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Published: 2017-06-26

New species and stages description of Ulmeritoides Traver, 1959 (Ephemeroptera: Leptophlebiidae) from Roraima State, Northern Brazil

Universidade Federal de Roraima (UFRR), Campus Paricarana, Centro de Estudos da Biodiversidade (Cbio), CEP 69310-000, Boa Vista, Roraima, Brazil.
Universidade Estadual do Piauí, Campus Heróis do Jenipapo, Laboratório de Biodiversidade, Cep 64280-000, Campo Maior, PI, Brazil.
Ephemeroptera Taxonomy Aquatic Insect Atalophlebiinae Mayflies Amazon Rainforest


A new species of Ulmeritoides Traver, 1959 is described based on all life stages from Roraima State, Northern Brazil. U. amajari sp. nov. has a forewing coloration and penis shape similar to that of the widespread species U. uruguayensis (Traver) but it differs from the latter by the shape of the penis lobe apex in the male imago and by the lines of pectinate setae on the ventral surface of tibia III, in the nymph. The descriptions of nymph and female imago of U. passorum Gama-Neto & Hamada, 2014 are also presented.



  1. Boldrini, R. & Cruz, P.V. (2013) Criação e Transporte de Ninfas de Ephemeroptera (Insecta) em Campo. EntomoBrasilis, 6 (2), 168–170.

    Coleman, C.O. (2003) “Digital inking”: How to make perfect line drawings on computers. Organisms Diversity & Evolution, Electronic Supplement, 1–14.

    Coleman, C.O. (2006) Substituting time-consuming pencil drawings in arthropod taxonomy using stacks of digital photographs. Zootaxa, 1360, 61–68.

    Domínguez, E. (1991) The status of the genus Ulmeritus (Ephemeroptera: Leptophlebiidae: Atalophlebiinae) and related taxa. In: Alba-Tercedor, J. & Sánchez-Ortega, A. (Eds.), Overview and strategies of Ephemeroptera and Plecoptera. Sandhill Crane Press, Gainesville, pp. 157–167.

    Domínguez, E. (1995) Cladistic analysis of the Ulmeritus-Ulmeritoides group (Ephemeroptera, Leptophlebiidae), with descriptions of five new species of Ulmeritoides. Journal of the New York Entomological Society, 103, 15–38.

    Domínguez, E., Molineri, C., Pescador, M.L., Hubbard, M.D. & Nieto, C. (2006) Ephemeroptera of South America. In: Adis, J., Arias, J.R., Rueda-Delgado, G. &Wantzen, K.M. (Eds.), Aquatic Biodiversity of Latin America. Vol. 2. Pensoft, Moscow-sofia, pp. 1–646.

    Gama Neto, J.L. & Hamada, N. (2014) Leptophlebiidae (Ephemeroptera) of the Serra do Tepequém, Roraima State, Brazil: new records and description of two new species. Zootaxa, 3900 (2), 279–286.

    Gama Neto, J.L. & Passos, M.A.B. (2016) Leptophlebiidae (Ephemeroptera) from Roraima State, Brazil: description of two new species of Ulmeritoides Domínguez, 1991. Zootaxa, 4178 (2), 295–300.

    Kluge, N.J. (1994) Pterothorax structure of mayflies (Ephemeroptera) and its use in systematics. Bulletin de la Société entomologique de France, 99 (1), 41–61.

    Salles, F.F. & Domínguez, E. (2012) Systematics and Phylogeny of Ulmeritus-Ulmeritoides revisited (Ephemeroptera: Leptophlebiidae). Zootaxa, 3571, 49–65.

    Salles, F.F., Boldrini R., Nascimento, J.M.C., Angeli, K.B., Massariol, F.C. & Raimundi, E. (2016) Ephemeroptera do Brasil. [Internet], Aug 2016. Available from: (accessed 29 May 2017)

    Souto, P.M., Da-Silva, E.R., Nessimian, J.L. & Gonçalves, I.C. (2016) Two new species of Ulmeritoides Traver (Ephemeroptera: Leptophlebiidae) from Southeastern Brazil. Zootaxa, 4078 (1), 127–136.

    Traver, J.R. (1959) Uruguayan mayflies. Family Leptophlebiidae: Part I. Revista de la Sociedad Uruguaya de Entomologia, 3, 1–13.