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Published: 2016-09-07

Whip spiders (Amblypygi, Arachnida) of the Western Palaearctic—a review

Callistus – Gemeinschaft für Zoologische & Ökologische Untersuchungen [association for zoological and ecological research], Heidloh 8, 95503 Hummeltal, Germany.
Institute of Zoology, Department of Integrative Biology and Biodiversity Research, University of Natural Resources and Life Sciences, Gregor Mendel Strasse 33, 1180 Vienna, Austria.
Amblypygi Arachnida


All records of the two amblypygid species occurring in the Western Palaearctic are mapped and both species (Charinus ioanniticus and Musicodamon atlanteus) are discussed. Charinus ioanniticus is known from the Eastern Mediterranean (Greece, Turkey, Israel and Egypt) from 10 localities and Musicodamon atlanteus is known from the Maghreb (Morocco, Algeria) from three localities. All records are mapped.



  1. Armas, L.F. de (2000) Parthenogenesis in Amblypygi (Arachnida). Avicennia, 12/13, 133–134.

    Armas, L.F. de, Palomino-Cárdenas, A.C. & Castillo-Espinoza, M. del (2016) Amblypygids of Cusco and Madre de Dios departments, Peru, with the description of a new Charinus (Amblypygi: Charinidae, Phrynidae). Revista Ibérica de Aracnología, 28, 45–50.

    Blick, T. & Harvey, M.S. (2011) Worldwide catalogues and species numbers of the arachnid orders (Arachnida). Arachnologische Mitteilungen, 41, 41–43.

    Delle Cave, L (1986) Biospeleology of the Somaliland. Amblypygi (Arachnida, Chelicerata) of the caves of Showli Berdi and Mugdile (Bardera, Somaliland). Redia, 69, 143–170.

    deMenocal, P.B. & Tierney, J.E. (2012) Green Sahara: African humid periods paced by earth's orbital changes. Nature Education Knowledge, 3 (10), 12. Available from: (29 July 2016)

    El-Hennawy, H. (2002) The first record of Amblypygi from Egypt. Journal of Arachnology, 30, 452–453.[0452:TFROAF]2.0.CO;2

    Fage, L. (1939) Sur une phryne du Sud Marocain pourvue d'un appareil stridulent Musicodamon atlanteus, n. gen., nov. sp. Bulletin de la Société Zoologique de France, 64, 100–114.

    Fage, L. (1954) Remarques sur la distribution géographique des Pédipalpes Amblypyges africains, accompagnées de la description d'une espèce nouvelle de Madagascar: Charinus madagascariensis, nov. sp. Annales du Musée Royal du Congo Belge, Nouvelle Série 4, Sciences Zoologiques, 1, 180–184.

    Giupponi, A.P. de L. & Miranda, G.S. de (2016) Eight new species of Charinus Simon, 1892 (Arachnida: Amblypygi: Charinidae) endemic for the Brazilian Amazon, with notes on their conservational status. PLoS ONE, 11 (e0148277), 1–33.

    Guzmán, A.A., Joya, D.C. & Francke, O.F. (2015) The first troglomorphic species of the genus Phrynus Lamarck, 1801 (Amblypygi: Phrynidae) from Mexico. Zootaxa, 3920 (3), 474–482.

    Harvey, M.S. (2003) Catalogue of the smaller arachnid orders of the world: Amblypygi, Uropygi, Schizomida, Palpigradi, Ricinulei and Solifugae. CSIRO Publishing Huntingdon, Collingwood, Victoria, 385 pp.

    Harvey, M.S. (2013) Whip spiders of the world, version 1.0. Western Australian Museum, Perth. Available from: (31 May 2016)

    Jolly, D., Prentice, I.C., Bonnefille, R., Ballouche, A., Bengo, M., Brenac, P., Buchet, G., Burney, D., Cazet, J.-P., Cheddadi, R., Edorh, T., Elenga, H., Elmoutaki, S., Guiot, J., Laarif, F., Lamb, H., Lezine, A.-M., Maley, J., Mbenza, M., Peyron, O., Reille, M., Reynaud-Farrera, I., Riollet, G., Ritchie, J.C., Roche, E., Scott, L., Ssemmanda, I., Straka, H., Umer, M., Van Campo, E., Vilimumbalo, S., Vincens, A. & Waller, M. (1998) Biome reconstruction from pollen and plant macrofossil data for Africa and the Arabian peninsula at 0 and 6000 years. Journal of Biogeography, 25, 1007–1027.

    Kovařík, F. & Vlasta, D. (1996) First report of Amblypygi (Charinidae: Charinus ioanniticus) from Turkey. Klapalekiana, 32, 57–58.

    Kraus, O. (1961) Charontidae aus Israel, ein zoogeographisch bemerkenswertes Vorkommen (Arach., Pedipalpi-Amblypygi). Senckenbergiana biologica, 42, 491–493.

    Kritscher, E. (1959) Ergebnisse der von Dr. O. Paget und Dr. E. Kritscher auf Rhodos durchgeführten zoologischen Exkursionen. II. Pedipalpi (Amblypygi). Annalen des Naturhistorischen Museums Wien, 63, 453–457.

    Lawrence, R.F. (1969) A collection of African Amblypygi with keys to the subfamilies, genera and species of the Ethiopian fauna. Revue de zoologie et de botanique africaines, 80, 80–87.

    Miranda, G.S. de, Giupponi, A.P. de L. & Wizen, G. (2016) Two new species of whip spider (Amblypygi): an epigean and a cave dwelling Charinus Simon, 1892 from Belize. Zootaxa, 4098 (3), 545–559.

    Prendini, L., Weygoldt, P. & Wheeler, W.C. (2005) Systematics of the Damon variegatus group of African whip spiders (Chelicerata: Amblypygi): evidence from behaviour, morphology and DNA. Organisms, Diversity & Evolution, 5, 203–236.

    Prendini, L. (2011) Phylum Arthropoda von Siebold, 1848. In: Zhang, Z.-Q. (Ed.) Animal biodiversity: an outline of higher-level classification and survey of taxonomic richness. Zootaxa, 3148, pp. 154.

    Réveillion, F. & Maquart, P.O. (2015) A new species of Charinus Simon, 1892 (Amblypygi, Charinidae) from termite nests in French Guiana. Zootaxa, 4032 (2), 190–196.

    Rosin, R. & Shulov, A. (1960) Representatlves of the order Amblypygi (Arachnida) found in Israel. Bulletin of the research council of Israel, 9B, 167–168.

    Seiter, M. & Hörweg, C. (2013) The whip spider collection (Arachnida, Amblypygi) held in the Natural History Museum Vienna, Austria. Arachnologische Mitteilungen, 46, 47–53.

    Seiter, M. & Wolff, J. (2014) Description of Sarax buxtoni (Gravely, 1915) (Arachnida: Amblypygi: Charinidae) and a new case of parthenogenesis in Amblypygi from Singapore. Journal of Arachnology, 42, 233–239.

    Seiter, M., Wolff, J. & Hörweg, C. (2015) A new species of the South East Asian genus Sarax Simon, 1892 (Arachnida: Amblypygi: Charinidae) and synonymization of Sarax mediterraneus Delle Cave, 1986. Zootaxa, 4012 (3), 542–552.

    Seyyar, O. & Demir, H. (2007) A new locality for Charinus ioanniticus (Kritscher,1959) (Amblypygi: Charnidae). Serket, 10, 109–111.

    Teruel, R. (2016) The genus Charinus Simon, 1892 (Amblypygi: Charinidae) on the island of Hispaniola, Greater Antilles. Revista Ibérica de Aracnología, 28, 3–12.

    Torres-Contreras, R., García, D.M.Á. & Armas, L.F. de (2015) A new species of Charinus Simon, 1892 (Amblypygi: Charinidae) from the Colombian Caribbean. Revista Ibérica de Aracnología, 27, 145–148.

    Vasconcelos, A.C.O., Giupponi, A.P. de L. & Ferreira, R.P. (2016) Description of a new troglomorphic species of Charinus Simon, 1892 from Brazil (Arachnida, Amblypygi, Charinidae). ZooKeys, 600, 35–52.

    Weygoldt, P. (1972) Charontidae (Amblypygi) aus Brasilien. Beschreibung von zwei neuen Charinus-Arten, mit Anmerkungen zur Entwicklung, Morphologie und Tiergeographie und mit einem Bestimmungsschlüssel für die Gattung Charinus. Zoologische Jahrbücher, Abteilung für Systematik, Ökologie und Geographie der Tiere, 99, 107–132.

    Weygoldt, P. (1997) The relationships of the south east African whip spiders Hemiphrynus machadoi Fage, 1951 and Phrynichus scullyi Purcell, 1901: introduction of the new generic names Xerophrynus and Phrynichodamon (Chelicerata: Amblypygi). Zoologischer Anzeiger, 235, 117–130. [for 1996]

    Weygoldt, P. (2000a) Whip spiders. Their biology, morphology and systematics. Apollo Books, Stenstrup, 163 pp.

    Weygoldt, P. (2000b) African whip spiders. Synopsis of the Amblypygi reported from Africa (Arachnida). Memorie della Societé Entomologica Italiana, 78, 339–359.

    Weygoldt, P. (2005) Biogeography, systematic position, and reproduction of Charinus ioanniticus (Kritscher 1959) with the description of a new species from Pakistan (Chelicerata, Amblypygi, Charinidae). Senckenbergiana biologica, 85, 43–56.

    Weygoldt, P. (2007) Parthenogenesis and reproduction in Charinus ioanniticus (Kritscher, 1959) (Chelicerata, Amblypygi, Charinidae). Arachnology, 14, 81–82.