Skip to main content Skip to main navigation menu Skip to site footer
Type: Article
Published: 2018-03-21
Page range: 261–271
Abstract views: 65
PDF downloaded: 2

Systematics of the Satyrium (Superflua) deria (Moore, 1865) species group with description of a new species from Afghanistan (Lepidoptera, Lycaenidae)

Department of Entomology, Faculty of Biology, Lomonosov Moscow State University, Leninskie gory, 1–12, Moscow 119991, Russia Severtsov Institute of Ecology and Evolution, Russian Academy of Sciences, Leninsky pr., 33, Moscow, 119071, Russia
Schmalhausen Institute of Zoology, B. Khmelnitsky Str., 15, Kiev, 01601, Ukraine
R. Luxemburg Str., 21-5, Donetsk, 83050, Ukraine
Lepidoptera Theclinae Eumaeini hairstreaks morphology genitalia taxonomy zoogeography Hindu Kush Palaearctic


Satyrium (Superflua) deria (Moore, 1865) species group is revised. The group contains four species: S. (S.) deria (northern Himalayan region and southwestern Pamirs), S. (S.) muksuria Churkin & Pletnev, 2010 (southwestern Alay Mountains and Peter the Great Range), S. (S.) zabirovi Churkin & Pletnev, 2010 (from northwestern Pamirs to Darvaz Range), and S. (S.) hazarajatica sp. n. (type locality: central Afghanistan, Bamyan Province, Band-e Amir valley, env. Yakawlang). The new species differs from the other members of the group in the male and female genitalia, wing pattern, and colouration. Female genitalia of the S. (S.) deria, S. (S.) muksuria and S. (S.) zabirovi are illustrated and discussed for the first time. S. (S.) muksuria is newly recorded from the Alay Mountains. A key to species of the S. (S.) deria group is provided.



  1. Braby, M.F., Eastwood, R., Murray, N. (2012) The subspecies concept in butterflies: has its application in taxonomy and conservation biology outlived its usefulness? // Biological Journal of the Linnean Society, 106 (4), 699–716.

    Charmeux, J-F. (2004) Une nouvelle espèce du sous-genre Superflua du Pakistan (Lepidoptera: Lycaenidae). Phegea, 32 (1), 9–16.

    Churkin, S.V. & Pletnev, V.A. (2010) A review of the Satyrium (Superflua) sassanides (Kollar, 1849) species group from “Russian” Central Asia (Lepidoptera, Lycaenidae). Atalanta, 41 (1/2), 207–220.

    Eckweiler, W. & ten Hagen, W. (2003) Eine neue Art des Subgenus Superflua Strand, 1910 aus Südiran. Nachrichten des Entomologischen Vereins Apollo, 23, 213–216.

    Huang, H. & Zhu, J-Q. (2016) Ahlbergia maoweiweii sp. n. from Shaanxi, China with revisional notes on similar species (Lepidoptera: Lycaenidae). Zootaxa, 4114 (4), 409–433
    . 10.11646/zootaxa.4114.4.3

    Korb, S.K. (2014) New and little known blues (Lepidoptera, Lycaenidae) from Middle Asia and Kazakhstan. Eversmannia, 40, 11–17. [in Russian with English summary]

    Krupitsky, A.V., Pljushtch, I.G. & Pak, O.V. (2015a) Taxonomic notes on the genus Satyrium Scudder, 1876 (Lepidoptera, Lycaenidae) of Afghanistan with description of two new taxa. Zootaxa, 3985 (3), 421–431.

    Krupitsky, A.V., Pljushtch I.G. & Pak, O.V. (2015b) A new species of the Callophrys paulae Pfeiffer, 1932 species group from Afghanistan (Lepidoptera, Lycaenidae). Zootaxa, 4027 (2), 281–286.

    Kryzhanovsky, O.L. (2002) Structure and distribution of the entomofauna of the world. KMK Press, Moscow, 237 pp. [in Russian]

    Lukhtanov, V.A. (1995) Eine Übersicht über die Arten der Untergattung Satyrium (Superflua Strand, 1910) mit der Beschreibung einer neuen Art aus Südwestgissar, Usbekistan (Lepidoptera: Lycaenidae, Theclinae). Nachrichten des Entomologischen Vereins Apollo, 16 (1), 47–58

    Miller, L.D. (1969) Nomenclature of wing weins and cells. Journal of Research on the Lepidoptera, 8 (2), 37–48.

    Moore, F. (1865) List of Diurnal Lepidoptera collected by Capt. A.M. Lang in the N. W. Himalayas. Proceedings, Zoological Society of London, 2, 485–511.

    Prieto, C. & Vargas, M.A. (2016) Elfin butterflies of the genus Rhamma Johnson (Lepidoptera: Lycaenidae: Theclinae): A review of the Colombian species. Zootaxa, 4093 (3), 323–342.

    Shorthouse, D.P. (2010) SimpleMappr, an online tool to produce publication-quality point maps. Available from: (accessed 10 December 2017)

    Sidhu, A.K. (2009) Taxonomic notes on the Satyrium sassanides (Kollar) (Lycaenidae: Theclinae: Eumaeini) from north-west India. Records of the Zoological Surveys of India, 109 (2), 55–59.

    Weidenhoffer, Z., Bozano, G.C., Zhdanko, A. & Churkin, S. (2004) Guide to the butterflies of the Palearctic Region. Lycaenidae part II. Omnes Artes, Milano, 94 pp.

    Weidenhoffer, Z., Bozano, G.C., Zhdanko, A. & Churkin, S. (2016) Guide to the butterflies of the Palearctic Region. Lycaenidae part II. Second edition. Omnes Artes, Milano, 106 pp.

    Zhdanko, A.B. (2000) Genus Superflua. In: Tuzov, V.K., Bogdanov, P.V., Churkin, S.V., Dantchenko, A.V., Devyatkin, A.L., Murzin, V.S., Samodurov, G.D. & Zhdanko, A.B. (Eds.), Guide to the butterflies of Russia and adjacent territories (Lepidoptera, Lycaenidae). Vol. 2. Libytheidae, Danaidae, Nymphalidae, Riodinidae, Lycaenidae. Pensoft, Sofia-Moscow, pp. 103–104.