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Type: Monograph
Published: 2012-02-22
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Revision of Flabelligera Sars, 1829 (Polychaeta: Flabelligeridae)

Departamento de Ecología Acuática El Colegio de la Frontera Sur, CONACYT Apdo. Postal 424, Chetumal, Quintana Roo, 77000 MEXICO
Annelida Siphonostoma Pherusa hard-bottoms soft-bottoms

Abstract

Flabelligera Sars, 1829 is the type genus for Flabelligeridae de Saint-Joseph, 1894 and includes species common in tem-perate or cold waters in the world oceans. Together with Flabelliderma Hartman, 1969 they are the only genera whoseneurohooks have multiarticulate handles and bending crests. However, Flabelliderma species differ because they formlarge notopodial lobes, and often have dorsal sediment tubercles. The revision of all type and non-type materials resultedin the distinction of five body patterns within Flabelligera; they can be recognized on the basis of body shape, tunic de-velopment, parapodial position, notopodial alignment, and shape of neurohooks. Flabelligera is restricted and four generaare proposed: Annenkova (neurohooks with crests tapered, apparently segmented), Flabegraviera (notopodia in laterallydescending series, notochaetae and neurochaetae in cylindrical sheaths, markedly longer than body width), Flabehlersia(body fusiform, notopodia ventrolateral) and Flabesymbios (body papillae not covered by tunic, notopodia dorsal with papillae in fan-shaped pattern, neuropodia ventral). Flabelligera contains 16 species with three newly described and a re-placement name is proposed: F. haswelli n. n. pro F. affine Haswell, F. nuniezi n. sp. from the Northeastern Atlantic andMediterranean Sea, F. orensanzi n. sp. from the Southwestern Atlantic, and F. salazarae n. sp. from the Eastern tropicalPacific. Annenkova is monotypic: A. mastigophora (Annenkova, 1952) n. comb. from the Northwestern Pacific. Flabe-graviera contains F. mundata (Gravier, 1906) n. comb. and F. profunda n. sp. both from the Antarctic. Flabehlersia in-cludes two species: F. induta (Ehlers, 1897) n. comb. from Tierra del Fuego and F. persimilis (Annenkova-Chlopina 1924)n. comb. from the Northwestern Pacific. Flabesymbios contains two species living on Northeastern Pacific sea-urchins: F. commensalis (Moore, 1909) n. comb. on Strongylocentrotus and F. roberti n. sp. on Centrostephanus.

References

  1. Agassiz, L. (1846(1848)) Nomenclatoris Zoologici: Index Universalis. Jent & Gasmann, Soloduri, 1135 pp.

    Amouroux, J.M. (1974) Sur la présence de Flabelligera affinis Sars (annélide polychète Flabelligeridae). Vie et Milieu, séries A, 24, 179–180.

    Annenkova, N. (1952) Noviye vidy mnogoschetinkovich chervei (Polychaeta). Trudy Zoologischeskogo Instituta Akademii Nauk SSSR, 12, 148–154.

    Annenkova-Chlopina, N. (1924) Neues über die Verbreitung einger Arten der Polychaeten. Comptes Rendus de l’Academie des Sciences de Russie, 1924, 125–128.

    Augener, H. (1918) Polychaeta. Beitrage zur Kenntnis des Meeresfauna West-Afrikas. Herausgegeben von W. Michaelsen. Z.L. Friedrerichsen & Co., Hamburg 2, 67–625, 6 pls.

    Baird, W. (1873) Description of some new species of Annelida and Gephyrea in the collection of the British Museum. Zoological Journal of the Linnean Society, London, 11, 94–97.

    Benham, W.B. (1915a) Notes on some New Zealand polychaetes. Transactions and Proceedings of the New Zealand Institute, 47, 161–170.

    Benham, W.B. (1915b) Preliminary report on the polychaetous annelids from the Kermadec islands. Transactions and Proceedings of the New Zealand Institute, 47, 170–174.

    Benham, W.B. (1921) Polychaeta. Australasian Antarctic Expedition 1911–14, Scientific Reports, series C, Zoology and Botany, 6(3), 1–128.

    Benham, W.B. (1927) Polychaeta. British Antarctic (“Terra Nova”) Expedition 1910, Zoology, 7(2), 47–182.

    Benham, W.B. (1950) Scientific results of the New Zealand Sub-Antarctic Expedition, 1941–45: Polychaeta. Bulletin of the Cape Expedition series,10, 3–26.

    Berkeley, E. & C. Berkeley. (1952) Canadian Pacific Fauna, 9. Annelida, 9b(2). Polychaeta Sedentaria. Univ. Toronto Press, 139 pp.

    Blake, J.A. (2000) Family Flabelligeridae Saint-Joseph, 1894; pp 1–23 In Taxonomic Atlas of the Benthic Fauna of the Santa Maria Basin and the Western Santa Barbara Channel. The Annelida 4, Polychaeta: Flabelligeridae to Sternaspidae. J.A. Blake, B. Hilbig & P.V. Scott (eds.), Santa Barbara Museum of Natural History, Santa Barbara, 7 vols.

    Chamberlin, R.V. (1919a) New polychaetous annelids from Laguna Beach, California. Journal of Entomology and Zoology, Pomona College, 11, 1–23.

    Chamberlin, R.V. (1919b) The Annelida Polychaeta of the Albatross Tropical Pacific Expedition, 1891–1905. Memoirs of the Museum of Comparative Zoology, Harvard College, 48, 1–514.

    Chamberlin, R.V. (1920) The polychaetes collected by the Canadian Arctic Expedition, 1913–18. Reports of the Canadian Arctic Expedition, 1913–18, 9. Annelids, Parasitic Worms, Protozoans, etc., B. Polychaeta, Southern Party 1913–16, 1B–41B.

    Claparède, E. (1869) Les Annélides Chétopodes du Golfe de Naples. Seconde Partie. Mémoires de la Société de Physique et d’Histoire Naturelle de Genève 20(1), 1–225.

    Costa, O.G. (1841) Description des quelques annélides nouvelles du golfe de Naples. Annales des Sciences Naturelles (Zoologie), Série 2, 16, 267–280.

    Cunningham, J.T. & G.A. Ramage. (1888) The Polychaeta Sedentaria of the Firth of Forth. Transactions of the Royal Society of Edinburgh, 33, 635–684.

    Cuvier, G. (1830) Le Règne Animal distribué après son organisation, pour servir de bae a l’Histoire naturelle des animaux et d’introduction a l’anatomie comparée. Deterville, Paris, vol. 2.

    Day, J.H. (1961) The Polychaeta Fauna of Sout Africa, 6. Sedentary species dredged off Cape coast with a few new records from the shore. Zoological Journal of the Linnean Society, London, 44, 463–560.

    Day, J.H. (1967) A Monograph on the Polychaeta of Southern Africa. British Museum (Natural History) Publications 656:38 + 878 pp.

    de Blainville, H. (1828) Vers (Entomoz.). Dictionnaire des Sciences Naturelles, 57, 365–625.

    de Quatrefages, A. (1849) Études sur les types inférieurs de l’embranchment des Annelés. Mémoire sur la Famille des Chlorèmiens, Chloraemea, Nob. Annales des Sciences Naturelles, troisième série, Zoologie, 12, 277–306, Pl. 9–10.

    de Quatrefages, A. (1866(1865)) Histoire Naturelle des Annelés Marins et d’Eau Douce: Annélides et Géphyriens. Tome Premier. Collection des Suites a Buffon formant avec les Oeuvres de cet auteur un Cours Complet d’Histoire Naturelle. Librairie Encyclopédique de Roret, Paris, 588 pp (Wright, E.P. 1866. Annelida. Zoological Record, Section 6 (Annelida, etc.), 578–600).

    de Saint-Joseph, A. (1894) Les annélides polychètes des côtes de Dinard, troisième partie (Nephtydiens – Serpuliens). Annales des Sciences Naturelles, Zoologie, 7ème série, 17, 1–395.

    delle Chiaje, S. (1841) Descrizione e notomia degli animali invertebrati della Sicilia cateriore osservati vivi negli anni 1822–1830. Napoli, vol. 3.

    Dujardin, F. (1839) Observations sur quelques annelids marines. Annals de Sciences Naturelles, Paris, série, 2 11, 287–294.

    Durham, J.W., C.D. Wagner & D.P. Abbott. (1980) Echinoidea: the sea urchins; pp 160–176 in Intertidal Invertebrates of California. R.H. Morris, D.P. Abbott & E.C. Haderlie (eds.), Stanford University Press, Stanford.

    Ehlers, E. (1897) Polychaeten. Hamburger Magalhaensische Sammelreise. Friederichsen, Hamburg, 148 pp.

    Ehlers, E. (1900) Magellanische Anneliden gesammelt während der schwedisch Expedition nach den Magellanländern. Koeniglichen Gesellschaft der Wissenschaften zu Göttingen, Mathematisch-physikalische Klasse, 1900(2), 1–18.

    Ehlers, E. (1901) Die Polychaeten des magellanischen und chilenischen Strandes: Ein faunisticher Versuch. Fetschrift zur Feier des 150jährigen Bestehens der Königlichen Gesellschaft der Wissenschaften zu Göttingen. Weidmannsche, Berlin, 232 pp.

    Ehlers, E. (1905(1904)) Nueseeländische Anneliden. Abhandlungen der Königlichen gesellschaft der wissenschaften zu Göttingen, Mathematisch-Physikalische Klasse, neue Folge, 3, 1–79.

    Ehlers, E. (1907) Neuseeländische Anneliden, 2. Abhandlungen der Königlichen gesellschaft der wissenschaften zu Göttingen, Mathematisch-Physikalische Klasse, neue Folge, 5, 1–31.

    Ehlers, E. (1913) Die Polychaeten – Sammlungen der Deutschen Südpolar-Expedition 1901–1903. Deutsche Südpolar-Expedition 1901–1903 im auftrage des reichsamtes des innern. 13, Zoologie, 5, 397–598.

    Fabricius, O. (1780) Fauna Groenlandica, systematice systems, Animalia Groenlandiae occidentalis hactenus indagata, quoad nomen specificum, triviale, vernaculumque; synonyma auctorum plurium, descriptionem, locum, victum, generationem, mores, usum, capturamque singuli; prout detegendi occasio fuit, maximaque parti secundum prorias observations. Hafniae et Lipsiae, 16 + 452 pp.

    Fauchald, K. (1977) The polychaete worms: Definitions and keys to the orders, families and genera. Natural History Museum of Los Angeles County, Science Series, 28, 1–190.

    Fauvel, P. (1927) Polychètes sédentaires & addenda aux polychètes errantes. Faune de France, 16, 1–494.

    Fauvel, P. & F. Rullier. (1957) Nouvelle contribution à la faune des annélides polychètes du Sénégal (deuxième partie). Bulletin de l’Institut Français d’Afrique Noire, série A 19, 375–399.

    Fauvel, P. & F. Rullier. (1959a) Golfe de Guinée, 6. Annélides polychètes. Résultats Scientifiques des Campagnes de la Calypso, 4, Annales de l’Institut Océanographique, 37, 143–205.

    Fauvel, P. & F. Rullier. (1959b) Contribution à la faune des annélides polychètes du Sénégal et de Mauritanie (2e partie). Bulletin de l’Institut Français d’Afrique Noire, série A 21, 934–987.

    Gravier, C. (1906) Sur les annélides polychètes recueillies par l’Expédition Antarctique Française (Aphroditiens, Amphinomiens, Flabelligériens, Maldaniens, Ampharétiens). Bulletin du Museum d’Histoire Naturelle, Paris 7, 535–540.

    Gravier, C. (1907) Annélides polychètes. Expédition Antarctique Française (1903–1905) commandée par le Dr. Jean Charcot. Sciences Naturelles, Vers, 12, 1–75.

    Gravier, C. (1911) Annélides polychètes. Deuxième Expédition Antarctique Française (1908–1910) commandée par le Dr Jean Charcot. Sciences Naturelles. Masson et Cie., Paris 1, 1–165.

    Gravier, C. (1918a) Sur un nouveau copépode (Flabellicola n.g. neapolitana n. sp.) parasite d’un annélide polychète [Flabelligera diplochaitos (Otto)]. Comptes Rendus de l'Académie des Science, Paris 166, 502–506.

    Gravier, C. (1918b) Contribution à l’étude d’un copépode (Flabellicola neapolitana Gravier) parasite d’un annélide polychète [Flabelligera (Siphonostoma) diplochaitos (Otto)]. Publicazioni della Stazione Zoologica di Napoli, 2, 209–222.

    Gravier, C. (1923a) Sur l’habitat du Flabellicola neapolitana Gravier. Bulletin du Muséum National d'Histoire Naturelle 39, 503–505.

    Gravier, C. (1923b) Sur un crustacé parasite (Flabellicola neapolitana Gravier) et sur sa fréquence chez un annélide polychète [Flabelligera (Siphonostoma) diplochaitos (Otto)]. Bulletin du Muséum National d'Histoire Naturelle, 29, 560–562.

    Gravier, C. (1924) Sur l’évolution d’un crustacé parasite (Flabellicola neapolitana Gravier). Comptes Rendus de l'Académie des Science, Paris 178, 163–168.

    Gravier, C. & Dantan, J.-L. (1928) Pêches nocturnes a la lumière dans la Baie d’Alger. Annales de l’Institut Océanographique, Paris, 5(1), 1–185.

    Grube, A.E. (1840) Actinien, Echinodermen und Würmer des Adriatischen- und Mittelmeers nach eigenen Sammlungen beschrieben. J.H. Bon, Königsberg, 92 pp.

    Grube, A.E. (1850) Die Familien der Anneliden. Archiv für Naturgeschichte, Berlin, 16, 249–364.

    Grube, A.E. (1877) Mittheilungen über die Familie der Chlorhaeminen. Jahres-Bericht der Schlesischen Gesellschaft für Vaterländische Cultur, 45, 60–73.

    Günther, K. (1912) Beiträge zur Systematik der Gattung Flabelligera und Studien über den Bau von Flabelligera (Siphonostoma) diplochaitus, Otto. Jenaische Zeitschrift für Naturwissenschaft herausgegeben von der Medizinisch-naturwissenschaftlichen Gesellschaft zu Jena 48, Neue Folge, 41, 93–186.

    Haase, P. (1915) Boreale und arktisch Chloraemiden. Wissenschaftliche Meeresuntersuchungen der Kommission zur Wissenschaftlichen Untersuchung der Deutschen Meere, Neue Folge, Kiel, 17, 169–228.

    Harrington, N.R. & B.B. Griffin. (1897) Notes upon the distribution and habits of some Puget Sound invertebrates. Transactions of the New York Academy of Sciences, 16, 152–165.

    Hartman, O. (1938) Annotated list of the types of polychaetous annelids in the Museum of Comparative Zoology. Bulletin of the Museum of Comparative Zoology, Harvard College, 85, 1–31.

    Hartman, O. (1952) The marine annelids of the United States Navy Antactic Expedition, 1947–48. Journal of the Washington Academy of Sciences, 42, 231–237.

    Hartman, O. (1953) Non-pelagic Polychaeta of the Swedish Antarctic Expedition 1901–1903. Further Zoological Result on the Swedish Antarctic Expedition 1901–1903, under the direction of Dr. Otto Nordenskjöld. N. H. Odhner (ed.) 4(11), 1–83.

    Hartman, O. (1959) Catalogue of the Polychaetous Annelids of the World, 2. Allan Hancock Foundation Occasional Papers, 23(2), 355–628.

    Hartman, O. (1961) Polychaetous annelids from California. Allan Hancock Pacific Expeditions, 25, 1–226.

    Hartman, O. (1966) Polychaeta Myzostomidae and Sedentaria of Antarctica. Antarctic Research Series, 7, 1–158.

    Hartman, O. (1967) Polychaetous annelids collected by the USNS Eltanin and Staten Islands Cruises, chiefly from Antarctic seas. Allan Hancock Monographs in Marine Biology, 2, 1–387.

    Hartman, O. (1969) Atlas of the Sedentariate Polychaetous Annelids from California. Allan Hancock Foundation, University of Southern California, Los Angeles, 812 pp.

    Hartman, O. (1978) Polychaeta from the Weddell Sea Quadrant, Antarctica. Antarctic Research Series, 26, 125–222.

    Hartmann-Schröder, G. (1965) Die Polychaeten des Sublitorals. Zur Kentnnis des Sublitorals der chilenischen Küste unter besonderer Berücksichtigung der Polychaeten und Ostracoden. (Mit Bemerkungen über den Einfluss sauerstoffarmer strömungen auf die Besiedlung von marinen Sedimenten). Mitteilungen aus dem Hamburgischen Zoologischen Museum und Institut, 62(Ergänzungsband), 59–305.

    Hartmann-Schröder, G. (1971) Annelida, Borstenwürmer, Polychaeta. Die Tierwelt Deutschlands, 58, 1–594.

    Hartmann-Schröder, G. (1996) Annelida, Borstenwürmer, Polychaeta. Die Tierwelt Deutschlands 58, 2nd ed., Fischer, Jena, 645 pp.

    Hartmann-Schröder, G. & P. Rosenfeldt. (1989) Die Polychaeten der “Polarstern”-Reise ANT III/2 in die Antarktis 1984, 2. Cirratulidae bis Serpulidae. Mitteilungen aus dem Hamburgischen Zoologischen Museum und Institut, 86, 65–106.

    Hartwich, G. (1993) Die Polychaeten-Typen des Zoologischen Museums in Berlin. Mitteilungen aus dem Zoologischen Museum in Berlin, 69, 73–154.

    Haswell, W.A. (1886) Observations on some Australian Polychaeta. Proceedings of the Linnean Society of New South Wales, 10, 733–756.

    Haswell, W.A. (1892) Observations on the Chloraemidae, with special reference to certain Australian forms. Proceedings of the Linnean Society of New South Wales, series 2, 6, 329–356.

    Hobson, K.D. & K. Banse. (1981) Sedentariate and Archiannelid Polychaetes of British Columbia and Washington. Canadian Bulletin of Fisheries and Aquatic Sciences, 209, 8 + 144.

    ICZN (International Commission on Zoological Nomenclature). (1999) International Code of Zoological Nomenclature, 4th ed. International Trust for Zoological Nomenclature (The Natural History Museum), London, 306 pp (www.iczn.org/iczn/index.jsp).

    Jirkov, I.A. & A.V. Filippova. (2001) Flabelligeridae Saint-Joseph, 1894; pp 348–363 In Polikhety Severnogo Ledovitogo Okeana. I.A. Jirkov. Yanus-K, Moskva, 632 pp.

    Johnson, H.P. (1901) The Polychaeta of the Puget Sound region. Proceedings of the Boston Society of Natural History, 29, 381–437.

    Jourdan, E. (1887) Étude anatomique sur le Siphonostoma diplochaetos, Otto. Annales du Musée d’Histoire Naturelle de Marseille, Zoologie, 3(2), 1–43.

    Kinberg, J.G.H. (1910) Konglia Svenska Fregatten Eugenies Resa imkring jorden under befall af C.A. Virgin åren 1851–1853. Vetenskapliga Iakttagelser På H. Maj: T. Konnung Oscar den Förstes befallning utgifna. Kongelige Svenska Vetenskaps-Akademien. Zoologie, 3. Annulata, 2, 33–78.

    Kirkegaard, J.B. (1959) The Polychaeta of West Africa, 1. Sedentary species. Atlantide Reports, 5, 7–117.

    Knox, G.A. & D.B. Cameron. (1998) The marine fauna of the Ross Sea: Polychaeta. National Institute of Water and Atmospheric Research, Biodiversity Memoir, 108, 1–125.

    Kretzler, J.E. (1984) Echinophilus xiphidion, new species (Ostracoda, Paradoxostomatidae) parasitic on regular echinoids of the Northeastern Pacific. Journal of Crustacean Biology, 4, 333–340.

    Lee, Y.H. (2003) Molecular phylogenies and divergence times of sea urchin species of Strongylocentrotidae, Echinoidea. Molecular Biology and Evolution, 20, 1211–1221.

    Levenstein, R.Y. (1966) Mnogotshchetinkov’e chervi (Polychaeta) zapadnoi tasti Beringova morya. Akademiya Nauk SSSR, Trudy Instituta Okeanologii, 81, 3–131.

    Lie, U. & D.S. Kisker. (1970) Species composition and structure of benthic infauna communities off the coast of Washington. Journal of the Fisheries Research Board, Canada, 27, 2273–2285.

    Light, W.J. (1978) Reexamination of the species referred to the genus Flabelliderma Hartman (Polychaeta: Flabelligeridae and Acrocirridae). Proceedings of the Biological Society of Washington, 91, 681–90.

    Loi, T.N. (1980) Catalogue of the types of polychaete species erected by J. Percy Moore. Proceedings of the Academy of Natural Sciences of Philadelphia, 132, 121–149.

    McIntosh, W.C. (1869) On the structure of the British nemerteans, and some new British annelids. Transactions of the Royal Society of Edinburgh, 25, 305–433.

    McIntosh, W.C. (1904(1903)) Marine annelids (Polychaeta) of South Africa, 1. Marine Investigations in South Africa, 3(19), 15–56.

    McIntosh, W.C. (1908) Notes from the Gatty Marine Laboratory, St. Andrews, 30: 1. On the stranding of an adult female Mesoplodon bidens, Sowerby, at St. Andrews; 2. On an abnormal plaice with a precaudal fin-frill on the left side; 3. On Orthogoriscus mola, Bl.; 4. On the British Sphaerodoridae, Chloraemidae, and Chaetopteridae; 5. On the same families dredged in the ‘Porcupine’ expeditions of 1869 and 1870; 6. On the foregoing families dredged in the Gulf of St. Lawrence, Canada, by Dr. Whiteaves; 7. On the same families dredged in Norwegian waters and in Finmark by Canon Norman. Annals and Magazine of Natural History, series 8, 2, 524–545.

    McIntosh, W.C. (1915) A Monograph of the British Annelids, 3. Polychaeta: Opheliidae to Ammocharidae. Ray Society, London, 368 pp.

    Monro, C.C.A.(1930) Polychaete worms. Discovery Reports, 2, 1–222.

    Monro, C.C.A. (1933) The Polychaeta Sedentaria collected by Dr. C. Crossland at Colón, in the Panama region, and the Galapagos Islands during the Expedition of the S.Y. "St. George". Proceedings of the Zoological Society, London, 1933(4), 1039–1092.

    Monro, C.C.A. (1936) Polychaete worms, 2. Discovery Reports, 12, 59–198.

    Monro, C.C.A. (1937) Polychaeta. Scientific Reports of the John Murray Expedition, 1933–34 4(8), 243–321.

    Monro, C.C.A. (1939) Polychaeta. Reports of the British Antarctic Research Expedition 1929–1931, series B 4(4), 87–156.

    Moore, J.P. (1909a) Polychaetous annelids from Monterey Bay and San Diego, California. Proceedings of the Academy of Natural Sciences of Philadelphia, 61, 235–295.

    Moore, J.P. (1909b) The polychaetous annelids dredged in 1908 by Mr. Owen Bryant off the coasts of Labrador and Newfoundland and Nova Scotia. Proceedings of the United States National Museum, 37, 133–146.

    Moore, J.P. (1923) The polychaetous annelids dredged by the USS "Albatross" off the coast of southern California in 1904, 4. Spionidae to Sabellariidae. Proceedings of the Academy of Natural Sciences of Philadelphia, 75, 179–259.

    Mueller, M. (1852) Observationes anatomicae de Vermibus quibusdam maritimis. Dissertatio inauguralis comparator-anatomica quam consensus et auctoritate Gratiosi Medicorum ordinis in Alma Litteratum Universitate Friderica Guilelma ut Summi in Medicina et Chirugia honores. Berolini, 32 pp.

    Müller, O.F. (1776) Zoologiae Danicae Prodromus, seu Animalium Daniae et Norvegiae indigenarum. Characteres, nomina et synonyma imprimis popularium. Hallageriis, Havniae, 274 pp.

    Müller, O.F. (1789) Zoologica Danica seu Animalium Daniae et Norwegiae rariorum ac minus notorum, Descriptiones et Historia 3, 1–71.

    Neave, S.A. (1939) Nomenclator Zoologicus: A List of the Names of Genera and Subgenera in Zoology from the tenth edition of Linneaeus 1758 to the end of 1935. Zoological Society of London, London, 4 vols. (uio.mbl.edu/NomenclatorZoologicus/).

    Newbigin, M.I. (1900) On British species of Siphonostoma. Annals and Magazine of Natural History, series 7, 5, 190–195.

    Oken, L. (1807) (Untitled). Göttingische gelehrte Anzeigen 1807(2), Stück 117 (23 Julius 1807), 1161–1168.

    Oken, L. (1815) Lehrbuch der Naturgeschichte, 3. Theil Zoologie, 1. Fleischlose Thiere und Atlas. August Schmid & Co., Jena.

    Otto, A.G. (1820) De Sternaspide thalassemoideo et Siphostomate diplochaito vermibus duobus marinis. Epistola Gratulatoria quam ad celebrandum diem laetissimum VI Marti MDCCCXX, Vratislaviae, 16 pp, 2 plates.

    Otto, A.G. (1821) Animalium maritorum nondum editorum, Genera duo. Verhandlungen der Kaiserlichen Leopoldinisch-Carolinischen Akademie der Naturforscher, 10(2), 617–634.

    Oug, E., T. Bakken & J.A. Kongsrud. (2011) Guide to identification of Flabelligeridae (Polychaeta) in Norwegian and adjacent waters. Norwegian Polychaete Forum Guides, 16 pp. www.polychaeta.no.

    Pettibone, M.H. (1954) Marine polychaete worms from Point Barrow, Alaska, with additional records from the North Atlantic and North Pacific. Proceedings of the United States National Museum, 108, 203–356.

    Pettibone, M.H. (1956) Marine polychaete worms from Labrador. Proceedings of the United States National Museum 105, 531–584.

    Pettibone, M.H. (1982) Annelida; pp 1–43 In Parker, S.P. (Ed.), Synopsis and Classification of Living Organisms. McGraw Hill, New York.

    Rathke, H. (1843) Beiträge zur Fauna Norwegens. Verhandlungen der Kaiserlichen Leopoldinisch-Carolinischen Akademie der Naturforscher, 20, 1–264.

    Rioja-LoBianco, E. (1931) Estudio de los poliquetos de la península Ibérica. Memorias de la Academia de Ciencias Físicas, Exactas y Naturales, Madrid, 2, 1–471.

    Rozbaczylo, N. (1985) Los anélidos poliquetos de Chile: Indice sinonímico y distribución geográfica de especies. Facultad de Ciencias Biológicas, Pontificia Universidad Católica de Chile, Monografías Biológicas, 3, 1-284.

    Rullier, F. & L. Amoureux. (1979) Campagne de la Calypso au large des côtes atlantiques de l’Amérique du Sud (1961–1962), 33. Annélides polychètes. Annales de l’Institut Océanographique, 55(Supplement), 145–206.

    Salazar-Vallejo, S.I. (2007) Revision of Flabelliderma Hartman, 1969 (Polychaeta: Flabelligeridae). Journal of Natural History, 41, 2037–2061.

    Salazar-Vallejo, S.I., L.F. Carrera-Parra & K. Fauchald. (2008) Phylogenetic affinities of the Flabelligeridae (Annelida: Polychaeta). Journal of Systematic Zoology and Evolutionary Research, 46, 203-215.

    Sars, G.O. (1872) Diagnoser af nye Annelider fra Christiana-fjorden, efter M. Sars efterladte Manuskripten. Forhandlinger i Videnskabsselskabet i Kristiania, 1871, 406–417.

    Sars, G.O. (1873) Bidrag til Jundskab om Christiana-fjordens Fauna, 3. Vaesentlig efter Prof. M. Sars’s efterladte Manuskripter. Nyt Magazin for Naturvidenskaberne, 19, 201–281.

    Sars, M. (1829) Bidrag til Söedyrenes Naturhistorie; Förste Hæfte. Dahl, Bergen, 60 pp.

    Sars, M. (1869) Fortsatte Bemærninger over det dryske Livs Udbredning i Havets Dybder. Forhandlinger i Videnskabs-Selskabet i Christiana, 1868, 246–275.

    Schmarda, L.K. (1861) Neue wirbellose Thiere beobachtet und gesammelt auf einer Reise um die Erde 1853 bis 1857. Erster Band: Turbellarien, Rotatorien und Anneliden, Zweite Halfte. Wilhelm Engelman, Leipzig, 165 pp.

    Sherborn, C.D. (1930) Index Animalium sive Index Nominum quae ab A.D. MDCCLVIII Generibus et Speciebus Animalium Imposita sunt. Sectio Secunda: A kalendis januariis, MDCCCI usque ad finem decembris, MDCCCL. 24. Index serratus -squamosus. pp 5911–6118. 1801–1850. Trustees of the British Museum, London.

    Spies, R.B. (1973) The blood system of the flabelligerid polychaete Flabelliderma commensalis (Moore). Journal of Morphology, 139, 465–490.

    Spies, R.B. (1975) Structure and function of the head in flabelligerid polychaetes. Journal of Morphology, 147, 187–208.

    Spies, R.B. (1977) Reproduction and larval development of Flabelliderma commensalis (Moore); pp 323–346 In Essays on the Polychaetous Annelids in Memory of Dr Olga Hartman. D.J. Reish & K. Fauchald (eds.), Allan Hancock Foundation, University of Southern California, Los Angeles, 604 pp.

    Stasek, C.R. (1966) Type specimens in the California Academy of Sciences, Department of Invertebrate Zoology. Occasional Papers of the California Academy of Sciences, 51, 1–38.

    Stebbins, T.D. (1988) Observations and experiments on the natural history and behavior of the commensal isopod Colidotea rostrata (Benedict, 1898) (Isopoda: Idoteidae). Journal of Crustacean Biology, 8, 539–547.

    Stebbins, T.D. (1989) Population dynamics and reproductive biology of the commensal isopod Colidotea rostrata (Crustacea: Isopoda: Idoteidae). Marine Biology, 101, 329–337.

    Stimpson, W. (1854) Synopsis of the marine Invertebrata of Grand Manan, or the region about the mouth of the Bay of Fundy, New Brunswick. Smithsonian Contributions to Knowledge, 6, 1–67.

    Stimpson, W. (1856(1855)) Descriptions of some new marine Invertebrata. Proceedings of the Academy of Natural Sciences of Philadelphia, 7, 385–395.

    Støp-Bowitz, C. (1948a) Les Flabelligériens Norvégiens. Bergens Museums Arbok, 1946(2), 1–59.

    Støp-Bowitz, C. (1948b) Sur les polychaetes arctiques, des famillies des glicériens, des ophéliens, des scalibregmiens et des flabelligériens. Tromsø Museums Årshefter, 66(2), 1–58.

    Théel, H.J. (1879) Les annélides polychetes des mers de la Nouvelle-Zemble. Kongelige Svenska Vetenskaps- Akademiens Handlingar, 16(3), 3–75.

    Uschakov, P.V. (1950) Mnogotschetinkvovye chervei (Polychaeta) Okhotskogo Morya. Issledovaniya Dal’nevostochnykh Morei SSSR, 2, 140–233.

    Uschakov, P.V. (1955) Polychaeta of the Far Eastern Seas of the USSR. Israel Program for Scientific Translations, Jerusalem, (transl. 1965), 26 + 419.

    von Marenzeller, E. (1888) Polychaten der Angra Pequena-Bucht. Zoologische Jahrbücher, Abteilung für Systematik, Geographie und Biologie der Tiere, 3, 1–24.

    Wesenberg-Lund, E. (1950) The Polychaeta of West Greenland with special reference to the Fauna of Nordre Strømfjord, Kvane-, and Bredefjord. Meddelelser om Grønland udgivne af Kommissionen for Videnskabelige undersøgelser I Grønland, 151(2), 1–171.

    Wesenberg-Lund, E. (1953) The Zoology of East Greenland: Polychaeta. Meddelelser om Grønland udgivne af Kommissionen for Videnskabelige undersøgelser I Grønland, 122(3), 1–169.