Skip to main content Skip to main navigation menu Skip to site footer
Type: Article
Published: 2018-05-31
Page range: 401–455
Abstract views: 183
PDF downloaded: 15

Stink bugs (Hemiptera: Pentatomidae) from Brazilian Amazon: checklist and new records

Museu Paraense Emílio Goeldi, Coordenação de Zoologia. Av. Perimetral, Terra Firme, CEP 66077-830, Belém, Pará, Brazil. Programa de Capacitação Institucional. CNPq Fellowship.
Museu Paraense Emílio Goeldi, Coordenação de Zoologia. Av. Perimetral, Terra Firme, CEP 66077-830, Belém, Pará, Brazil.
Universidade Federal do Pará, Instituto de Ciências Biológicas, Av. Augusto Correa, CEP 66075-110, Belém, Pará, Brazil.
Hemiptera biodiversity Heteroptera Neotropical Region

Abstract

A list of the stink bugs (Pentatomidae) species of the Brazilian Amazon is provided for the first time, based on literature review and identification of the specimens of Pentatomidae deposited in the entomological collection of the Museu Paraense Emílio Goeldi (MPEG). Amazonian data base presented here has 324 species of Pentatomidae belonging to 102 genera, ten tribes and five subfamilies: Asopinae, Cyrtocorinae, Discocephalinae, Edessinae and Pentatominae. The Pentatominae has the largest number of species registered to Brazilian Amazon (144 species). The MPEG entomological collection has 122 species distributed in 53 genera, nine tribes and five subfamilies. Forty species from eight tribes and four subfamilies are recorded for the first time to Brazilian Amazon, including species from Podisus Herrich-Schäffer (Asopinae); Adoxoplatys Breddin, Macropygium Spinola, Neoadoxoplatys Kormilev, Ochlerus Spinola and Platycarenus Fieber (Discocephalinae); Edessa Fabricius (Edessinae); Arocera Spinola, Arvelius Spinola, Banasa Stål, Dichelops Spinola, Euschistus Dallas, Mormidea Amyot & Serville, Oebalus Stål, Pellaea Stål, Piezodorus Fieber, Proxys Spinola and Thyanta Stål (Pentatominae). Three of these species–Podisus maculiventris (Say, 1831) Neoadoxoplatys saileri Kormilev, 1956 and Edessa laticornis Stål, 1872are new records to South America. The number of species of Pentatomidae to Brazilian Amazon represents approximately 50% of the species registered to Brazil. Such list of species is important to better understand the distributional range and diversity of the species of Pentatomidae in the Amazon region.

 

References

  1. Almeida, F.R.A.A., Nunes, B.M. & Fernandes, J.A.M. (2018) A new genus and new species of Edessinae (Hemiptera: Heteroptera: Pentatomidae). Zootaxa, 4377 (2), 254–268.
    https://doi.org/10.11646/zootaxa.4377.2.6

    Arismendi, N. & Thomas, D.B. (2003) Pentatomidae (Heteroptera) of Honduras, a checklist with description of a new ochlerine genus. Insecta Mundi, 17, 1–18.

    Barcellos, A. (2006) Hemípteros terrestres. In: Becker, F.G., Ramos, R.A. & Moura, L.A. (Orgs.), Região da Lagoa do Casamento e dos Butiazais de Tapes, Planície Costeira do Rio Grande do Sul. Ministério do Meio Ambiente, Brasília, pp. 198–209.

    Barcellos, A. & Grazia, J. (2003) Revision of Brachystethus (Heteroptera, Pentatomidae, Edessinae). Iheringia, Série Zoologia, 93 (4), 413–446.
    https://doi.org/10.1590/S0073-47212003000400008

    Barcellos, A. & Grazia, J. (2008) Revision of the genus Poriptus Stål (Hemiptera: Heteroptera: Pentatomidae: Pentatominae). Zootaxa, 1821, 25–36.
    https://doi.org/10.5281/zenodo.183013

    Becker, M. (1977a) A review of the genus Colpocarena Stål (Heteroptera, Pentatomidae, Discocephalinae). Revista Brasileira de Biologia, 37 (2), 367–373.

    Becker, M. (1977b) The genus Abascantus Stål, with the description of two new species (Heteroptera, Pentatomidae, Discocephalinae). Revista Brasileira de Biologia, 37 (2), 385–393.

    Becker, M. & Grazia, J. (1970) Sobre os gêneros Lopadusa Stål e Bothrocoris Mayr (Hemiptera, Pentatomidae, Pentatomini). Revista Brasileira de Biologia, 30 (2), 217–232.

    Becker, M. & Grazia-Vieira, J. (1971a) Sobre o gênero Rhyncholepta Bergroth, 1911, com a descrição de uma nova espécie (Hemiptera, Pentatomidae, Pentatominae). Revista Brasileira de Biologia, 31 (3), 389–399.

    Becker, M. & Grazia-Vieira, J. (1971b) Contribuição ao conhecimento da superfamília Pentatomoidea na Venezuela (Heteroptera). Iheringia, Série Zoologia, 40, 3–26.

    Becker, M. & Grazia-Vieira, J. (1977) The Pentatomoidea [Heteroptera] collected in French Guiana by the expedition of the Muséum National D’Histoire Naturelle. Annales de la Société Entomologique de France, 13 (1), 53–67.

    Becker, M. & Grazia, J. (1971) Contribuição ao conhecimento da Superfamilia Pentatomoidea na Venezuela (Heteroptera). Iheringia, Série Zoologia, 40, 3–26.

    Becker, M. & Grazia, J. (1985) Revisão do gênero Dinocoris Burmeister, 1835 (Heteroptera, Pentatomidae, Discocephalinae). Revista Brasileira de Zoologia, 3 (2), 65–108.
    https://doi.org/10.1590/S0101-81751985000200001

    Becker, M. & Grazia, J. (1989a) Novas contribuições ao gênero Ablaptus Stål, 1864 (Heteroptera, Pentatomidae, Discocephalinae). Memórias do Instituto Oswaldo Cruz, 84 (IV), 57–68.
    https://doi.org/10.1590/S0074-02761989000800015

    Becker, M. & Grazia, J. (1989b) The Discocephaline genus Alcippus Stål, 1867 (Heteroptera, Pentatomidae). Revista Brasileira de Entomologia, 33 (1), 49–55.

    Becker, M. & Grazia, J. (1992) Revisão do gênero Agaglitus Stål (Heteroptera, Pentatomidae, Discocephalinae). Revista Brasileira de Entomologia, 36 (4), 831–842.

    Becker, M. & Grazia, J. (1997) A new combination and a new synonymy in Discocephalini (Heteroptera, Pentatomidae). Revista brasileira de Zoologia, 14 (2), 319–321.
    https://doi.org/10.1590/S0101-81751997000200005

    Becker, M. & Ruckes, H. (1969) A Review of the Genus Sympiezorhincus Spinola (Heteroptera, Pentatomidae, Discocephalinae). American Museum Novitates, 2391, 1–17.

    Berg, C. (1878) Hemiptera Argentina: Ensayo de una monografía de los Hemípteros Heterópteros y Homópteros de la República Argentina. Anales de la Sociedad Científica Argentina, 5 (6), 297–314.

    Bernardes, J.L.C., Grazia, J. & Barcellos, A. (2006) New Species of Neotibilis Grazia & Barcellos (Hemiptera: Pentatomidae: Pentatomini). Neotropical Entomology, 35 (3), 344–348.
    https://doi.org/10.1590/S1519-566X2006000300008

    Bianchi, F., Gonçalves, V.R., Souza, J.R. & Campos, L.A. (2016) Description of three new species of Glyphepomis Berg (Heteroptera: Pentatomidae: Pentaminae). Zootaxa, 4103 (5), 443–452.
    https://doi.org/10.11646/zootaxa.4103.5.2

    Bianchi, F. M., Deprá, M., Ferrari, A., Grazia, J., Valente, V.L.S. & Campos, L.A. (2017) Total evidence phylogenetic analysis and reclassification of Euschistus Dallas within Carpocorini (Hemiptera: Pentatomidae: Pentatominae). Systematic Entomology, 42, 399–409.
    https://doi.org/10.1111/syen.12224

    Bonatto, S.L. & Grazia, J. (1989) Contribuição ao estudo do gênero Mormidea Amyot & Serville, 1843. I. M. speciosa Haglund, 1868 (Heteroptera, Pentatomidae, Pentatomini). Iheringia, Série Zoologia, 69, 3–8.

    Brailovsky, H. (1981) Revisión del género Arvelius Spinola (Hemiptera-Heteroptera-Pentatomidae-Pentatomini). Anales del Instituto de Biología Universidad Nacional Autónoma de México, Serie Zoología, 51 (1), 239–298.

    Breddin, G. (1901) Neu netropische Wanzen und Zirpen. Societas entomologica, 16 (16), 123–124.

    Breddin, G. (1903) Neu Arten der Gattung Edessa Fab. Societas entomologica, 18 (18), 139–140.

    Breddin, G. (1904 a) Beiträge zur Systematik der Rhynchoten. Sitzungsberichte der Gesellschaft Naturforschender Freunde zu Berlin, 6, 135–153.

    Breddin, G. (1904b) Neue Rhynchotenausbeute aus Süd-Amerika. Societas entomologica, 19 (7), 49–50.

    Breddin, G. (1905) Neue Rhynchoten (Vorläufige Diagnosen). Societas entomologica, 20 (12), 26–27.

    Breddin, G. (1907) Beiträge zur Systematik der Pentatomiden Südamerikas. Sitzungs-Berichte der Gesellschaft naturforschender freunde zu Berlin, 10, 327–348.

    Brown Jr., K.S. (1997) Diversity, disturbance, and sustainable use of Neotropical forests: insects as indicators for conservation monitoring. Journal of Insect Conservation, 1, 25–42.
    https://doi.org/10.1023/A:1018422807610

    Bunde, P.R.S., Grazia, J. & Junior, M.S.M. (2006) Nova espécie de Euschistus (Mitripus) da Argentina e sul do Brasil (Hemiptera, Pentatomidae, Pentatominae). Iheringia, Série Zoologia, 96 (3), 289–291.
    https://doi.org/10.1590/S0073-47212006000300003

    Bunde, P.R.S., Grazia, J., Mendonça Júnior, M.S., Schwertner, C.F., Silva, E.J.E. & Garcia, É.N. (2010) Pentatomidae (Hemiptera: Heteroptera) of the Pampa biome: Serra do Sudeste and Parque de Espinilho da Barra do Quaraí, Rio Grande do Sul, Brazil. Biota Neotropical, 10 (3), 83–88.
    https://doi.org/10.1590/S1676-06032010000300008

    Bundy, C.S., Esquivel, J.F., Panizzi, A.R., Eger, J.E., Davis, J.A. & Jones, W.A. (2018) Piezodurus guildinii. In: McPherson, J.E. (Ed.), Invasive Stink Bugs and Related Species (Pentatomoidea) Biology, Higher Systematics, Semiochemistry, and Management. CRC Press, Boca Raton, Florida, pp. 425–451.

    Burmeister, H. (1835) Handbuch der Entomologie. Zweiter Band. Eimer, G., Berlin, 1050 pp.

    Cambra, R.A., Carranza, R., Ramos, Y.J.A. & Murgas, A.S. (2018) Los Pentatómidos (Hemiptera: Heteroptera) de Panamá. Revista Nicaraguense de Entomologia, 149, 1–21.

    Campos, L.A. (2005) A new species of Cromata Rolston from Brazilian and Venezuelan Amazon Forest (Hemiptera: Pentatomidae: Discocephalinae). Zootaxa, 820 (1), 1–6.
    https://doi.org/10.11646/zootaxa.820.1.1

    Campos, L.A. & Grazia, J. (1995) Paratibraca, um novo gênero de Pentatomini (Heteroptera, Pentatomidae). Iheringia, Série Zoologia, 79, 163–171.

    Campos, L.A. & Grazia, J. (1998) Revisão de Glyphepomis Berg, 1891 (Heteroptera, Pentatomidae). Revista Brasileira de Entomologia, 41 (2–4), 203–212.

    Campos, L.A. & Grazia, J. (2000) Descrição de um novo gênero de Ochlerini Rolston (Heteroptera, Pentatomidae, Discocephalinae). Revista Brasileira de Zoologia, 17 (3), 789–793.
    https://doi.org/10.1590/S0101-81752000000300022

    Campos, L.A. & Roell, T. (2017) Description of four new species of Ochlerini from South America (Hemiptera: Pentatomidae: Discocephalinae). Zootaxa, 4236 (3), 507–520.
    https://doi.org/10.11646/zootaxa.4236.3.5

    Campos, L.A., Teixeira, R.A. & Martins, F.S. (2007) Três padrões de coloração de ninfas de Arvelius albopunctatus (De Geer) (Hemiptera: Pentatomidae). Neotropical Entomology, 36 (6), 972–975.
    https://doi.org/10.1590/S1519-566X2007000600022

    Castro-Huertas, V., Schwertner, C.F. & Fernández, F. (2015) New records of stink bugs (Hemiptera: Pentatomidae) from Colombia. Zootaxa, 3973 (3), 553–566.
    https://doi.org/10.11646/zootaxa.3973.3.9

    Cervantes-Peredo, L. & Ortega-León, G. (2014) Description of a New Species of Neoadoxoplatys and Immature Stages of Neodoxoplatys saileri Kormilev (Heteroptera: Pentatomidae) Associated with Bamboo. Systematics, Morphology and Physiology, 43, 236–244.
    https://doi.org/10.1007/s13744-014-0203-3

    Chocorosqui, V.R. & Panizii, A.R. (2004) Impact of cultivation systems on Dichelops melacanthus (Dallas) (Heteroptera: Pentatomidae) population and damage and its chemical control on wheat. Neotropical Entomology, 33 (4), 487–492.
    https://doi.org/10.1590/S1519-566X2004000400014

    Correia, A.O. & Fernandes, J.A.M. (2016) Grammedessa, a new genus of Edessinae (Hemiptera: Heteroptera: Pentatomidae). Zootaxa, 4107 (4), 541–565.
    https://doi.org/10.11646/zootaxa.4107.4.4

    Costa, E.C. & Corseuil, E. (1979) Avaliação dos danos causados por Piezodorus guildinii (Westwood, 1937) (Hemiptera, Pentatomidae) em soja. Revista Centro Ciências Rurais, 9 (4), 403–408.

    Couturier, G. (1988) Alguns insetos do cubiu (Solanum sessiliflorum Dunal Var. sessiliflorum Dunal, Solanaceae) na região de Manaus—Am. Acta Amazonica, 18 (3–4), 93–103.
    https://doi.org/10.1590/1809-43921988183103

    Dallas, W.S. (1851) List of the specimens of hemipterous insects in the collection of the British Museum. Parte 1. Printed by order of the Trustees, London, 390 pp., 15 pls.

    De Clercq, P. (2000) Predaceous stink bugs (Pentatomidae: Asopinae). In: Schaefer, C.W. & Panizzi, A.R. (Eds.), Heteroptera of economic importance. CRC Press, Boca Raton, pp. 737–789.
    https://doi.org/10.1201/9781420041859.ch32

    De Clercq, P. (2008) Predatory Stink Bugs (Hemiptera: Pentatomidae, Asopinae). In: Capinera, L.J. (Ed.), Encyclopedia of Entomology, Springer Science, Business Media B.V, pp 3042–3045.
    https://doi.org/10.1007/978-1-4020-6359-63115

    Distant, W.L. (1880–1893) Insecta. Rhynchota, Hemiptera—Heteroptera. In: Godman, F.D & Salvin, O. (Eds.), Biologia Centrali-Americana, London, Part 1, pp. 1–462.

    Eger, J.E. (1978) Revision of the Genus Loxa (Hemiptera: Pentatomidae). Journal of The New York Entomological Society, 86 (3), 224–259.

    Erichson, W.F (1848) Insecten. In: von Schomburgk, R. (Ed.). Reisen in British-Guiana In den Jahren 1840–1844. Weber, J. J., Leipzig, pp. 553–617.

    Esquivel, J.F., Musolin, D.L., Jones, W.A., Rabitsch, W., Greene, J.K., Toews, M.D., Schwertner, C.F., Grazia, J. & McPherson, R.M. (2018) Nezara viridula (L) In: McPherson, J.E. (Ed.), Invasive Stink Bugs and Related Species (Pentatomoidea): Biology, Higher Systematics, Semiochemistry, and Management. CRC Press, Boca Raton, London, New York, pp. 351–423.

    Eva, H.D., Miranda, E.E., Di Bella, C.M., Gond, V., Huber, O., Sgrenzaroli, M., Jones, S., Coutinho, A., Dorado, A., Guimarães, M., Elvidge, C., Achard, F., Belward, A.S., Bartholomé, E., Baraldi, A., De Grandi, G., Vogt, P., Fritz, S. & Hartley, A. (2002) A vegetation map of South America. Office for Official Publications of the European Communities, Luxembourg, 34 pp.

    Faúndez, E. & Rider, D. (2015) First record of Mormidea speciosa Haglund, 1868 (Heteroptera: Pentatomidae: Carpocorini) from French Guiana. Boletín del Museo Nacional de Historia Natural del Paraguay, 19 (2), 70–71.

    Favetti, B.M., Krinski, D., Butnariu, A.R. & Loiácono, M.S. (2013) Egg parasitoids of Edessa meditabunda (Fabricius) (Pentatomidae) in lettuce crop. Revista Brasileira de Entomologia, 57 (2), 236–237.
    https://doi.org/10.1590/S0085-56262013005000014

    Fernandes, J.A.M. (2010) A new genus and species of Edessinae from Amazon Region (Hemiptera: Heteroptera: Pentatomidae). Zootaxa, 2662, 53–65.

    Fernandes, J.A.M. & van Doesburg P.H. (2000a) The E. dolichocera-group of Edessa Fabricius, 1803 (Heteroptera: Pentatomidae: Edessinae). Zoologische Mededelingen Leiden, 73, 305–315.

    Fernandes, J.A.M. & van Doesburg, P.H. (2000b) The E. beckeri-group of Edessa Fabricius, 1803 (Heteroptera: Pentatomidae: Edessinae). Zoologische Mededelingen Leiden, 74 (7), 143–150.

    Fernandes, J.A.M. & van Doesburg, P.H. (2000c) The E. cervus-group of Edessa Fabricius, 1803 (Heteroptera: Pentatomidae: Edessinae). Zoologische Mededelingen Leiden, 74 (8), 151–165.

    Fernandes, J.A.M & Grazia, J. (1996) Revisão do gênero Hypatropis Bergroth, 1891 (Heteroptera, Pentatomidae). Revista Brasileira de Entomologia, 40 (3/4), 341–352.

    Fernandes, J.A.M. & Grazia, J. (1998a) Amauromelpia, a new northern Neotropical genus (Heteroptera, Pentatomidae). Iheringia, Série Zoologia, 84, 153–160.

    Fernandes, J.A.M. & Grazia, J. (1998b) Revision of the genus Tibraca Stâl (Heteroptera, Pentatomidae, Pentatominae). Revista Brasileira de Zoologia, 15 (4), 1049–1060.
    https://doi.org/10.1590/S0101-81751998000400022

    Fernandes, J.A.M. & Grazia, J. (2006) Revisão do gênero Antiteuchus Dallas (Heteroptera, Pentatomidae, Discocephalinae). Revista Brasileira de Entomologia, 50 (2), 165–231.
    https://doi.org/10.1590/S0085-56262006000200004

    Fernandes, J.A.M., Grazia, J. & Campos, L.D. (2011) Redescription of Callostethus Ruckes, 1961 (Hemiptera: Heteroptera: Pentatomidae: Discocephalinae) with the description of C. flacolineatus sp. nov. Zootaxa, 2866, 55–60.

    Fernandes, J.A.M., Grazia, J. & Lobo, D. (2008) Revision of the genus Psorus Bergroth, 1914 (Hemiptera: Pentatomidae: Discocephalinae). Zootaxa, 1696, 48–56.

    Fernandes, J.A.M., Silva, V.J., Correia, A.O. & Nunes, B.M. (2015) New species of Edessa Fabricius, 1803 (Hemiptera: Pentatomidae) from Costa Rica. Zootaxa, 3999 (4), 511–536.
    https://doi.org/10.11646/zootaxa.3999.4.3

    Ferrari, A., Schwertner, C.F. & Grazia, J. (2010) Review, cladistic analysis and biogeography of Nezara Amyot & Serville (Hemiptera: Pentatomidae). Zootaxa, 2424, 1–41.

    Firmino, J.V.L., Mendonça Junior, M.D.S, Lima, I.M.M. & Grazia, J. (2017) Pentatomidae (Hemiptera: Heteroptera) in Herbaceous and Shurb Strata of Atlantic Forest Remnants in Northeastern Brazil. Environmental Entomology, 43 (3), 480–486.

    https://doi.org/10.1093/ee/nvx062

    Fortes, N.D.F. & Grazia, J. (1998) Um novo gênero e novas espécies de Pentatomini do Brasil (Heteroptera, Pentatomidae). Revista Brasileira de Zoologia, 15 (4), 867–874.
    https://doi.org/10.1590/S0101-81751998000400004

    Fortes, N.D.F. & Grazia, J. (2000) Novas espécies do gênero Rio (Heteroptera, Pentatomidae). Iheringia, Série Zoologia, 88 (88), 67–102.

    Fortes, N.D.F. & Grazia, J. (2005) Revisão e análise cladística de Serdia Stål Heteroptera, Pentatomidae, Pentatomini). Revista Brasileira de Entomologia, 49 (3), 294–339.

    https://doi.org/10.1590/S0085-56262005000300002

    Frey-da-Silva, A. & Grazia, J. (2001) Novas espécies de Acrosternum subgênero Chinavia (Heteroptera, Pentatomidae, Pentatomini). Iheringia, Série Zoologia, 90, 107–126.
    https://doi.org/10.1590/S0073-47212001000100011

    Frey-da-Silva, A., Grazia, J. & Fernandes, J.A.M. (2002) Revision of the genus Ogmocoris Mayr, 1864 (Heteroptera, pentatomidae, Pentatomini) Bulletin Zoological Museum University of Amsterdam, 52 (10), 179–185.

    Frey-da-Silva, A., Grazia, J. & Fernandes, J.A.M. (2014) Revisão do gênero Paramecocephala Benvegnú, 1968 (Heteroptera, Pentatomidae). Revista Brasileira de Entomologia, 46 (2), 209–225.
    https://doi.org/10.1590/S0085-56262002000200013

    Froeschner, R.C. (1988) Family Pentatomidae Leach, 1815: the stink bugs. In: Henry, T. & Froeschner, R. (Eds.), Catalog of the Heteroptera, or True Bugs, of Canda and the United States. E.J. Brill, Leiden, pp. 544–597.

    Gaedike, H. (1971) Katalog der in Sammlungen des ehemaligen Deutschen Entomologischen Institutes aufbewahrten Typen—V. Heteroptera. Beiträge zur Entomologie, 21, 79–159.

    Garbelotto, T.A. & Campos, L.A. (2014) Pentatominae do Sul de Santa Catarina Sociedade Brasileira de Zoologia, Curitiba, 80 pp. Zoologia: guias e manuais de identificação series. SciELO Books ISBN 978–85–98203–08–9. Available from: http://books.scielo.org/id/fs5j4/pdf/garbelotto-9788598203089.pdf (accessed 15 May 2018)

    Garbelotto, T.A., Campos, L.A. & Grazia, J. (2014) Xynocoris, new genus of Ochlerini from Central and South America (Hemiptera: Pentatomidae: Discocephalinae). Zootaxa, 3869 (3), 281–305.
    https://doi.org/10.11646/zootaxa.3869.3.4

    Genevcius, B.C. & Schwertner, C.F. (2014) Review and phylogeny of the geniculata group, genus Chinavia (Heteroptera: Pentatomidae), with notes on biogeography and morphological evolution. Zootaxa, 3847 (1), 033–056.
    https://doi.org/10.11646/zootaxa.3847.1.2

    González, U., Juárez, G. & Faúndez, E.I. (2016) Nuevo registro de Pellaea stictica Dallas, 1851 (Hemiptera: Heteroptera: Pentatomidae) para Perú. Boletin de la Sociedad Entomológica Aragonesa (S.E.A.), 58, 235–236.

    Grazia, J. (1967) Estudos sobre o gênero Galedata Amyot & Serville, 1843 (Hemiptera-Heteroptera, Pentatomidae). Iheringia, Série Zoologia, 35, 45–59.

    Grazia, J. (1978) Revisão do gênero Dichelops Spinola, 1837 (Heteroptera, Pentatomidae, Pentatomini). Iheringia, Série Zoologia, 53, 3–119.

    Grazia, J. (1980) Revisão do gênero Pallantia Stål, 1862 (Heteroptera, Pentatomidae). Revista Brasileira de Entomologia, 24 (1), 15–27.

    Grazia, J. (1983) Sobre O Gênero Phalaecus Stål, 1862 Com Descrição de Quatro Novas Espécies (Heteroptera, Pentatomidae). Revista Brasileira de Entomologia, 27 (2), 177–187.

    Grazia, J. (1989) Contribuições ao conhecimento dos gêneros Elanela e Nocheta (Heteroptera, Pentatomini). Revista Brasileira de Zoologia, 6 (1), 57–62.
    https://doi.org/10.1590/S0101-81751989000100008

    Grazia-Vieira, J. (1972) O gênero Mayrinia Horváth, 1925 (Heteroptera, Pentatomidae, Pentatomini). Revista Peruana de Entomologia, 15 (1), 117–124.

    Grazia, J., & Barcellos, A. (1994) Neotibilis, um gênero de Pentatomini (Heteroptera). Iheringia, Série Zoologia, 76, 55–94.

    Grazia, J., & Barcellos, A. (2005) Revision of Taurocerus (Heteroptera, Pentatomidae, Pentatomini). Iheringia, Série Zoologia, 95 (2), 173–181.
    https://doi.org/10.1590/S0073-47212005000200007

    Grazia, J., Barros, L.D. & Barão, K.R. (2016) Elanela Rolston revisited (Heteroptera: Pentatomidae): new distributional records and description of new species. Zootaxa, 4092 (4), 561–571.
    https://doi.org/10.11646/zootaxa.4092.4.6

    Grazia, J., Becker, M. & Thomas, D.B. (1994) A review of the genus Pseudevoplitus Ruckes, (Heteroptera: Pentatomidae) with the description of three new species. Journal of the New York Entomological Society, 102, 442–455.

    Grazia, J., Bolze, G.J. & Barão, K.R. (2016) There and back again: contributions on Pseudevoplitus Ruckes (Heteroptera: Pentatomidae). Zootaxa, 4078 (1), 161–167.

    https://doi.org/10.11646/zootaxa.4078.1.15

    Grazia, J. & Campos, L. (1996) Hypanthracos, a new genus of Pentatomini (Heteroptera: Pentatomidae). Iheringia, Série Zoologia, 80, 13–19.

    Grazia, J., Campos, L.A. & Becker, M. (2000) Revision of Cataulax Spinola, with Architas Distant as a new synonymy (Heteroptera: Pentatomidae: Discocephalini). Anais da Sociedade Entomológica do Brasil, 29 (3), 475–488.
    https://doi.org/10.1590/S0301-80592000000300010

    Grazia, J., Campos, L.A., Greve, C. & Rocha, F.S. (2002) Notas sobre Pseudevoplitus (Heteroptera, Pentatomidae) e descrição de duas espécies novas. Iheringia, Série Zoologia, 92, 53–61.
    https://doi.org/10.1590/S0073-47212002000100003

    Grazia, J., Campos, L.A., Fernandes, J.A.M., Schwertner, C.F. & Garbelotto, T.A. (2017) Pentatomidae in Catálogo Taxonômico da Fauna do Brasil. PNUD. Available from: http://fauna.jbrj.gov.br/fauna/faunadobrasil (accessed 4 August 2017)

    Grazia, J. & Greve, C. (2011) Contributions to the knowledge of Elanela: Elanela jordi sp. nov., from Amazonas, Brazil (Hemiptera, Heteroptera, Pentatomidae). Heteropterus Revista de Entomologia, 11, 261–266.

    Grazia, J., Fernandes, J.A.M. & Schwertner, C.F. (1999) Stysiana, a new genus and four new species of Pentatomini (Heteroptera, Pentatomidae) of the Neotropical region. Acta Societatis Zoologicae Bohemicae, 63 (1–2), 71–83.

    Grazia, J. & Frey-da-Silva, A. (2003) Rideriana amazonica gen. nov. and sp. nov. of Pentatomini (Heteroptera, Pentatomidae). Revista Brasileira de Zoologia, 20 (3), 507–510.
    https://doi.org/10.1590/S0101-81752003000300023

    Grazia, J. & Hildebrand, R. (1982) Revisão do gênero Berecynthus Stål, 1862 (Heteroptera, Pentatomidae, Pentatomini). Revista Brasileira de Entomologia, 26 (2), 173–182.

    Grazia, J., Schuh, R.T. & Wheeler, W.C. (2008) Phylogenetic relationships of family groups in Pentatomoidea based on morphology and DNA sequences (Insecta: Heteroptera). Cladistics, 24, 932–976.
    https://doi.org/10.1111/j.1096-0031.2008.00224.x

    Grazia, J. & Schwertner, C.F. (2011) Checklist dos percevejos–do–mato (Hemiptera: Heteroptera: Pentatomoidea) do Estado de São Paulo, Brasil. Biota Neotropical, 11 (1), 1–12.
    https://doi.org/10.1590/S1676-06032011000500034

    Grazia, J., Schwertner, C.F. & Greve, C. (2008) Two new species of the genus Chloropepla (Hemiptera: Pentatomidae: Pentatominae) from Brazil. Acta Entomologica Musei Nationalis Pragae, 48 (2), 533–542.

    Grazia, J. & Silva, A.N. (2006) Elanela kerzhneri sp.n., a new species of Pentatomidae from Brazil (Hemiptera: Heteroptera: Pentatominae). Russian Entomological Journal, 15 (2), 159–160.

    Grazia, J., Simões, F.L. & Panizzi, A.R. (2015) Morphology, Ontogeny, Reproduction, and Feeding of True Bugs. In: Panizzi, A.R. & Grazia, J. (Eds.), True Bugs (Heteroptera) of the Neotropics. Entomology in Focus. Vol. 2. Springer, Dordrecht, pp. 21–55.
    https://doi.org/10.1007/978-94-017-9861-7_2

    Grazia, J. & Teradaira, C.T. (1980) Nova espécie de Chloropepla Stål, 1867 coletada em Tucurui, Pará, Brasil (Heteroptera, Pentatomidae). Anais da Sociedade Entomológica do Brasil, 9, 123–131.

    Greve, C., Schwertner, C.F. & Grazia, J. (2013) Cladistic analysis and synopsis of Chloropepla Stål (Hemiptera: Heteroptera: Pentatomidae) with the description of three new species. Insect Systematics & Evolution, 44, 1–43.
    https://doi.org/10.1163/1876312X-04401002

    Greve, C., Fortes, N.D.F. & Grazia, J. (2003) Estágios imaturos de Oebalus poecilus (Heteroptera, Pentatomidae). Iheringia, Série Zoologia, 93 (1), 89–96.
    https://doi.org/10.1590/S0073-47212003000100010

    Herrich-Schäeffer, G.W.A. (1840) Die Wanzensrtigen Insecten. C. M. Zelischer, Nürnberg, 127 pp.

    Kirkaldy, G.W. (1909) Catalogue of the Hemiptera (Heteroptera) with biological and anatomical references, lists of foodplants and parasites, etc. Vol. I. Cimicidae. Felix L. Dames, Berlin, xl + 392 pp.

    Kormilev, N.A. (1949) Dos especies nuevas del género Adoxoplatys Breddin (1903) de Bolivia (Hemiptera, Pentatomidae). Notas Mus. Plata, 14, 313–324.

    Klein, J.T., Barcellos, A., Grazia, J. & Redaelli, L.R. (2012) Contributions to the knowledge of Dichelops (Dichelops) with the description of a new species (Hemiptera: Heteroptera: Pentatomidae: Pentatominae: Carpocorini). Zootaxa, 3157, 61–68.

    Krinski, D., Favetti, B.M. & Butnariu, A.R. (2012) First Record of Edessa meditabunda (F.) on Lettuce in Mato Grosso State, Brazil. Neotropical Entomology, 41 (1), 79–80.
    https://doi.org/10.1007/s13744-011-0012-x

    Krinski, D., Foerster, L.A. & Grazia, J. (2015) Hypatropis inermis (Hemiptera, Pentatomidae): first record on rice crops. Revista Brasileira de Entomologia, 59 (1), 12–13.
    https://doi.org/10.1016/j.rbe.2014.11.001

    Lepeletier, A.L.M. & Serville, J.G.A. (1825) In: Olivier, G.A. (Ed.), Encyclopédie Méthodique. Vol. 10. Mme veuve Agasse, Paris, pp. 1–344.

    Lewinsohn, T.M., Prado, P.K.L. & Almeida, A.M. (2001) Inventários Bióticos Centrados em recursos: Insetos fitófagos e plantas hospedeiras. In: Dias, B.F.S. & Garay, I. (Orgs.), Conservação da biodiversidade em ecossistemas tropicais. Avanços conceituais e revisão de novas metodologias de avaliação e Monitoramento. Vozes, Petrópolis, pp. 174–189.

    Lima, S.O.F., Martins, M.B., Prudente, A.L.C., Montag, L.F.A., Monnerat, M.C., Cabral, P.R. & Rosário, D.A.P. (2008) Biodiversidade na Província petrolífera de Urucu. Petrobrás, Rio de Janeiro, 194 pp.

    Lima, M.L., Fernandes, J.A.M. & Lima, I.M.M. (2010) Record of Edessa scabriventris Stål (Hemiptera: Pentatomidae) Associated to Eugenia uniflora (Brazilian-Cherry) and Psidium guajava (Guava) (Myrtaceae), in North-Northeastern Brazil. Neotropical Entomology, 39 (4), 674–675.
    https://doi.org/10.1590/S1519-566X2010000400032

    Lupoli, R. (2017) Edessa amazonica Fernandes & Doesburg, 2000, espèce nouvelle pour la Guyane française et le Suriname, et première description de la femelle (Hemiptera, Pentatomidae, Edessinae). Bulletin de la Société entomologique de France, 122 (2), 199–202.

    Maciel, A.S., Garbelotto, T.A., Winter, I.C., Roell, T. & Campos, L.A. (2015) Description of the males of Lincus singularis and Lincus incisus (Hemiptera: Pentatomidae: Discocephalinae). Zoologia, 32 (2), 157–161.
    https://doi.org/10.1590/S1984-46702015000200007

    Maes, J.M. (1994) Catalogo de los Pentatomoidea (Heteroptera) de Nicaragua. Revista Nicaraguense de Entomologia, 28, 1–29.

    Mendonça Jr, M.S., Schwertner, C.F. & Grazia, J. (2009) Diversity of Pentatomoidea (Hemiptera) in riparian forests of Southern Brazil: taller forests, more bugs. Revista Brasileira de Entomologia, 53 (1), 121–127.
    https://doi.org/10.1590/S0085-56262009000100026

    Museu Paraense Emílio Goeldi (2016) Censo da biodiversidade da Amazônia Brasileira. Available from: http://www.museu-goeldi.br/censo/ (accesed 6 September 2017)

    Nascimento, D.R., Mendonça, M.T.S. & Fernandes, J.A.M. (2017) Description of a new group of species of Edessa (Hemiptera: Pentatomidae: Edessinae). Zootaxa, 4254 (1), 136–150.
    https://doi.org/10.11646/zootaxa.4254.1.10

    Packauskas, R.J. & Schaefer, C.W. (1998) Revision of the Cyrtocoridae (Hemiptera: Pentatomoidea). Annals of the Entomological Society of America, 91 (4), 363–386.
    https://doi.org/10.1093/aesa/91.4.363

    Panizzi, A.R., Smith, J.G., Pereira, L.A.G. & Yamashita, J. (1979) Efeitos dos danos de Piezodorus guildinii (Westwood, 1837) no rendimento e qualidade da soja. Anais do I Seminário Nacional de Pesquisa de Soja, 2, 59–68.

    Panizzi, A.R., McPherson, J.E., James, D.G., Javahery, M. & McPherson, R.M. (2000) Stink Bugs (Pentatomidae). In: Schaefer, C.W. & Panizzi, A.R. (Eds.), Heteroptera of economic importance. CRC Press, Boca Raton, London, New York, pp 421–474.

    Pook-da-Silva, P., Barão, K.R. & Grazia, J. (2013) Contributions to the knowledge of Dichelops Spinolsa: description of a new species of Dichelops (Diceraeus) and of the male of Dichelops (Prodichelops) divisus (Heteroptera: Pentatomidae: Pentatomine: Carpocorini). Zootaxa, 3609 (1), 060–068.
    https://doi.org/10.11646/zootaxa.3609.1.4

    Ribeiro, R.C., Lemos, P.W., Bernardino, A.S., Buecke, J. & Müller, A.A. (2010) Primeira Ocorrência de Alcaeorrhynchus grandis (Dallas) (Hemiptera: Pentatomidae) Predando Lagartas Desfolhadoras do Dendezeiro no Estado do Pará. Neotropical Entomology, 39 (1), 131–132.
    https://doi.org/10.1590/S1519-566X2010000100018

    Rider, D.A. (1992) Revision of Arocera Spinola, with description of two new species (Heteroptera: Pentatomidae). Journal of the New York Entomological Society, 100 (1), 99–136.

    Rider, D.A. (1994) A generic conspectus of the tribe Procleticini Pennington (Heteroptera: Pentatomidae), with the description of Parodmalea rubella new genus and species. Journal of the New York Entomological Society, 102 (2), 193–221.

    Rider, D.A. & Chapin, J.B. (1991) Revision of the genus Thyanta Stål, 1862 (Heteroptera: Pentatomidae). I. South America. Journal of the New York Entomological Society, 99 (1), 1–77.

    Rider, D.A. & Eger, Jr. J. (2008) Two new genera of Pentatomini for species previously placed in Mormidea Amyot & Serville (Hemiptera: Heteroptera: Pentatomidae: Pentatominae). Proceedings of the Entomological Society of Washington, 110 (4), 1050–1058.
    https://doi.org/10.4289/0013-8797-110.4.1050

    Rider, D.A. & Fischer, C. (1998) Zorcadium Bergroth, an objective junior synonym of Pseudobebaeus Fallou (Heteroptera: Pentatomidae). Entomological News, 109 (4), 274–276.

    Rider, D.A. & Rolston, L.H. (1987) Review of the genus Agroecus Dallas, with the description of a new species (Hemiptera: Pentatomidae). Journal of the New York Entomological Society, 95, 428–439.

    Rider, D.A., Schwertner, C.F., Vilímová, J., Rédei, D., Kment, P. & Thomas, D.T. (2018) Higher Systematics of the Pentatomoidea In: McPherson, J.E. (Ed.), Invasive Stink Bugs and Related Species (Pentatomoidea) Biology, Higher Systematics, Semiochemistry, and Management. CRC Press, Boca Raton, Florida, pp. 24–201.

    Rodrigues, H.D.D. & Ferreira-Keppler, R.L. (2013) Catalog of type specimens of invertebrates in the collection of the Instituto Nacional de Pesquisas da Amazônia, Manaus, Brazil. VI. Hexapoda: Hemiptera: Heteroptera. Zootaxa, 3716 (2), 192–206.
    https://doi.org/10.11646/zootaxa.3716.2.4

    Roell, T., Winter, I.C., Asipuela, R. & Campos, L.A. (2017) New records and update on the geographic distribution of fifteen species of Lincus Stål, 1867 (Insecta: Hemiptera: Pentamidae) associated with palms and coconut trees in the Neotropics. Check List, 13 (2), 2086.
    https://doi.org/10.15560/13.2.2086

    Rolston, L.H. (1974) Revision of the genus Euschistus in Middle America (Hemiprtera: Pentatomidae: Pentatomini). Entomologica Americana, 48 (1), 1–102. [http://biodiversitylibrary.org/page/50638954]

    Rolston, L.H. (1975) A New Species and Review of Sibaria (Hemiptera: Pentatomidae). Journal of the New York Entomological Society, 83 (4), 218–225. [http://biodiversitylibrary.org/page/50819908]

    Rolston, L.H. (1978a) A revision of the genus Odmalea Bergroth (Hemiptera: Pentatomidae). Journal of The New York Entomological Society, 86 (1), 20–36. [http://biodiversitylibrary.org/page/52171346]

    Rolston, L.H. (1978b) A new subgenus of Euschistus (Hemiptera: Pentatomidae). Journal of the New York Entomological Society, 86 (2), 102–120. [http://biodiversitylibrary.org/part/214345]

    Rolston, L.H. (1978c) A revision of the genus Mormidea (Hemiptera: Pentatomidae). Journal of the New York Entomological Society, 86 (3), 161–219. [http://biodiversitylibrary.org/page/52171487]

    Rolston, L.H. (1982) A revision of Euschistus Dallas subgenus Lycipta Stål (Hemiptera: Pentatomidae). Proceedings of the Entomological Society of Washington, 84, 281–296.

    Rolston, L.H. (1983a) A Revision of the Genus Lincus Stål (Hemiptera: Pentatomidae: Discocephalinae: Ochlerini) Journal of the New York Entomological Society, 91 (1), 1–47.

    Rolston, L.H. (1983b) The genus Paralincus (Hemiptera: Pentatomidae). Journal of the New York Entomological Society, 91 (2), 183–187.

    Rolston, L.H. (1984) Key to the Males of the Nominate Subgenus of Euschistus in South America, with Descriptions of Three New Species (Hemiptera: Pentatomidae). Journal of the New York Entomological Society, 92 (4), 352–364.

    Rolston, L.H. (1987) Two new genera and species of Pentatomini from Peru and Brazil (Hemiptera: Pentatomidae). Journal of the New York Entomological Society, 95 (1), 62–68. [http://biodiversitylibrary.org/page/50767384]

    Rolston, L.H. (1988) The genus Ablaptus Stål (Pentatomidae: Discocephalinae: Discocephalini). Journal of the New York Entomological Society, 96 (3), 284–290. [http://biodiversitylibrary.org/page/50779467]

    Rolston, L.H. (1990) Key and diagnoses for the genera of “broadheaded” discocephalines (Hemiptera: Pentatomidae). Journal of the New York Entomological Society, 98 (1), 14–31.

    Rolston, L.H. (1992) Key and diagnoses for the genera of Ochlerini (Hemiptera: Pentatomidae: Discocephalinae). Journal of the New York Entomological Society, 100 (1), 1–41.

    Rolston, L.H. & McDonald, F.J.D. (1981) Conspectus of the Pentatomini genera of the Western Hemisphere. Part 2 (Hemiptera: Pentatomidae). Journal of the New York Entomological Society, 88, 257–272.

    Rolston, L.H., McDonald, F.J.D. & Thomas Junior, D.B. (1980) A conspectus of Pentatomini genera of the Western Hemisphere. Part I (Hemiptera: Pentatomidae). Journal of the New York Entomological Society, 88 (2), 120–132.

    Ruckes, H. (1965) Several new genera and species of Discocephaline Pentatomids (Heteroptera: Pentatomidae). Journal of the New York Entomological Society, 73 (3), 114–134.

    Ruckes, H. (1966a) The Genus Eurystethus Mayr, with the Descriptions of New Species (Heteroptera, Pentatomidae, Discocephalinae). American Museum Novitates, 2254 (18), 1–38.

    Ruckes H. (1966b) An analysis and a breakdown of the genus Platycarenus Fieber (Heteroptera, Pentatomidae, Discocephalinae). American Museum Novitates, 2255, 1–42.

    Ruckes, H. (1966c) An analysis and a breakdown of the genus Platycarenus Fieber (Heteroptera, Pentatomidae, Discocephallinae). American Museum Novitates, 2255 (18), 1–42.

    Ruckes, H. (1966d) The Genus Dryptocephala Laporter (Heteroptera, Pentatomidae, Discocephalinae). American Museum Novitates, 2256 (18), 1–32.

    Ruckes, H. (1966e) Review of the Bug Genus Pelidnocoris Stål (Heteroptera, Pentatomidae, Discocephalinae). American Museum Novitates, 2257, 18, 1–8.

    Ruckes, H. & Becker, M. (1970) The Bug Genus Coriplatus White (Heteroptera, Pentatomidae, Discocephalinae). American Museum Novitates, 2409 (25), 1–11.

    Santos, G. de & Bastardo, R. (2013) La familia Pentatomidae (Hemiptera: Heteroptera) en las colleciones de referencia de la República Dominica. Novitates Caribaea, 6, 1–15.

    Santos, B.T.S., Silva, V.J. & Fernandes, J. (2015) A Revison of Ascra with proposition of the bifida species group and description of two new species (Hemiptera: Pentatomidae: Edessinae). Zootaxa, 4303 (3), 445–470.
    https://doi.org/10.11646/zootaxa.4034.3.2

    Schmidt, L.S. & Barcellos, A. (2007) Abundância e riqueza de espécies de Heteroptera (Hemiptera) do Parque Estadual do Turvo, sul do Brasil: Pentatomoidea. Iheringia, Série Zoologia, 97 (1), 73–79.
    https://doi.org/10.1590/S0073-47212007000100011

    Schuh, R.T. & Slater, J.A. (1995) True bugs of the world (Hemiptera: Heteroptera). Classification and natural history. Cornell University Press, Ithaca, New York, xii + 336 pp.

    Schwertner, C.F. & Grazia, J. (2006) Descrição de seis espécies de Chinavia (Hemiptera, Pentatomidae, Pentatominae) da América do Sul. Iheringia, Serie Zoologia, 96 (2), 237–248.
    https://doi.org/10.1590/S0073-47212006000200015

    Schwertner, C.F. & Grazia, J. (2007) O gênero Chinavia Orian (Hemiptera, Pentatomidae, Pentatominae) no Brasil, com chave pictórica para os adultos. Revista Brasileira de Entomologia, 51 (4), 416–435.
    https://doi.org/10.1590/S0085-56262007000400005

    Schwertner, C.F. & Grazia, J. (2012) Review of the Neotropical genus Aleixus Mcdonald (Hemiptera: Heteroptera: Pentatomidae: Procleticini), with description of a new species and cladistic analysis of the tribe Procleticini. Entomologica Americana, 118 (1–4), 252–262.
    https://doi.org/10.1664/12-RA-034.1

    Segarra-Carmona, A.E., Franqui, R.A., Pérez-Martínez, H. & Morales-Grajales. (2015) New records for stink bugs (Heteroptera: Pentatomidae) in Puerto Rico. The Journal of Agriculture of University of Puerto Rico, 99 (2), 167–178.

    Silva, E.J.E., Fernandes, J.A.M. & Grazia, J. (2004) Variações morfológicas em Edessa rufomarginata e revalidação de E. albomarginata e E. marginalis (Heteroptera, Pentatomidae, Edessinae). Iheringia, Série Zoologia, 94 (3), 261–268.
    https://doi.org/10.1590/S0073-47212004000300006

    Silva, E.J.E., Fernandes, J.A.M. & Grazia, J. (2006) Caracterização do grupo Edessa rufomarginata e descrição de setes novas espécies (Heteroptera, Pentatomidae, Edessinae). Iheringia, Série Zoologia, 96 (3), 345–362.
    https://doi.org/10.1590/S0073-47212006000300012

    Silva, V.J. & Fernandes, J.A.M. (2012) A new species group in Edessa Fabricius, 1803 (Heteroptera: Pentatomidae: Edessinae). Zootaxa, 3313, 12–22.

    Silva, V.J., Nunes, B.M. & Fernandes, J.A.M. (2013) Paraedessa, a new genus of Edessinae (Hemiptera: Heteroptera: Pentatomidae). Zootaxa, 3716 (3), 395–416.
    https://doi.org/10.11646/zootaxa.3716.3.4

    Silva, V.J., Rider, D. & Fernandes, J.A.M. (2017) Reevaluation of the type species and redescription of Edessa (Heteroptera: Pentatomida: Edessinae). Zootaxa, 4347 (2), 255–274.
    https://doi.org/10.11646/zootaxa.4347.2.3

    Simões, F.L. & Campos, L.A. (2015) Breddin’s types of Ochlerus (Hemiptera, Pentatomidae, Discocephalinae). Contributions to Entomology: Beiträge zur Entomologie, 65 (2), 213–222.

    Smaniotto, L.F. & Panizzi, A.R. (2015) Interactions of selected species of stink bugs (Hemiptera: Heteroptera: Pentatomidae) from leguminous crops with plants in the Neotropics. Florida Entomologist, 98 (1), 7–17.
    https://doi.org/10.1653/024.098.0103

    Smithsonian Institution (2010) Smithsonian Tropical Research Institute. Available from: http://biogeodb.stri.si.edu/bioinformatics/dfm/metas/view/39026 (accessed 31 July 2017)

    Stål, C. (1872) Enumeratio Hemipterorum. Bidrag till em förteckning öfver alla kittilskånda Hemiptera, Jemte systematiska meddelanden. 2. Kongliga Svenska Vetenskaps—kademien Handlingar, Neue Folge, 10 (4), 1–159.

    Thomas D.B (1985) Revision of the Genus Chlorocoris Spinola (Hemiptera: Pentatomidae). Annals of the Entomological Society of America, 78 (5), 674–690.
    https://doi.org/10.1093/aesa/78.5.674

    Thomas D.B. (1992) Taxonomic synopsis of the Asopine Pentatomidae (Heteroptera) of the Western Hemisphere. The Thomas Say Foundation Series. Vol. 16. Entomological Society of America, Lanham, 156 pp.

    Thomas, D.B. (2000) Pentatomidae (Hemiptera). In: Llorente, J.E., Gonzalez, E. & Papavero, N. (Eds.), Biodiversidad, Taxonomia y Biogeografia de artrópodos de Mexico: hacia un sintesis de su conocimiento. Vol. 2. Universidad Nacional Autonoma de Mexico, Mexico D.F., pp. 335–352.

    Thomas, D.B. & Brailovsky, H. (1993) The genus Tibilis Stål in Mexico (Heteroptera: Pentatomidae). Pan-Pacific Entomologist, 69, 199–204.

    Thomas, D.B. & Yonke, T.R. (1988) Review of the genus Banasa Stål, 1860 (Hemiptera: Pentatomidae) for Mexico, Central America, and the Antilles. Annals of the Entomological Society of America, 81, 28–49.
    https://doi.org/10.1093/aesa/81.1.28

    Thomas, D.B. & Yonke, T.R. (1990) Review of the genus Banasa (Hemiptera: Pentatomidae) in South America. Annals of the Entomological Society of America, 83, 657–688.
    https://doi.org/10.1093/aesa/83.4.657

    Vivian, L.M. & Panizzi, A.R. (2006) Geographical distribution of genetically determined types of Nezara viriula (L.) (Heteroptera: Pentatomidae) in Brazil. Neotropical Entomology, 35 (2), 175–181.
    https://doi.org/10.1590/S1519-566X2006000200004

    Walker, F. (1868) Catalogue of the specimens of Heteropterus Hemiptera in the collection of the British Museum. Part III. British Museum, London, 180 pp.

    Weiler, L., Bianchi, F.M. & Grazia, J. (2012) Percevejos-do-mato (Hemiptera, Heteroptera, Pentatomoidea) da Floresta Nacional de São Francisco de Paula, RS, Brazil. Revista Brasileira de Biociências, 10 (2), 186–191.

    Weiler, L., Ferrari, A. & Grazia, J. (2016) Phylogeny and biogeography of the South American subgenus Euschistus (Lycipta) Stål (Heteroptera: Pentatomidae: Carpocorini). Insect Systematics & Evolution, 47 (4), 1–34.
    https://doi.org/10.1163/1876312X-47032145

    Westwood, J.O. (1837) A Catalogue of Hemiptera in the collection of the Rev. F, W. Hope, M. A. with short Latin descriptions of the new species. J. C. Bridgewater, London, 46 pp.

    Zwetsch, A. & Grazia, J. (2001) Revisão do gênero Runibia (Heteroptera, Pentatomidae, Pentatomini). Iheringia, Série Zoologia, 91, 5–28.
    https://doi.org/10.1590/S0073-47212001000200001