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Type: Article
Published: 2018-06-22
Page range: 443–470
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Cicada fossils (Cicadoidea: Tettigarctidae and Cicadidae) with a review of the named fossilised Cicadidae

Australian Museum Research Institute, 1 William Street, Sydney N.S.W. 2010.
Hemiptera Eocene Cretaceous Jurassic Miocene Oligocene Paleocene Quaternary Pleistocene Pliocene Tertiary

Abstract

The Cicadoidea comprise two families, the Cicadidae and the Tettigarctidae. This paper evaluates the status and taxonomy of all named Cicadoidea fossils belonging to the Cicadidae. Shcherbakov (2009) has previously revised the Tettigarctidae. Two new genera are described, Camuracicada gen. n. and Paleopsalta gen. n., for Camuracicada aichhorni (Heer, 1853) comb. n. and Paleopsalta ungeri (Heer, 1853) comb. n. A lectotype is designated for Cicada emathion Heer, 1853.

          Cicada grandiosa Scudder, 1892 is transferred to Hadoa Moulds, 2015 as Hadoa grandiosa comb. n.; Oncotympana lapidescens J. Zhang, 1989 is transferred to Hyalessa China, 1925 as Hyalessa lapidescens comb. n.; Meimuna incasa J. Zhang, Sun & X. Zhang, 1994 and Meimuna miocenica J. Zhang & X. Zhang, 1990 are transferred to Cryptotympana Stål, 1861 as Cryptotympana incasa comb. n. and Cryptotympana miocenica comb. n.; Tibicen sp. aff. japonicus Kato, 1925 is transferred to Auritibicen as Auritibicen sp. aff. japonicus comb. n., and Terpnosia sp. aff. vacua Olivier, 1790 is transferred to Yezoterpnosia Matsumura, 1917 as Yezoterpnosia sp. aff. vacua comb. n. The generic placement of two other fossils is changed to reflect current classification, those species now being Auritibicen bihamatus (Motschulsky, 1861) and Yezoterpnosia nigricosta (Motschulsky, 1866).

         Two species, Davispia bearcreekensis Cooper, 1941 and Lithocicada perita Cockerell, 1906, are transferred from the subfamily Cicadinae to the Tibicininae, tribe Tibicinini. Cicadatra serresi (Meunier, 1915) is also transferred from the Cicadinae to the Cicadettinae because the Cicadatrini have recently been transferred from the Cicadinae to the Cicadettinae (Marshall et al. 2018).

         Miocenoprasia grasseti Boulard and Riou, 1999 is transferred from the tribe Prasiini to the Lamotialnini. Tymocicada gorbunovi Becker-Migdisova, 1954 is transferred from the Dundubiini to the Cryptotympanini; Paracicadetta oligocenica Boulard & Nel, 1990 is transferred from the Cicadettini to the Pagiphorini and Minyscapheus dominicanus Poinar et al., 2011 is assigned to the Taphurini. Names of species once considered to belong in Cicadidae, but now excluded, are listed with explanation.

 

References

  1. Beattie, R.G. & Avery, S. (2012) Palaeoecology and palaeoenvironment of the Jurassic Talbragar Fossil Fish Bed, Gulgong, New South Wales, Australia. Alcheringa, 36, 451–465.
    https://doi.org/10.1080/03115518.2012.671675

    Becker-Migdisova, E.E. (1954) Nakhodka yuzhnoi tsikady v miotsene Zapadnoi Sibiri [The discovery of a southern cicada in the Miocene of Western Siberia]. Voprosy Geologii Azii [Geological Problems of Asia], 1, 799–805, 1 pl. [in Russian]

    Bode, A. (1953) Die Insektenfauna des ostneidersachsischen oberen Lias. Paleontographica, Stuttgart, Series A, 103, 1–375, pls. 1–15.

    Boulard, M. & Nel, A. (1990) Sur deux cigales fossiles des terrains tertiaires de la France (Homoptera, Cicadoidea). Revue Française d'Entomologie, New Series, 12, 37–45.

    Boulard, M. & Riou, B. (1989) Tibicina gigantea n. sp. cigale fossile de la Montagne d'Andance (Homoptera, Tibicinidae). Nouvelle Revue d'Entomologie, New Series, 5 (4), 349–351. [published 28 February,1989, not 1988 as sometimes quoted]

    Boulard, M. & Riou, B. (1999) Miocenoprasia grasseti n. g., n. sp., grande cigale fossile du Miocene de la Montagne d'Andance (Cicadomorpha Cicadidae Tibicininae). Ecole pratique des hautes Etudes, Biologie et Evolution des Insectes, 11/12, 135–140.

    Brodie, P.B. (1845) A history of the fossil insects in the Secondary rocks of England. accompanied by a particular account of the strata in which they occur, and of the circumstances connected with their preservation. van Voorst, London, 130 pp., 11 pls.

    Brodie, P.B. (1853) On the occurrence of the remains of insects in the Tertiary clay of Dorsetshire. Proceedings of the Geological Society of London, 9, 53–54.
    https://doi.org/10.1144/GSL.JGS.1853.009.01-02.17

    Buckton, G.B. (1890–91) Monograph of the British Cicadidae or Tettigidae. 2 Volumes published in 7 parts. Macmillan, London and New York, 312 pp.

    Carpenter, F.M. (1992) Superclass Hexapoda. In: Kaesler, R.L. (Ed.), Treatise on Invertebrate Paleontology. Part R. Arthropoda 4. Vol. 3. The Geological Society of America and the University of Kansas, Boulder, pp. i–xxii + 1–655.

    Cockerell, T.D.A. (1906) A fossil cicada from Florissant, Colorado. Bulletin of the American Museum of Natural History, 22, 457–458.

    Cockerell, T.D.A. (1908) Descriptions of Tertiary insects; (2) a cicada from Florissant, Colorado. American Journal of Science, New Haven, Series 4, 25, 51–52.
    https://doi.org/10.2475/ajs.s4-25.145.51

    Cockerell, T.D.A. (1915) A visit to Oeningen. The Scientific Monthly, 1 (3), 287–291.

    Cohen, K.M., Finney, S.C., Gibbard, P.L. & Fan, J.-X. (2013) The ICS International Chronostratigraphic Chart. Episodes, 36, 199–204. Updated Version 2017/02. Available from: http://www.stratigraphy.org/ (accessed 18 March 2018)

    Cooper, K.W. (1941) Davispia bearcreekensis Cooper, a new cicada from the Paleocene, with a brief review of the fossil Cicadidae. American Journal of Science, 239, 286–304.
    https://doi.org/10.2475/ajs.239.4.286

    Etheridge, R. & Olliff, A.S. (1890) The Mesozoic and Tertiary insects of New South Wales. Memoirs Geological Survey of New South Wales, Palaeontology, 7, i–xi + 1–15, pls. 1–2.

    Evanoff, E., McIntosh, W.C. & Murphey, P.C. (2001) Stratigraphic summary and 40Ar/39Ar geochronology of the Florissant Formation, Colorado. Proceedings of the Denver Museum of Natural Science, 4, 1–16.
    https://doi.org/10.1071/ZO9560165

    Evans, J.W. (1956) Palaeozoic and Mesozoic Hemiptera (Insecta). Australian Journal of Zoology, 4, 165–258.

    Fossilworks (2018) Gateway to the Paleobiology Database. Available from: http://fossilworks.org/ (accessed 18 March 2018)

    Fujiyama, I. (1969) A Miocene cicada from Nasu, with an additional record of a Pleistocene cicada from Shiobara, Japan (Tertiary Insect Fauna of Japan, 3). Bulletin of the National Science Museum, Tokyo, 12, 863–73.

    Fujiyama, I. (1979) Some late Cenozoic cicadas from Japan. Bulletin of the National Science Museum of Tokyo, Series A (Zoology), 5, 139–152.

    Fujiyama, I. (1982) Some fossil cicadas from Neogene of Japan. Bulletin of the National Science Museum Series C (Geology and Paleontology), 8, 181–187.

    Giebel, C.G.A. (1856) Fauna der Vorwelt, mit steter Berücksichtigung der lebenden Thiere. Monographisch dargestelit. Vol. 2. Insekten und Spinnen der Vorwelt mit Steter Berücksichtigung der lebenden Insekten und Spinnen, monographisch dargestellt. F.U. Brodhaus, Leipzig, xviii + 511 pp.

    Grimaldi, D. & Engel, M.S. (2005) Evolution of the insects. Cambridge University Press, New York, xv + 755 pp.

    Handlirsch, A. (1906–1908) Die Fossilen Insekten und die Phylogenie der rezenten Formen. Ein Handbuch fúr Paläontologen und Zoologen. W. Engelmann, Leipzig, ix + 1430 pp., 51 pls.

    Handlirsch, A. (1925) Palaeontologie. In: Schröder, C. (Ed.), Handbuch der entomologie. Vol. 3. Geschichte, literature, technic, paläontologie, phylogenie, systematic. Gustav Fischer, Jena, pp. 1–1201.

    Heer, O. (1853) Die Insektenfauna der Tertiargebilde von Oeningen und von Radoboj in Croatien. Neue Denkschriften der Allgemeinen Schweizerischen Gesellschaft für die Gesammten Naturwissenschaften, Zürich, 13 (4), 1–138, pls. 1–15.

    Hill, K.B.R., Marshall, D.C., Moulds, M.S. & Simon C. (2015) Molecular phylogenetics, diversification, and systematics of Tibicen Latreille 1825 and allied cicadas of the tribe Cryptotympanini, with three new genera and emphasis on species from the USA and Canada (Hemiptera: Auchenorrhyncha: Cicadidae). Zootaxa, 3985 (2), 219–251.
    https://doi.org/10.11646/zootaxa.3985.2.3

    Jell, P.A. (2004) The fossil insects of Australia. Memoirs of the Queensland Museum, 50 (1), 1–124.

    Kato, M. (1925) Japanese Cicadidae, with descriptions of new species. Transactions of the Formosa Natural History Society, 15, 1–47, 1 pl. [in Japanese]

    Kinugasa, Y. & Miyataki, Y. (1976) A neogene cicada from Tatsumi-toge, Tottori Pref., Japan. Bulletin of the Osaka Museum of Natural History, 30, 5–10, pls. 1–2.

    Kinugasa, Y. & Miyataki, Y. (1979) The second Neogene cicada from Tatsumi-tôge, Tottori Prefecture, Japan (Homoptera: Cicadidae). Bulletin of the Osaka Museum of Natural History, 32, 1–6, pls. 1–2.

    Krause, J.M., Bown, T.M., Bellosi, E.S. & Genise, J.F. (2008) Trace fossils of cicadas in the Cenozoic of Central Patagonia, Argentina. Paleontology, 51 (2), 405–418.
    https://doi.org/10.1111/j.1475-4983.2008.00753.x

    Lameere, A. & Severin, G. (1897) Les insects de Bernissart. Annales de la Societe Entomologique de Belgique, 41, 35–38.

    Lee, Y.J. (2009) Taxonomic notes on the genus Haphsa (Hemiptera: Cicadidae) with descriptions of two new species. Florida Entomologist, 92 (2), 330–337.
    https://doi.org/10.1653/024.092.0218

    Lee, Y.J. (2012) Resurrection of the genus Yezoterpnosia Matsumura (Hemiptera: Cicadidae: Cicadini) based on a new definition of the genus Terpnosia Distant. Journal of Asia-Pacific Entomology, 15, 255–258.
    https://doi.org/10.1016/j.aspen.2011.12.006

    Lee, Y.J. (2015) Description of a new genus, Auritibicen gen. nov., of Cryptotympanini (Hemiptera: Cicadidae) with redescriptions of Auritibicen pekinensis (Haupt, 1924) comb. nov. and Auritibicen slocumi (Chen, 1943) comb. nov. from China and a key to the species of Auritibicen. Zootaxa, 3980 (2), 241–254.
    https://doi.org/10.11646/zootaxa.3980.2.5

    Lee, Y.J. & Emery, D. (2013) New genera and species of Leptopsaltriini (Hemiptera: Cicadidae: Cicadinae) from India and Tibet, with the descriptions of five new subtribes. Zoosystema, 35 (4), 525–535.
    https://doi.org/10.5252/z2013n4a6

    Marshall, D.C. (2015) Comments on Tibicina Amyot, 1847 and Lyristes Horvath, 1926 (Insecta, Hemiptera, Homoptera): proposed conservation by the suppression of Tibicen Berthold, 1827 [?Latreille, 1825], and concerning the type species of Cicada Linnaeus, 1758 (Case 239; see BZN 41: 163–184; 71: 103–131; 179–180. (1). Bulletin of Zoological Nomenclature, 72, 217–219.

    Marshall, D.C., Moulds, M.S.. Hill, K.B.R., Price, B.W., Wade, E.J., Owen, C.L., Goemans, G., Marathe, K., Sarkar, V., Cooley, J.R., Sanborn, A.F., Kunte, K., Villet, M.H. & Simon, C. (2018) A molecular phylogeny of the cicadas (Hemiptera: Cicadidae) with a review of tribe and subfamily classification. Zootaxa, 4424 (1), 1–64.

    https://doi.org/10.11646/zootaxa.4424.1.1

    Matsumura, S. (1917) A list of the Japanese and Formosan Cicadidae, with descriptions of new species and genera. Transactions of the Sapporo Natural History Society, 6, 186–212.

    Martynov, A.V. (1937) Liassovye nasekomye Shuraba i Kizil Kii [Liassic insects from Shurab and Kisyl-Kiya], Part I, Various orders except Blattodea and Coleoptera. Akademiya Nauk SSSR, Trudy Paleontologicheskogo Instituta, 7 (1), 1–180. [in Russian. Part 2 comprises pp. 181–232.]

    Meischner, D. (2000). Der pliozäne Teich von Willershausen am Harz. In: Pinna, G. & Meischner, D. (Eds.), Europäische Fossillagerstätten. Springer-Berlin, Heidelberg, New York, pp. 223–228, 261.
    https://doi.org/10.1007/978-3-642-57198-5_25

    Metcalf, Z. P. & Wade, V. (1963) General catalogue of the Homoptera. In: A catalogue of the fossil Homoptera (Homoptera: Auchenorhyncha). Supplement to Fascicle 1. University of North Carolina State College, Raleigh, pp. i–v, 1–245.

    Meunier, F. (1915) Nouvelles recherches sur quelques insectes des plattrières d'Aix-en-Provence. Verhandelingen van de Koninklijke Nederlandse Akademie van Wetenschappen te Amsterdam, Series 2, 18 (5), 1–17, pls. 1–15.

    Motschulsky de, V.I. (1861) Insectes du Japon [continuation]. Etudes Entomologiques, 10, 4–24.

    Motschulsky de, V.I. (1866) Catalogue des insectes reçus du Japon. Bulletin de la Société Impériale des Naturalistes de Moscou, Biologique, 39, 163–200.

    Moulds, M.S. (2005) An appraisal of the higher classification of cicadas (Hemiptera: Cicadoidea) with special reference to the Australian fauna. Records of the Australian Museum, 57, 375–446.
    https://doi.org/10.3853/j.0067-1975.57.2005.1447

    Moulds, M.S. (2012) A review of the genera of Australian cicadas (Hemiptera: Cicadoidea). Zootaxa, 3287, 1–262.

    Nel, A. (1996) Un Tettigarctidae fossile du Lias européen (Cicadomorpha, Cicadoidea Tettigarctidae). Ecole pratique des hautes Etudes, Biologie et Evolution des Insectes, 9, 83–94.

    Olivier, G.A. (1790) Cigale, Cicada. In: Encyclopedie methodique histoire naturelle, Insectes, 5, pp.735–760.

    Piton, L.E. (1940) Paléontologie du Gisement Eocéne de Menat (Puy-de-Dôme) (Flore et Faune). Memoires de la Societe d'Histoire Naturelle d'Auvergne, 1, 1–303, pls. 1–26.

    Poinar, G. & Kritsky, G. (2011) Morphological conservatism in the foreleg structure of cicada hatchlings, Burmacicada protera n. gen., n. sp. in Burmese amber, Dominicicada youngi n. gen., n. sp. in Dominican amber and the extant Magicicada septendecim (L.) (Hemiptera: Cicadidae). Historical Biology, 24, 461–466.
    https://doi.org/10.1080/08912963.2011.603421

    Poinar, G., Kritsky, G. & Brown, A. (2011) Minyscapheus dominicanus n. gen., n. sp. (Hemiptera: Cicadidae), a fossil cicada in Dominican amber. Historical Biology, 24, 329–333.
    https://doi.org/10.1080/08912963.2011.635791

    Prokop, J. & Boulard, M. (2000) Tibicina sakalai n. sp., cigale fossile du Miocene inferieur de Tchecoslovaquie (Auchenorhyncha [Auchenorrhyncha], Cicadidae, Tibicininae).[Tibicina sakalai n. sp., fossil cicada of the Lower Miocene of Czechoslovakia (Auchenorrhyncha, Cicadidae, Tibicininae)]. Ecole Pratique des Hautes Etudes, Travaux du Laboratoire Biologie et Evolution des Insectes, 13, 127–131.

    Prokop, J. & Fikáček, M. (2007) Early Oligocene insect fauna from Seifhennersdorf (Saxony, Germany). Acta Musei Nationalis Pragae, Series B, 63 (2–4), 205–213.

    Puissant, S., Boulard, M., Lee, Y.J., Hayashi, M., Wei, C. & Sueur, J. (2015) Comments on Tibicina Amyot, 1847 and Lyristes Horvath, 1926 (Insecta, Hemiptera, Homoptera): proposed conservation by the suppression of Tibicen Berthold, 1827 [?Latreille, 1825], and concerning the type species of Cicada Linnaeus, 1758 (Case 239; see BZN 41: 163–184; 71: 103–131; 179–180. (2). Bulletin of Zoological Nomenclature, 72, 219–220.

    Qiu, Z.-X., Qiu, Z.-D., Deng, T., Li, C.-K., Zhang, Z.-Q., Wang, B.-Y. & Wang, X. (2013) Neogene land mammal stages/ages of China: toward the goal to establish an Asian land mammal stage/age scheme. In: Wang, X., Flynn, L.J. & Fortelius, M. (Eds.), Fossil mammals of Asia neogene biostratigraphy and chronology. Columbia University Press, New York, pp. 29–90.
    https://doi.org/10.7312/columbia/9780231150125.003.0001

    Rasnitsyn, A.P. & Zherikhin, V.V. (2002) Appendix: alphabetical list of selected insect fossil sites. Impression fossils. In: Rasnitsyn, A.P. & Quick, D.L.J. (Eds.), History of insects. Kluwer Academic Publishers, Dordrecht, Boston and London, pp. 437–444.
    https://doi.org/10.1007/0-306-47577-4_4

    Reik, E.F. (1970) Fossil history. In: The insects of Australia: a textbook for students and research workers. 1st Edition. Melbourne University Press, Carlton, pp. 168–186.

    Riou, B. (1995) Lyristes renei n. sp., cigale fossile du Miocène ardéchois (Homoptera Cicadidae). Ecole pratique des hautes Etudes, Biologie et Evolution des Insectes, 7/8, 73–76.

    Sanborn, A.F. (2013). Catalogue of the Cicadoidea (Hemiptera: Auchenorrhyncha). Academic Press, San Diego, 1001 pp. [printed with publication date 2014 but published on 14 November 2013, according to Elsevier website: https://www.elsevier.com/books/catalogue-of-the-cicadoidea-hemiptera-auchenorrhyncha/sanborn/978-0-12-416647-9 (accessed 25 September 2017)]

    Scopoli, J.A. (1763) Entomologia carniologica exhibens Insecta carnioliae indigena et distributa in ordines, genera, species, varietates. Methodo Linnaeana. Trattner, Vindobonae, [xxxii] + 423 pp.
    https://doi.org/10.5962/bhl.title.119976

    Scopoli, J.A. (1772) Observationes Zoologicae. Annus Historico Naturalis, 5, 70–125.

    Scudder, S.H. (1892) Some insects of special interest from Florissant, Colorado, and other points in the Tertiaries of Colorado and Utah. United States Geological Survey Bulletin, 93, 3–35, pls. 1–3.

    Serres, M. de (1829) Geognosie des terrains Tertiaires, ou tableau des principaux animaux invertebres des terrains marins Tertiaires, du midi de la France. Montpellier, Paris, xcii + 276 pp.

    Shcherbakov, D.E. (1984) Systematics and phylogeny of Permian Cicadomorpha (Cimicida and Cicadina). Paleontological Journal, 18 (2), 87–97.

    Shcherbakov, D.E. (1985) Homopterans (Cimicida = Hemiptera: Aphidina, Cicadina) in the Jurassic of Southern Siberia and Western Mongolia. In: Rasnitsyn, A.P. (Ed.), Jurassic Insects of Siberia and Mongolia. Transactions of Paleontological Institute, USSR Academy of Sciences, 211, pp. 23–28. [in Russian]

    Shcherbakov, D.E. (1986) Homoptera. Cicadina (=Auchenorrhyncha). In: Rasnitsyn, A.P. (Ed.), Insects in the Early Cretaceous ecosystems of West Mongolia. Transactions of Joint Soviet-Mongolian Paleontological Expedition, 28, pp. 47–50. [in Russian]

    Shcherbakov, D.E. (2000) Permian faunas of Homoptera (Hemiptera) in relation to phytogeography and the Permo-Triassic crisis. Paleontological Journal, 34 (Supplement 3), S251–S267.

    Shcherbakov, D.E. (2009) Review of the fossil and extant genera of the cicada family Tettigarctidae (Hemiptera: Cicadoidea). Russian Entomological Journal, 17, 343–348. [dated 2008 but published 2009]

    Shcherbakov, D.E. (2012) A new subfamily of Mesozoic Hylicellidae (Homoptera Cicadomorpha). Russian Entomological Journal, 21, 441–444.

    Shcherbakov, D.E. & Popov, Y.A. (2002) Superorder Cimicidea Laicharting, 1781 order Hemiptera Linné, 1758. The bugs, cicadas, plantlice, scale insects, etc. (=Cimicida Laicharting, 1781, =Homoptera Leach, 1815 + Heteroptera Latreille, 1810). In: Rasnitsyn, A.P. & Quicke, D.L.J. (Eds.), History of Insects. Kluwer Academic Publishers, Dordrecht, pp. xii + 517.

    Shi, G., Grimaldi, D.A,, Harlow, G.E., Wang, J., Wang, J., Yang, M., Lei, W., Li, Q. & Li, X. (2012) Age constraint on Burmese amber based on U-Pb dating of zircons. Cretaceous Research, 37, 155–163.
    https://doi.org/10.1016/j.cretres.2012.03.014

    Stål, C. (1861) Genera nonnulla nova Cicadinorum. Annales de la Société Entomologique de France, Series 4, 1, 613–622.

    Straus, A. (1952) Thallophyten, Kryptogamen und Gymnospermen aus dem Pliozan von Willershausen. Berichte de Deutschen Botanischen Gesellschaft, 65 (4), 74–79, pls. 3 and 4.

    Théobald, N. (1937) Les insects fossils des terrains Oligocenes de France. G. Thomas, Nancy, 473 pp., 29 pls.

    Tietz, O., Berner, T. & Matting, E. (1998) Insekten aus dem Unteroligozän von Seifhennersdorf in der Oberlausitz. Abhandlungen und Berichte des Naturkundemuseums, Gorlitz, 70 (2), 139–154.

    Wagner, W. (1967) Die Singzikaden (Homoptera, Cidadae [sic]) aus dem Pliozän von Willershausen. Bericht der Naturhistorischen Gesellschaft zu Hannover, 111, 91–94.

    Walther, H. & Kvaček, Z. (2007) Early Oligocene flora of Seifhennersdorf (Saxony). Acta Musei Nationalis Pragae, Series B, 63 (2–4), 85–174.

    Wang, B., Zhang, H., Wappler, T. & Rust, J. (2010) Palaeontinidae (Insecta: Hemiptera: Cicadomorpha) from the Upper Jurassic Solnhofen Limestone of Germany and their phylogenetic significance. Geological Magazine, 147, 570–580.
    https://doi.org/10.1017/S0016756809990896

    Wang, B., Zhang, H., Jarzembowski, E.A., Fang, Y. & Zheng, D. (2013) Taphonomic variability of fossil insects: a biostratinomic study of Palaeontinidae and Tettigarctidae (Insecta: Hemiptera) from the Jurassic Daohugou Lagerstätte. Palaios, 28, 233–242.
    https://doi.org/10.2110/palo.2012.p12-045r

    Whalley, P.E.S. (1983) A survey of Recent and fossil cicadas (Insecta, Homoptera–Homoptera) in Britain. Bulletin of the British Museum Natural History, Geology, 37,139–147.
    https://doi.org/10.5962/bhl.part.5530

    Zhang, J.-F. (1989) Fossil insects from Shanwang, Shandong, China. Shandong Science and Technology, Jinan, 459 pp., 92 pls. [in Chinese with English summary]

    Zhang, J.-F. & Zhang, X.-Y. (1990) Fossil insects of cicada (Homoptera) and true bugs (Heteroptera) from Shanwang, Shandong. Acta Palaeontologica Sinica, 29 (3), 337–348, pls. 1–3. [in Chinese with English summary]

    Zhang, J.-F., Sun, B. & Zhang, X.-Y. (1994) Miocene insects and spiders from Shanwang, Shandong. Science Press, Beijing, 298 pp., 44 pls. [in Chinese with English summary]