Abstract
The generic name Lanchnophorus Reuter, 1887, deemed for a long time to be unavailable as incorrect original spelling of Lachnophorus (in fact Lachnophorus Distant, 1903 is an unjustified emendation of the former), is restored as a valid name of the genus. Lachnesthus Bergroth, 1915, syn. nov. (new name for the preoccupied Lachnophorus Distant, 1903) is considered junior synonym of Lanchnophorus. The following nomenclatural changes are proposed: Lanchnophorus flavus (Scudder, 1971) comb. nov. = Lachnesthus chinai Scudder, nomen nudum; Lanchnophorus guttulatus Reuter, 1887, comb. restit. = Lachnophorus albidomaculatus Distant, 1913, syn. nov. = Lachnesthus rodriguezensis China, 1925, syn. nov.; Lanchnophorus leucospilus (Walker, 1872) comb. nov.; Lanchnophorus merula (Distant, 1903) comb. nov.; and Lanchnophorus singalensis (Dohrn, 1860) comb. nov. Three new species are described: Lanchnophorus gaoqingae Kment & Jindra sp. nov. from China (Yunnan), Lanchnophorus seminitens Kment & Carapezza sp. nov. from Socotra Island (Yemen), and Lanchnophorus webbi Kondorosy sp. nov. from India: Tamil Nadu. Bibliographies and known distribution of all the included species are reviewed. The following new country and state records are provided: L. flavus from Central African Republic, Ethiopia, Ghana, Mali, Malawi, Niger, Zambia and Zimbabwe; L. leucospilus from China (Yunnan) and Laos, L. merula from India (Kerala/Tamil Nadu) and Thailand; L. singalensis from Angola, Benin, Mozambique, Namibia, Senegal, Sierra Leone, Tanzania, Togo, Uganda, Zambia, Zimbabwe, China (Hainan), Iran (Sistan and Ba-luchestan), Oman, Pakistan, India (Himachal Pradesh, Karnataka, Kerala, Rajasthan), Malaysia, Philippines, and Thailand.
References
Al-Houty, W. (2011) Recent insect fauna recorded in Kuwait. Kuwait Journal of Science and Engineering, 38 (1a), 81–92.
Atkinson, E.T. (1882) Notes on the zoology of the N.W.P. India. E.T. Atkinson, St. Leonard’s-on-Sea, 266 pp.
Aukema, B., Rieger, Ch. & Rabitsch, W. (2013) Catalogue of the Heteroptera of the Palaearctic Region. VI. Supplement. The Netherlands Entomological Society, Amsterdam, xxiii + 629 pp.
Báez, M., Heiss, E., García, A. & Cabrera, A. (2005) Hemiptera: Heteroptera. In: Archavaleta, M., Zurita, N., Marrero, M.C. & Martín, J.L. (Eds), Lista preliminar de especies silvestres de Cabo Verde (hongos, plantas y animales terrestres). Consejería de Medio Ambiente y Ordenación Territorial, Gobierno de Canarias, pp. 71–78.
Bergroth, E. (1915) Hemiptera from the Bombay Presidency. Journal of the Bombay Natural History Society, 24 (1), 170–179.
Bezděk, J., Purchart, L., Král, K. & Hula, V. (2012) List of local Socotran geographical names used in entomological literature, pp. 27–67. In Hájek, J. & Bezděk, J. (eds): Insect biodiversity of the Socotra Archipelago. Acta Entomologica Musei Nationalis Pragae 52 (supplementum 2), i–vi + 1–557.
Bhasin, G.D. (1953) Part. 15.—Order Hemiptera (continued). Families Coreidae, Berytidae, Lygaeidae, Pyrrhocoridae and Aradidae. In: Roomwal, M.L., Mathur, R.N., Bhasin, G.D., Singh, B., Pant, G.D., Chandra, A., Katiyar, K.N. & Mathur, R.K.: A systematic catalogue of the main identified entomological collection at the Forest Research Institute, Dehra Dun. Parts 9–21.—Orders Isoptera, Embioptera, Psocoptera, Anoplura, Ephemeroptera, Odontata, Thysanoptera and Hemiptera. Indian Forest Leaflet, 121 (3), pp. 102–117.
Bonhag, P.F. & Wick, J.R. (1953) The functional anatomy of the male and female reproductive systems of milkweed bug, Oncopeltus fasciatus (Dallas) (Heteroptera: Lygaeidae). Journal of Morphology, 93, 177–283.
https://doi.org/10.1002/jmor.1050930202Breddin, G. (1907) Berytiden und Myodochiden von Ceylon aus der Sammelausbeute von Dr. W. Horn (Rhynch. het. [sic!]). Deutsche Entomologische Zeitschrift, 1907, 203–220.
Chatterjee, N.C. (1937) Entomological investigations on the spike disease of sandal (32). Lygaeidae (Hemipt.). Indian Forest Records, 3, 106–121.
China, W.E. (1924) The Hemiptera-Heteroptera of Rodriguez, together with the description of a new species of Cicada from that Island. Annals and Magazine of Natural History, Series 9, 15, 427–453.
China, W.E. (1925) Additional notes on the Heteroptera of Rodriguez. Annals and Magazine of Natural History, Series 9, 15, 163–165.
https://doi.org/10.1080/00222932508633195Ðang, D.K. & Nguyen, T.T.H. (2009) Ket qua nghien cuu ve thanh phan loai cu abo box it (Heteroptera) o vuon quoc gia Phong Nha—Ke Bang, tinh Quang Binh. (Study results on species composition of bugs (Heteroptera) in Phong Nha—Ke Bang National Park, Quang Binh Province). Tap Chi Sinh Hoc—Journal of Biology, 31(3), 41–45. [in Vietnamese, English summary]
Dejean, P.F.M.A. (1831) Species général des coléoptères, de la Collection de M. le Comte Dejean. Tome cinquième. Méquignon-Marvis, Paris, viii + 833 pp.
Distant, W.L. (1901) Rhynchotal notes.- XI. Heteroptera: Fam. Lygaeidae. Annals and Magazine of Natural History, Series 7, 8, 464–486 + 497–510.
Distant, W.L. (1903) Rhynchota.–Vol. II. (Heteroptera), pp. 1–242. In: Blanford, W.T. (Ed.), The Fauna of British India, including Ceylon and Burma. Taylor & Francis, London, i–xvii, 503 pp. [Published in two parts: pp. 1–242. (December 1903), pp. 243–503 (April 1904)–see the Preface, p. iii.]
Distant, W.L. (1913) Rhynchota. Part I: Suborder Heteroptera. In: Reports of the Percy Sladen Trust Expedition to the Indian Ocean in 1905, under the leadership of Mr J. Stanley Gardiner, M.A. Volume V. Transactions of the Linnean Society of London, Series 2, 16, 139–191 + plates 11–13.
Distant, W.L. (1918) Contributions to a further knowledge of the Rhynchotal family Lygaeidae. Annals and Magazine of Natural History, Series 9, 2, 257–270.
Dohrn, A. (1860) Zur Heteropteren-Fauna Ceylon’s. Stettiner Entomologische Zeitung, 21, 399–409.
Gerlach, J. (2013) Odonata, Hemiptera, Hymenoptera and other insects of the Seychelles islands. Siri Scientific Press, Manchester, 394 pp + 4 plates.
Goel, S.C. (1970) Taxonomic significance of the metathoracic wing-venation in the landbugs (Heteroptera). Bulletin of Entomology, 11 (1), 1–10.
Hoberlandt, L. (1954) Contribution à l’étude de la zone d’inondation du Niger (Mission G. Remaudière). XI.—Hémiptères-Hétéroptères. Bulletin de l’Institut Français d’Afrique Noire, 16, 917–934.
Hoberlandt, L. (1961) Ergebnisse der Deutschen Afghanistan-Expedition 1956 der Landessammlungen für Naturkunde in Karlsruhe. Beiträge zur Naturkundlichen Forschung in Südwestdeutschland, 19 (3), 197–222.
ICZN (International Commission on Zoological Nomenclature) (1999) International Code of Zoological Nomenclature. Fourth edition. International Trust for Zoological Nomenclature, London, 306 pp.
Kirby, W.F. (1891) Catalogue of the described Hemiptera Heteroptera and Homoptera of Ceylon, based on the collection formed (chiefly at Pundaluoya) by Me. E. Ernest Green. Journal of the Linnean Society of London, 24, 72–176.
https://doi.org/10.1111/j.1096-3642.1891.tb02479.xKment, P., Hemala, V. & Baňař, P. (2016) Rhyparoclava pyrrhocoroides, a new genus and species of autapomorphic Rhyparochromidae with clavate antennae from Madagascar (Hemiptera: Heteroptera). Acta Entomologica Musei Nationalis Pragae, 56, 517–545.
Krüger, A. & Deckert, J. (2016) True bugs (Hemiptera-Heteroptera) of Botswana—Bibliographical inventory and new records. Zootaxa, 4114 (1), 33–63.
https://doi.org/10.11646/zootaxa.4114.1.2Lafont, A. (1910) Sur la présence d’un Leptomonas, parasite de la classe des Flagellés, dans le latex de trois Euphorbiacées. Annales de l’Institut Pasteur, 24, 205–219.
LePelley, R.H. (1960) Agricultural insects of East Africa. East Africa High Commission, Nairobi, 307 pp.
Lethierry, L. & Severin, G. (1894) Cataloque général des Hémiptères. Tome II, Hétéroptères, Coreidae, Berytidae, Lygaeidae, Pyrrhocoridae. Musée royal d’histoire naturelle de Belgique, Bruxelles, 277 pp.
Lindberg, H. (1959) Hemiptera Insularum Caboverdensium. Systematik, Ökologie und Verbreitung der Heteropteren und Cicadinen der Kapverdischen Inseln. In: Lindberg, H. (ed.): Ergebnisse der Zoologischen Expedition von Professor Håkan Lindberg nach den Kapverde-Inseln im Winter 1953–54. Commentationes Biologicae, 19 (1) [1958], 1–246.
Linnavuori, R.E. (1978) Hemiptera of the Sudan, with remarks on some species of the adjacent countries 6. Aradidae, Meziridae, Aneuridae, Pyrrhocoridae, Stenocephalidae, Coreidae, Alydidae, Rhopalidae, Lygaeidae. Acta Zoologica Fennica, 153, 1–108.
Linnavuori, R.E. (2004) Heteroptera of the Hormozgan province in Iran. III. Cimicomorpha (Anthocoridae, Cantacaderidae, Tingidae), Pentatomorpha [sic!]. Acta Universitatis Carolinae, Biologica, 48, 273–286.
Linnavuori, R.E. (2011) Studies on the Cimicomorpha and Pentatomomorpha (Hemiptera: Heteroptera) of Khuzestan and the adjacent provinces of Iran. Acta Entomologica Musei Nationalis Pragae, 51, 21–48.
Linnavuori, R.E. & Harten, A. van (2000) Additional notes on the Heteroptera (Insecta: Hemiptera) of Yemen. Fauna of Arabia, 18, 165–175.
Linnavuori, R.E. & Harten, A. van (2002) Additional notes on the Heteroptera (Insecta: Hemiptera) of Yemen II. Esperiana, Buchreihe zur Entomologie, 9, 157−188.
Linnavuori, R.E. & Harten, A. van (2006) Additional notes on the Heteroptera (Insecta: Hemiptera) of Yemen IV. Pentatomomorpha. Acta Universitatis Carolinae, Biologica, 49 [2005], 161–174.
Linnavuori, R.E., Kment, P. & Carapezza, A. (2014) Order Hemiptera, suborder Heteroptera. Infraorder Pentatomomorpha, superfamily Lygaeoidea, pp. 192–246. In: Harten, A. van (Ed.), Arthropod fauna of the United Arab Emirates. Vol. 5. Multiply Marketing Consultancy Services, Abu Dhabi, 744 pp.
Maxwell-Lefroy, H. (1909a) The insect fauna of Tirhut. I.—Rhynchota Heteroptera. Records of the Indian Museum, 3, 301–338.
Maxwell-Lefroy, H. (1909b) Indian insect life. Thacker, Spink & Co., Calcutta and Simla, xii + 786 pp.
Medler, J.T. (1980) Insects of Nigeria—check list and bibliography. Memoirs of the American Entomological Institute, 30, i–vii + 1–919.
Mukhopadhyay, A. (1988) Taxonomic study of Lygaeidae (Heteroptera: Insecta) from West Bengal (India). Records of the Zoological Survey of India Occasional Paper, 107, 1–72 + 12 pls.
Nadkarni, Sh.N.T. (1975) Chapter 3 – Insects, pp. 238–458. In: Maharashtra state gazetteers. General series: Fauna. Government Printing, Stationary, Maharashtra State, Bombay, 707 pp + 5 pls.
Parshad, R. (1957a) Post-reductional m-chromosomes in the male Lanchnophorus singalensis Dohrn. (Lygaeidae-Heteroptera). Journal of Genetics, 55 (3), 503–506.
https://doi.org/10.1007/BF02984067Parshad, R. (1957b) Cytological studies in Heteroptera. IV. Chromosome complement and meiosis in twenty six species of the Pentatomoidea, Lygaeoidea and Coreoidea with a consideration of the cytological bearing on the status of these super-families. Research Bulletin of the Panjab University, Zoology, 133, 521–559.
Parshad, R. (1958) Structure of the Heteropteran kinetochore. The behaviour of the long chromosomes in some lygaeid and coreid bugs during mitosis and meiosis. Cytologia, 23 (1), 25–32.
https://doi.org/10.1508/cytologia.23.25Péricart, J. (2001) Family Lygaeidae Schilling, 1829—seed-bugs. In: Aukema, B. & Rieger, Ch. (Eds.), Catalogue of the Heteroptera of the Palaearctic Region. Volume 4. The Netherlands Entomological Society, Amsterdam, pp. 35–220.
Priesner, H. & Alfieri, A. (1953) A review of the Hemiptera-Heteroptera known to us from Egypt. Bulletin de la Société Fouad Ier de Entomologie, 37, 1–119.
Priesner, H. & Wagner, E. (1961) Supplement to “A review of the Hemiptera Heteroptera known to us from Egypt”. Bulletin de la Société Entomologique d’Egypt, 45, 323–339.
Reuter, O.M. (1887) Ad cognitionem Heteropterorum Madagascariensium. Entomologisk Tidskrift, 8, 77−109.
Schouteden, H. (1957) Contributions à l’étude de la faune entomologique du Ruanda-Urundi (Mission P. Basilewsky 1953). CXXV. Heteroptera Lygaeidae, Pyrrhocoridae et Berytidae. Annales du Musée du Congo Belge, Tervuren, in-8°, Zoologie, 58, 247–268.
Schumacher, F. (1913) Ein Beitrag zur Kenntnis der Rhynchoten-Fauna Südafrikas, insbesondere von Deutsch-Südwestafrika, Klein-Namaland und dem Kalaharigebiet, auf Grund der Expeditionsergebnisse von Herrn Professor L. Schultze und dem Material des Königl. Zoologischen Museums zu Berlin. In: Schultze, L. (Ed.), Zoologische und anthropologische Ergebnisse eine Forschungsreise in Südafrika. Bd. V. Denkschriften der Medizinisch-Naturwissenschaftlichen Gesellschaft zu Jena, 17, 49–88.
Scudder, G.G.E. (1957) The higher classification of the Rhyparochrominae (Hem., Lygaeidae). Entomologist’s Monthly Magazine, 93, 152−156.
Scudder, G.G.E. (1962). The World Rhyparochrominae (Hemiptera, Lygaeidae). I. New synonymy and generic changes. Canadian Entomologist, 94, 764−773.
Scudder, G.G.E. (1967) Rhyparochrominae types in the British Museum (Natural History) (Hemiptera: Lygaeidae). Bulletin of the British Museum (Natural History), Entomology, 20 (6), 253–285.
Scudder, G.G.E. (1968) Air-borne Lygaeidae (Hemiptera) trapped over the Atlantic, Indian and Pacific Oceans, with the description of a new species of Appolonius Distant. Pacific Insects, 10, 155–160.
Scudder, G.G.E. (1969) The Lygaeidae (Hemiptera) collected by Dr. M. Mrázek and Mrs. I. Korecká in the Republic of Guinea. Acta Musei Moraviae, Scieantiae Naturales, 54, 169−182.
Scudder, G.G.E. (1971) New Lygaeidae [Hemiptera] from the Niokolo-Koba National Park. Bulletin de l’Institut Français d’Afrique Noire, Série A, 33, 718−736.
Scudder, G.G.E. (1982) IX. Hemiptera Lygaeidae du Parc National du Niokolo-Koba. In: Recherches scientifiques dans les parcs nationaux du Sénégal. Mémoires de lʼInstitut Fondamental dʼAfrique Noire, 92, 141−154.
Slater, J.A. (1964a) A catalogue of the Lygaeidae of the World. Volume I & II. University of Connecticut, Storrs, i–xviii, 1–778 [I] & 779–1668 [II] pp.
Slater, J.A. (1964b) Hemiptera (Heteroptera): Lygaeidae, pp. 16−228. In: Hanström, B., Brinck, P. & Rudebeck, G. (Eds.), South African Animal Life. Vol. X. Swedish Natural History Research Council, Stockholm, 496 pp.
Slater, J.A. (1972) The Lygaeidae of Upemba National Park (Hemiptera: Heteroptera). In: Parc National de l’Upemba.—Mission G. F. de Witte en collaboration avec W. Adam, A. Janssens, L. van Meel et R. Verheyen (1946–1949). Fascicule 72. Institut des Parcs Nationaux du Congo belge, Bruxelles, pp. 17−81.
Slater, J.A. (1979) Hemiptera, Heteroptera: Lygaeidae from Sri Lanka (Ceylon). Report No. 45. Lund University Ceylon Expedition 1962, 27 pp. [Preprint, also published in Entomologica Scandinavica, Supplement, 11 (1982), 1−27.
Slater, J.A. & Lattin, J.D. (1965) Lachnestes singalensis (Dohrn), a lygaeid new to the Western Hemisphere (Hemiptera). Pan-Pacific Entomologist, 41, 58−60.
Slater, J.A. & O’Donnell, J.E. (1995) A catalogue of the Lygaeidae of the World (1960–1994). New York Entomological Society, New York, xv + 410 pp.
Stål, C. (1874) Enumeratio Hemipterorum. Bidrag till en förteckning öfver alla hittils kända Hemiptera, jemte systematiska meddelanden. 4. Kungliga Svenska Vetenskapsakademiens Handlingar, 12 (1), 1–186.
Suman, V. & Kaur, H. (2012) Meiotic studies in seven Heteropteran species. Cytologia, 77, 311–322.
https://doi.org/10.1508/cytologia.77.311Sweet, M.H. (1967) The tribal classification of the Rhyparochrominae (Heteroptera: Lygaeidae). Annals of the Entomological Society of America, 60, 208–226.
https://doi.org/10.1093/aesa/60.1.208Thangavelu, K. & Ananthasubramanian, K.S. (1981) Natural enemies of Rhyparochrominae (Lygaeidae: Heteroptera) from Southern India. Proceedings of Indian National Science Academy, B47 (5), 632–636.
Tsai, J.-F. & Rédei, D. (2014) A revision of the genus Amblycara (Hemiptera: Heteroptera: Pentatomidae). Acta Entomologica Musei Nationalis Pragae, 54, 133–155.
Tsai, J.-F. & Rédei, D. (in press) The genus Arocatus in Taiwan (Hemiptera: Heteroptera: Lygaeidae). Zootaxa.
Tsai, J.-F., Rédei, D., Yeh, G.-F. & Yang, M.-M. (2011) Jewel bugs of Taiwan (Heteroptera: Scutelleridae). National Chung Hsing University, Taichung, 309 pp.
Ueda, K. & Yoshiyasu, Y. (2001) A list of Japanese Insect Collection by P. F. von Siebold and H. Bürger preserved in Nationaal Natuurhistorisch Museum, Leiden, the Netherlands. Bulletin of the Kitakyushu Museum of Natural History, 20, 81–143.
Ueshima, N. & Ashlock, P.D. (1980) Cytotaxonomy of the Lygaeidae (Hemiptera-Heteroptera). University of Kansas Science Bulletin, 51 (26), 717–801.
https://doi.org/10.5962/bhl.part.3259Walker, F. (1872) Catalogue of the specimens of Hemiptera Heteroptera in the collection of the British Museum. Part V. Trustees of the British Museum, London, 202 pp.
Wilcox, D.B. & Slater, J.A. (1969) Contribution a la faune du Congo (Brazzaville). Mission A. Villiers et A. Descarpentries. LXXXVII. Hétéroptere Lygaeidae. Bulletin de l’Institute de Afrique Noire, Série A, 31, 686−701.
Zrzavý, J. (1990) Evolution in antennal sclerites in Heteroptera (Insecta). Acta Universitatis Carolinae Biologica, 34, 189–227.