonian Miscellaneous Collections, 6 (141), 1–221.
    https://doi.org/10.5962/bhl.title.57906

    Loew, H. (1864) Ueber zu Durren berg beobachtete halophile Dipteren. Zeitschrift für die Gesammten Naturwissenschaft, 23, 336–347.

    Loew, H. (1869) Ueber Dypteren der Augsburger Umgegend. Bericht des Naturhistorischen Vereins in Augsburg, 20, 39–59.

    Loew, H. (1878) Neue nordamerikanische Ephydrinen. Zeitschrift fur die Gesammten Natunvissenschaften, 51, 192–203.

    Macquart, M.J. (1835) Diptères. In: Histoire Naturelle des Insectes. In: Roret, N.E. (Ed.), Collection des suites à Buffon, Formant avec les oeuvres de cet auteur un cours complet d'histoire naturelle. Tome deuxième. Vol. 2. Pourrat Frères, Paris, pp. 1–703.

    Macquart, P.J.M. (1843) Diptères exotiques nouveaux ou peu connus. Tome deuxième, 3e partie. Roret, Paris, 460 pp.

    Macquart, J. (1844) Dipteres exotiques nouveaux ou peu connus. Memoires de la Société Royale des Sciences, de l’Agriculture et des Arts a Lille, 1840, 162–460.

    Macquart, J. (1851) Diptères exotiques nouveaux ou peu connus. Suite du 4e supplement publié dans les memoires de 1849. Mémoires de la Société Royale des Sciences, de l’Agriculture et des Arts de Lille, 1850, 134–294.

    Malloch, J.R. (1932) A new species of the genus Zaprionus, Coq. (Diptera, Drosophilidae). Stylops, 1, 10–11.
    https://doi.org/10.1111/j.1365-3113.1932.tb01328.x

    Malloch, J.R. (1933) Some Acalyptrate Diptera from the Marquesas Islands. Bulletin of the Bernice P. Bishop Museum, 114, 3–31.

    Mathis, W.N. (1978) New synonymy in Ephydridae (Diptera). Entomological News, 89 (2–3), 105–106.

    Mathis, W.N. (1979) Studies of Ephydrinae (Diptera: Ephydridae), II: Phylogeny, classification, and zoogeography of Nearctic Lamproscatella Hendel. Smithsonian Contributions to Zoology, 295, 1–41.
    https://doi.org/10.5479/si.00810282.295

    Mathis, W.N. (1982) Studies of Ephydrinae (Diptera: Ephydridae), VII: Revision of the genus Setacera Cresson. Smithsonian Contributions to Zoology, 350, 1–57.
    https://doi.org/10.5479/si.00810282.350

    Mathis, W.N. (1984a) A revision of the genus Asmeringa Becker (Diptera: Ephydridae). Israel Journal of Entomology, 17 (1983), 67–79.

    Mathis, W.N. (1984b) A revision of the shore-fly genus Homalometopus Becker (Diptera: Ephydridae). Proceedings of the Biological Society of Washington, 9 (2), 251–262.

    Mathis, W.N. (1986) A revision of the subgenus Eremotrichoma Soika of the shore-fly genus Allotrichoma Becker (Diptera, Ephydridae). Israel Journal of Entomology, 19, 127–139.

    Mathis, W.N. (1993) Studies of Gymnomyzinae (Diptera: Ephydridae), IV: A revision of the shore-fly genus Hecamede Haliday. Smithsonian Contributions to Zoology, 541, 1–46.
    https://doi.org/10.5479/si.00810282.541

    Mathis, W.N. & Deeming, J.C. (1987) A revision of the shore-fly genus Elephantinosoma Becker (Diptera: Ephydridae). Proceedings of the Entomological Society of Washington, 89 (3), 562–570.

    Mathis, W.N., Rodrigues, F. de A. & Couri, M. S. (2016) Family Ephydridae. Zootaxa, 4122 (1), 752–770.
    https://doi.org/10.11646/zootaxa.4122.1.64

    Mathis, W.N. & Zatwarnicki, T. (1990) A revision of the Western Palearctic species of Athyroglossa (Diptera: Ephydridae). Transactions of the American Entomological Society, 116 (1), 103–133.

    Mathis, W.N. & Zatwarnicki, T. (1993) A revision of the shore-fly genus Chlorichaeta Becker (Diptera: Ephydridae). Tijdschrift voor Entomologie, 136 (1), 55–76.

    Mathis, W.N. & Zatwarnicki, T. (1995) A world catalog of the shore flies (Diptera: Ephydridae). Memoirs on Entomology, International, 4, 1–423.

    Mathis, W.N. & Zatwarnicki, T. (2002) A phylogenetic study of the tribe Dryxini Zatwarnicki (Diptera: Ephydridae). Smithsonian Contributions to Zoology, 617, 1–101.

    Mathis, W.N. & Zatwarnicki, T. (2004) A review of the species of Hyadina Haliday (Diptera: Ephydridae) occurring in Israel. Israel Journal of Entomology, 34, 35–57.

    Mathis, W.N., Zatwarnicki, T. & Krivosheina, M.G. (1993) Studies of Gymnomyzinae (Diptera: Ephydridae), V: A revision of the shore-fly genus Mosillus Latreille. Smithsonian Contributions to Zoology, 548, 1–38.

    Mathis, W.N., Zatwarnicki, T, Stuke, J.-H. & Deeming, J.C. (2017) Order Diptera, family Ephydridae: A conspectus on shore-flies from the United Arab Emirates. Arthropod fauna of the UAE, 6, 636–761.

    Matsumura, S. (1915) Konchu-bunruigaku. Part 2. Keiseisha shoten, Tokyo, 316 pp.

    Megeed, M.S, Galal, O.A. & Abdel-Razek, M.M. (2015) Phylogenetic relationship of an invasive drosophilid, Zaprionus indianus and closely related species of Drosophilidae (Diptera) based on esterase patterns. Egyptian Journal of Genetics and Cytology, 44, 281–290.

    Meigen, J.W. (1803) Versuch einer neuen Gattungs-Eintheilung der europäischen zweiflügligen Insekten. Magazin für Insektenkunde, Illiger, 2, 259–281.

    Meigen, J.W. (1830) Systematische Beschreibung der bekannten europäischen zweiflügeligen Insekten. Sechster Theil. Schulz-Wundermann, Hamm, 401 pp.

    Meigen, J.W. (1838) Systematische Beschreibung der bekannten europäischen zweiflügeligen Insekten. Tome 7. Bei Friedrich Wilhelm Forstmann, Gedrukt bei Beaufort Sohn, Aachen, xii + 435 pp.

    Mercier, L. (1927) Discocerinella omonvillea genre nouveau, espèce nouvelle d'Ephydridae (Dipt.) de la région paléarctique. Bulletin and Annales de la Société Entomologique de Belgique, 67 (3–4), 123–125.

    Mik, J. (1886) Eine neue Drosophila aus Nieder-Oesterreich und den Aschanti-Ländern. (Ein dipterologischer Beitrag). Wiener entomologische Zeitung, 5, 328–331.
    https://doi.org/10.5962/bhl.part.20609

    Miyagi, I. (1966) Descriptions of three new species of Ephydridae, with one unrecorded species from Japan (Diptera, Ephydridae). Kontyu, 34 (2), 136–140.

    Mourad, A.M. & Mallah, G.S. (1960) Chromosomal polymorphism in Egyptian populations of Drosophila melanogaster. Evolution, 14, 166–170.
    https://doi.org/10.1111/j.1558-5646.1960.tb03075.x

    Ninomiya, E. (1929) On a new Species Brachydeutera ibari (Ephydridae). Oyo Dobutsugaku Zasshi, 1, 190–193.

    Nitzsch, C.L. (1818) Die Familien und Gattungen der Thierinsekten (insecta epizoica); als Prodromus einer Naturgeschichte derselben. Magazin der Entomologie, 3, 261–316.

    Oken, L. (1815) Okens Lehrbuch der Naturgeschichte, Dritter Theil, Zoologie, Erste Abtheilung Fleischlose Thiere. August Schmid, Jena, 850 pp.

    Oldenberg, L. (1914) Beitrag zur Kenntnis der europäischen Drosophiliden (Dipt.). Archiv fur Naturgeschichte, 80, 1–42.

    Oldenberg, L. (1923) Neue Acalyptraten (Dipt.) meiner Ausbeute. Deutsche Entomologische Zeitschrift, 1923, 307–319.
    https://doi.org/10.1002/mmnd.48019230314

    Olafsson, E. (1991) Taxonomic revision of western Palaearctic species of the genera Scatella R.-D. and Lamproscatella Hendel, and studies on their phylogenetic position within the subfamily Ephydrinae (Diptera: Ephydridae). Entomologica Scandinavica, 37 (Supplement), 1–100.

    d’Orbigny, C.V.D. (1845) Dictionnaire Universel d’Histoire Naturelle. Vol. 6. C. Renard, Paris, 792 pp.

    Örösi Pal, Z. (1939) Mehellensegek es a kopu allatvilaga. Orsz. Magyar Meheszeti Egyesulet, Budapest, 163 pp.

    Örösi Pal, Z. (1963) Pchelinyye brauly [Braula of bees]. Pchelovodstvo, 40 (6), 28–30. [in Russian]

    Örösi Pal, Z. (1966) A new bee louse: Braula coeca angulata. Journal of Apicultural Research, 5, 27–31.
    https://doi.org/10.1080/00218839.1966.11100128

    Ozerov, A.L. (1984) New Species of the Genus Risa (Diptera, Risidae) from Middle Asia. Zoologicheskii Zhurnal, 6, 945–948. [in Russian with an English title and abstract]

    Panzer, G.W.F. (1794) Favnae insectorvm germanicae initia oder Devtschlands Insecten. Heft 17. Felsecker, Nürnberg, 24 pp.

    Pape, T., Blagoderov, V. & Mostovski, M.B. (2011) Order Diptera Linnaeus, 1758. In: Zhang, Z.Q. (Ed.), Animal biodiversity: An outline of higher–level classification and survey of taxonomic richness. Zootaxa, 3148, 222–229.

    Papp, L. (1977) Notes on Some Becker's Types (Diptera, Carnidae and Risidae fam. n.). Annales Historico-Naturales Musei Nationalis Hungarici, 69, 185–189.

    Papp, L. (1979) A contribution to the revision of the Palaearctic Ephydridae (Diptera). Folia Entomologica Hungarica, New Series, 32 (1), 97–104.

    Papp, L. (1984) Family Braulidae. In: Soós, A. & Papp, L. (Eds.), Catalogue of Palaearctic Diptera. Vol. 10. Elsevier, Amsterdam, Oxford, New York & Tokyo, pp. 178–180.

    Patterson, J.T. & Wheeler, M.R. (1942) Description of new species of the subgenera Hirtodrosophila and Drosophila. The University of Texas Publication, 4213, 67–109.

    Pavan, C. (1952) Relações entre populações naturais de Drosophila e o meio ambiente. Universidade de Sao Paulo, Sao Paulo, 107 pp. [PhD Thesis]

    Raffone, G. (2011) Philotelma ulianai sp. nov. from Egypt (Insecta, Diptera, Ephydridae). Societa Veneziana di Scienze Naturali Lavori, 36, 15–17.

    Robineau-Desvoidy, J.B. (1830) Essai sur les Myodaires. Mémoires Preséntes par divers Savans à l'Académie Royale des Sciences de l'Institut de France, et Imprimés par son Ordre Sciences Mathématiques et Physiques, 2 (2), 1–813.

    Rocha Pité, M.T. & Tsacas, L. (1980) Catalogue of the European species of the genus Drosophila. Boletim da Sociedade Portuguesa de Ciencias Naturais, 19, 37–47.

    Rondani, C. (1847) Communications: Nota septima ad inserviendum dipterologiae italicae. Nova species generis Ochtherae Latr., descripta a Camillo Rondani. Annales de la Société Entomologique de France, Series 2, 5, xxix–xxxi.

    Rondani, C. (1856) Dipterologiae Italicae prodromus. Vol. I. Genera Italica ordinis dipterorum ordinatim disposita et distincta et in familias et stirpes aggregata. A. Stocchi, Parmae [= Parma], 226 pp.
    https://doi.org/10.5962/bhl.title.8160

    Rondani, C. (1875) Notizie sul moscherino dell'uva. Bollettino del Comizio Agrario Parmense, 8, 145–149.

    Sawaby, R.F., El Hamouly, H. & Abo-El Ela, R.H. (2018) Diagnosis and keys of the main dipterous families and species collected from rabbit and guinea pig carcasses in Cairo, Egypt, The Journal of Basic and Applied Zoology, 79 (10), 1–14.
    https://doi.org/10.1186/s41936-018-0018-6

    Schmitz, H.J., Valente, V.L.S. & Hofmann, P.R.P. (2007) Taxonomic Survey of Drosophilidae (Diptera) from Mangrove Forests of Santa Catarina Island, Southern Brazil. Neotropical Entomology, 36, 53–64.
    https://doi.org/10.1590/S1519-566X2007000100007

    Séguy, E. (1933) Mission Saharienne Augiéras-Draper 1927–1928. Insectes Diptères. Bulletin du Muséum national d'Histoire naturelle, Paris, 5, 122–127

    Séguy, E. (1951) Description d'un nouveau genre d'Ephydrine (Dipt. Ephydridae). Bollettino della Società Entomologica Italiana, 81 (1), 4–7.

    Sherif, M.R., Abdallah, F.E. & Soliman, A.M. (1999) Major insects of rice plants inEgypt and their management. Advances in Agricultural Research in Egypt, 2 (3), 188–219.

    Sherif, M.R., Hendawy, A.S. & El-Habashy, M.M. (2005) Management of rice insect pests. Egyptian Journal of Agricultural Research, 83 (5A), 111–130.

    Sherif, M.R., Khodair, I. & El-Habashy, M. (1997) Cultural practices to manage the rice leaf miner, Hydrellia prosternalis (Diptera: Ephydridae) in Egypt. Egyptian Journal of Agricultural Research, 75 (3), 611–622.

    Soliman, A.M., Abd-El-Basir, S. & Abou-El-Ela, R.G. (1997) Varietal resistance: field evaluation and screening of certain rice materials against the rice leaf miner, Hydrellia prosternalis Deem. and role of some factors in plant relative resistance. Egyptian Journal of Agricultural Research, 75 (1), 121–136.

    Stenhammar, C. (1844) Försök till Gruppering och Revision af de Svenska Ephydrinae. Kongliga Vetenskaps-Akademiens Handlingar, Series 3, 1843, 75–272.

    Steyskal, G.C. (1968) Notes and descriptions of Egyptian Acalyptrate Diptera (Ephydridae, Milichiidae, Otitidae). Bulletin de la Société Entomologique d'Egypte, 50, 109–125.

    Steyskal, G.C. & El-Bialy, S. (1967) A list of Egyptian Diptera with a bibliography and key to families. Ministry of Agriculture Technical Bulletin, 3, 12–18.

    Strobl, G. (1906) Spanische Dipteren. II. Beitrag. Memorias de la Real Sociedad Española de Historia Natural, 3 (5), 271–422.

    Stuke, J.-H. (2012a) Neue Uferfliegen-Funde (Diptera: Ephydridae) vom Madeira-Archipel, mit der Beschreibung einer neuen Art. Studia Dipterologica, 17 (2010), 211–221.

    Stuke, J.-H. (2012b) Funde von Ephydridae (Diptera) aus Jordanien, mit der Beschreibung von fünf neuen Arten [Records of Ephydridae (Diptera) from Jordan, with the description of five new species]. Studia dipterologica, 18 (2011), 197–224.

    Sturtevant, A.H. (1919) A new species closely resembling Drosophila melanogaster. Psyche, 26, 153–155.
    https://doi.org/10.1155/1919/97402

    Sturtevant, A.H. (1921) The North American Species of Drosophila. Carnegie Institute of Washington Publication, 301, 1–150.
    https://doi.org/10.5962/bhl.title.33564

    Sturtevant, A.H. (1939) On the subdivision of the genus Drosophila. Proceedings of the National Academy of Sciences of the United States, 25, 137–141.
    https://doi.org/10.1073/pnas.25.3.137

    Tsacas, L. (1980) L’identité de Zaprionus vittiger Coquillett et revision des espéces afrotropicales affines (Dipt. Drosophilidae). Bulletin de la Société entomologique de France, 85, 141–154.

    van der Wulp, F.M. (1891) Eenige uitlandsche Diptera. Tijdschrift voor entomologie, 34, 193–218.

    van der Wulp, F.M. (1897) Zur Dipteren-Fauna von Ceylon. Természetrajzi Füzetek, 20, 136–144.

    Vilela, C.R. (1983) A revision of the Drosophila repleta species group (Diptera, Drosophilidae). Revista Brasileira de Entomologia, 27, 1–114.

    Villeneuve, J. (1911) Description de deux nouveaux Diptères. Wiener entomologische Zeitung, 30, 81–84.

    Walker, F. (1849) List of the specimens of dipterous insects of the collection of the British Museum. Part 4. British Museum (Natural History), London, 484 pp.

    Walsh, D.B., Bolda, M.P., Goodhue, R.E., Dreves, A.J., Lee, J., Bruck, D.J., Walton, V.M., O'Neal, S.D. & Zalom, F.G. (2011) Drosophila suzukii (Diptera: Drosophilidae): Invasive pest of ripening soft fruit expanding its geographic range and damage potential. Journal of Integrated Pest Management, 2 (1), 1–7.
    https://doi.org/10.1603/IPM10010

    Waltl, J. (1837) Neue Gattungen von Mucken bey Passau. Isis (Oken's), 4, 279–283.

    Westwood, J.O. (1840) Order XIII. Diptera Aristotle (Antilata Fabricius. Halteriptera Clairv.). In: An Introduction to the Modern Classification of Insects; Founded on the Natural Habits and Corresponding Organisation of the Different Families. Synopsis of the genera of British Insects. Orme, Brown, Green and Longmans, London, pp. 125–154.

    Wiedemann, C.R.W. (1824) Munus rectoris in Academia Christiana Albertina aditurus Analecta entomologica ex Museo Regio Havniensi maxime congesta profert iconibusque illustrat. Eregio typoguapheo scholarum, Kiliae [= Kiel], 60 pp.
    https://doi.org/10.5962/bhl.title.77322

    Wiedemann, C.R.W. (1830) Aussereuropäische zweiflügelige Insekten. Zweiter Theil. Schulz, Hamm, 684 pp.

    Willcocks, F.C. (1916) Miscellaneous insect notes, Metamorphose of Ephydra macellaria Egger. Bulletin de la Société Entomologique d'Egypte, 9, 100–105.

    Wirth, W.W. (1955) East African Ephydridae (Ins., Diptera). Mitteilungen ausdem Zoologischen Museum in Berlin, 31, 48–58.
    https://doi.org/10.1163/187631275X00325

    Wirth, W.W. (1975) A revision of the brine flies of the genus Ephydra of the Old World (Diptera: Ephydridae). Entomologica Scandinavica, 6 (1), 11–44.

    Wollaston, T.V. (1858) Brief diagnostic characters of undescribed Madeiran insects. Annals and Magazine of Natural History, 1, 18–28.
    https://doi.org/10.1080/00222935808696865

    Yassin, A. & Abou-Youssef, A.Y. (2004) A new front for a global invasive drosophilid: the colonization of the Northern-Western desert of Egypt by Zaprionus indianus Gupta, 1970. Drosophila Information Service, 87, 67–68.

    Yassin, A., Borai, F., Capy, P., David, J.R., Elias, E., Riad, S.A., Shalaby, H.G., Serour, S. & Abou-Youssef, A.Y. (2009) Evolutionary Genetics of Zaprionus. II. Mitochondrial DNA and chromosomal variation of the invasive drosophilid Zaprionus indianus in Egypt. Mitochondrial DNA, 20, 34–40.
    https://doi.org/10.1080/19401730902890042

    Zatwarnicki, T. (1987) New synonyms and records of Palearctic Scatophila (Diptera, Ephydridae). Polskie Pismo Entomologiczne, 57 (2), 277–298.

    Zatwarnicki, T. (1991) Changes in nomenclature and synonymies of some genera and species of Ephydridae (Diptera). Deutsche Entomologische Zeitschrift, 38 (4–5), 295–333.
    https://doi.org/10.1002/mmnd.19910380403

    Zatwarnicki, T., Cielniak, M. & Pochrząst, K. (2016) A revised phylogeny of the tribe Discocerinini (Diptera: Ephydridae) with an emphasis on structures of the proboscis. Annales Zoologici, 66 (1), 1–34.
    https://doi.org/10.3161/00034541ANZ2016.66.1.001

    Zatwarnicki, T. & Irwin, A. (2018) Taxonomic notes on the genera Scatella and Scatophila (Diptera: Ephydridae) with a remark on Trixoscelis chilensis (Trixoscelididae). Zootaxa, 4377 (1), 91–109.
    https://doi.org/10.11646/zootaxa.4377.1.6

    Zatwarnicki, T. & Mathis, W.N. (2007) A revision of the Palearctic species of the shore-fly genus Discomyza Meigen (Diptera: Ephydridae). Insect Systematics and Evolution, 38, 241–266.
    https://doi.org/10.1163/187631207788754448

    Zatwarnicki, T. & Mathis, W.N. (2012) Heterogeneity in shore-flies—the case of Glenanthe Haliday (Diptera: Ephydridae) in the Old World. Annales de la Société Entomologique de France, New Series, 47 (3–4), 418–443.

    Zetterstedt, J.W. (1838) Insecta Lapponica. Editora, Lipzia (= Leipzig), 1139 pp.

    Zetterstedt, J.W. (1847) Diptera scandinaviae disposita et descripta. Tomus Sextus. Lundberg, Lund, 418 pp. [pp. 2163–2580]