Abstract
The land snails of the family Amphibulimidae occurring within northwestern South America are critically examined and notes on their distribution are given. The total number of taxa discussed is 61 (7 Dryptus, 54 Plekocheilus); of these, four Colombian Dryptus and 32 Plekocheilus species are included. The following new species are described: Plekocheilus (P.) bigener, P. (P.) incognitus, Plekocheilus (Eurytus) camaritagua, P. (E.) labiosus, and P. (E.) paraguas. The following nominal taxa are now synonymized: Plekocheilus (P.) speciosus (Pfeiffer, 1854) = P. (P.) plectostylus (Pfeiffer, 1848); Plekocheilus (Eurytus) couturesi Ancey, 1900 = P. (P.) glandiformis (Lea, 1838); Plekocheilus (Aeropictus) latilabris (Pfeiffer, 1855) = P. (A.) succineoides (Petit de la Saussaye, 1840); Plekocheilus (Eurytus) virgatus (Pilsbry, 1935) = P. (E.) mabillei (Crosse, 1867) = P. (E.) pulicarius (Reeve, 1848). Also, Simpulopsis fulguratus Miller, 1878 is now considered a junior subjective synonym of Simpulopsis (Eudioptus) citrinovitrea (Moricand, 1836). For the following species, precise localities are given for the first time: Dryptus stuebeli (Martens, 1885), Plekocheilus (Aeropictus) cathcartiae (Reeve, 1848), P. (E.) cardinalis (Pfeiffer, 1853), P. (Eurytus) episcopalis auriformis (Da Costa, 1904), P. (E.) episcopalis corticosus (Sowerby, 1895), and P. (E.) lynciculus (Deville & Huppé, 1850). Newly recorded for the Colombian malacofauna are the following taxa: Plekocheilus (P.) couturesi (Ancey, 1900), P. (Eurytus) piperitus (Sowerby I, 1837). The following taxa are excluded from the Colombian fauna: Dryptus funckii (Nyst, 1843), Plekocheilus (Aeropictus) veranyi (Pfeiffer, 1848), Plekocheilus (Eurytus) jimenezi (Hidalgo, 1872).
References
Abbott, R.T. (1989). Compendium of Landshells. American Malacologists Inc., Melbourne, Florida, 240 pp.
Albers, J.C. (1852) Diagnoses heliceorum novorum. Zeitschrift für Malakozoologie, 9, 30–32.
Albers, J.C. (1854a) Diagnosen neuer Bulimus-Arten. Malakozoologische Blätter, 1, 30–32.
Albers, J.C. (1854b) Novorum Heliceorum diagnoses. Malakozoologische Blätter, 1, 213–221.
Albers, J.C. (1860) Die Heliceen nach natürlicher Verwandtschaft systematisch geordnet, 2 Ausgabe (ed. E. von Martens). Engelmann, Leipzig, xviii + 359 pp.
Ancey, C.F. (1900) Description of a land shell from South America. The Nautilus, 14, 42.
Baker, H.B. (1926) The Mollusca collected by the University of Michigan-Williamson expedition in Venezuela, part IV. Occasional Papers of the Museum of Zoology, University of Michigan (Ann Arbor), 167, 1–49, pls. 7–9.
Baker, H.B. (1963) Type land snails in the Academy of Natural Sciences of Philadelphia Part II. Land Pulmonata, exclusive of North America north of Mexico. Proceedings of the Academy of Natural Sciences of Philadelphia, 115, 191–259.
Bouchet, P., Fryda, J., Hausdorf, B., Ponder, W., Valdés, A. & Warén, A. (2005) Working classification of the Gastropoda. Malacologia, 47, 239–283.
Breure, A.S.H. (1974a) Caribbean land molluscs: Bulimulidae: I. Bulimulus. Studies on the Fauna of Curaçao and other Caribbean Islands, 45, 1–80.
Breure, A.S.H. (1974b) Notes on the genus Gaeotis Shuttleworth, 1854 (Mollusca, Gastropoda, Bulimulidae). Netherlands Journal of Zoology, 24, 236–252.
Breure, A.S.H. (1975) Types of Bulimulidae (Mollusca, Gastropoda) in the Muséum national d’Histoire naturelle, Paris. Bulletin du Muséum national d’Histoire naturelle Paris (3) 331, Zoologie, 233, 1137–1187.
Breure, A.S.H. (1976) Types of Bulimulidae (Gastropoda, Euthyneura) in the Zoologisches Museum, Universität Zürich. In: Malacologische Opstellen. Backhuys, Rotterdam, 1–4, pls. 1–3.
Breure, A.S.H. (1977) Notes on Bulimulidae (Gastropoda, Euthyneura), 5. On some collections from Colombia. Archiv für Molluskenkunde, 107, 257–270.
Breure, A.S.H. (1978) Notes on and descriptions of Bulimulidae (Mollusca, Gastropoda). Zoologische Verhandelingen, 164, 1–255.
Breure, A.S.H. (1979) Systematics, phylogeny and zoogeography of Bulimulinae (Mollusca). Zoologische Verhandelingen, 168, 1–215.
Breure, A.S.H. (1984) Survey of the land Mollusca of the Sierra Nevada de Santa Marta, Colombia. In: Hammen, T. van der & Ruiz, P.M. (eds.), Studies on Tropical Andean ecosystems, 2, La Sierra Nevada de Santa Marta (Colombia) Transecto Buritaca─La Cumbre. Cramer, Berlin/Stuttgart, pp. 487–500.
Breure, A.S.H. (2008) Mysterious or confusing: enigmatic species in the Orthalicidae (Gastropoda, Pulmonata). Basteria, 72, 241–252.
Breure, A.S.H. (2009) New Orthalicidae (Mollusca, Gastropoda) from Venezuelan Guayana: unravelling secrets from the Lost World. Zootaxa, 2065, 25–50.
Breure, A.S.H. (2011) Annotated type catalogue of the Orthalicoidea (Mollusca, Gastropoda) in the Royal Belgian Institute of Sciences, Brussels, with descriptions of two new species. ZooKeys, 101, 1–50.
Breure, A.S.H. & Borrero, F.J. (2008) An annotated checklist of the land snail family Orthalicidae (Gastropoda: Pulmonata: Orthalicoidea) in Ecuador, with notes on the distribution of the mainland species. Zootaxa, 1768, 1–40.
Breure, A.S.H., Groenenberg, D.S.J. & Schilthuizen, M. (2010) New insights in the phylogenetic relations within the Orthalicoidea (Gastropoda: Stylommatophora) based on 28S sequence data. Basteria, 74, 25–31.
Breure, A.S.H. & Schlögl, J. (2010) Additional notes on Orthalicidae from the Chimantá massif, Venezuelan Guayana, with descriptions of new species of Plekocheilus Guilding, 1828. Zootaxa, 2416, 51–60.
Breure, A.S.H. & Schouten, J.R. (1985) Notes on and descriptions of Bulimulidae (Mollusca, Gastropoda), III. Zoologische Verhandelingen, 216, 1–98.
Bruguière, J.C. (1789–1792) Histoire naturelle des Vers, Tome premier. In: Encyclopédie méthodique, ou par ordre de matière, 11. Panckoucke, Paris, 1e Partie, pp. i–xviii + 1–344 (1789), 2e Partie, pp. 345–757 (1992).
Clench, W.J. & Turner, R.D. (1962) New names introduced by H.A. Pilsbry in the Mollusca and Crustacea. Special Publications, Academy of Natural Sciences, Philadelphia, 4, 1–218.
Crosse, H. (1867) Descriptions d’espèces nouvelles. Journal de Conchyliologie, 15, 195–203, pls. 5–6.
Da Costa, S.I. (1898) Remarks on some species of Bulimulus, Sect. Drymaeus, and descriptions of land-shells from Bolivia, Ecuador, and the U.S. of Colombia. Proceedings of the Malacological Society of London, 3, 80–84, pl. 6.
Da Costa, S.I. (1904) Descriptions of new species of land-shells from Central and South America. Proceedings of the Malacological Society of London, 6, 5–6.
Dance, S.P. (1966) Shell collecting. An illustrated history. University of California Press, Berkeley/Los Angeles, 344 pp, 35 pls.
Dall, W.H. (1912) New species of landshells from the Panama Canal Zone. Smithsonian Miscellaneous Collections, 59(18), 1–3, pls. 1–2.
Dautzenberg, P. (1900) Croisières du yacht Chazalie dans l’Atlantique. Mollusques. Mémoires de la Société Zoologique de France, 13, 145–156, pl. 9–10.
Deville, E. & Huppé, H. (1850) Description de quelques coquilles nouvelles provenant de l’expédition de M. de Castelnau. Revue et Magasin de Zoologie, (2) 2, 638–644, pls. 14–16.
Dohrn, H. (1875). Ueber einige von Herrn G. Wallis in Neugranada gesammelte Landconchylien. Jahrbücher der deutschen malakozoologischen Gesellschaft, 2, 292–309, pl. 10.
Dohrn, H. (1882) Beiträge zur Kenntnis der südamerikanischen Landconchylien. Jahrbücher der deutschen malakozoologischen Gesellschaft, 9, 97–115, pl. 3.
Dunker, W. (1875) Zwei neue Bulimi aus der Sierra Nevada (Columbia). Jahrbücher der deutschen malakozoologischen Gesellschaft, 2, 220–221, pl. 6 figs. 1–4.
Dunker, W. (1882) De molluscis nonnullis terrestribus Americae australis. Jahrbücher der deutschen malakozoologischen Gesellschaft, 9, 377–380, pl. 11.
Férussac, A.E.J.d’A. (1821–1822) Tableaux systématiques des animaux mollusques suivis d'un Prodrome général pour tous les mollusques terrestres ou fluviatiles vivants ou fossiles. Première partie, Tableaux systématiques généraux, pp. i–xlvii. Deuxième partie. Tableaux particuliers des mollusques terrestres et fluviatiles. Classe des Gastéropodes, 114 pp [Folio ed.], 111 pp. [Quarto ed.]. Arthus-Bertrand, Paris.
Fulton, H.C. (1908) Description of a new species of Strophocheilus. Proceedings of the Malacological Society of London, 8, 86–87.
Gmelin, J.F. (1791) Caroli a Linné … systema naturae per regna tria naturae ..., editio decima tertia, aucta, reformata, vol. 1(6). G.E. Beer, Lipsiae, 3021–4120.
Götting, K.-J. (1978) Lista preliminar de los caracoles terrestres de la region septentrional de Colombia. Anales del Instituto de Investigaciones Marinas de Punta de Betín, 10, 101–110.
Haas, F. (1955) On some small collections of inland shells from South America. Fieldiana, Zoology, 34, 361–387.
Harvey, M.B. (2008) New and poorly known Dipsas (Serpentes: Colubridae) from northern South America. Herpetologica, 64, 422–451.
Harvey, M.B., Fuenmayor, G.R., Portilla, J.R.C. & Rueda-Almonacid, J.V. (2008) Systematics of the enigmatic dipsadine snake Tropidodipsas perijanensis Alemán (Serpentes: Colubridae) and review of morphological characters of Dipsadini. Herpetological Monographs, 22, 106–132.
Hidalgo, J.G. (1869) Diagnoses molluscorum novorum. Journal de Conchyliologie, 17, 188–189.
Hidalgo, J.G. (1872) Moluscos del viaje al Pacífico verificado de 1862 a 1865 por una comision de naturalistas enviado por el Gobierno Español, Parte Primera, Univalvos terrestres. Carlos Bailly-Bailliere, Madrid, 152 pp., 8pls.
Jonas, J.H. (1844) Vorläufige Diagnosen neuer Conchylien welche ausführlicher beschrieben und abgebildet nächstens erscheinen werden. Zeitschrift für Malakozoologie, 1, 33–37.
Jonas, J.H. (1846) Molluskologische Beiträge. D. Beschreibungen neuer Arten. Abhandlungen aus dem Gebiete der Naturwissenschaften, 1, 109–130, pl. 7 figs. 1–2, pls. 4–5, 8–9, pl. 10 figs. 11, 14–15, pl. 11.
Jousseaume, F. (1889) Voyage de M. Eugène Simon au Venezuela. Mémoires du Sociéte zoologique de France, 2, 232–259, pl. 9.
Jousseaume, F. (1900) Mollusques terrestres: Clausilia, Rhodea et Bulimus Sud-Américanae. Bulletin de la Société philomathique de Paris (9) 2, 5–44, pl. 1.
Köhler, F. (2007) Annotated type catalogue of the Bulimulinae (Pulmonata, Orthalicoidea, Bulimulidae) in the Museum für Naturkunde Berlin. Mitteilungen aus dem Museum für Naturkunde Berlin, Zoologische Reihe, 83, 125–159.
Lea, I. (1838) Descriptions of new freshwater and land shells. Transactions of the American Philosophical Society, (n.s.) 6, 1–154, pls. 1–24.
Lynch, J.D. (1992) Distribution and variation in a Colombian frog, Eleutherodactylus erythropleura (Amphibia: Leptodactylidae). Studies on Neotropical Fauna and Environment, 27, 211–226.
Martens, E. von (1873) Die Binnenmollusken Venezuela’s. Festschrift zur Feier des 100jährigen Bestehens der Gesellschaft für naturforschende Freunde zu Berlin, 157–225, pls.1–2.
Martens, E. von (1885) Uebersicht der von Herrn Dr. Alfred Stübel im nördlichen Theil von Süd-Amerika gesammelten Binnen-Conchylien. Conchologische Mittheilungen, 2, 155–179, pl. 31 figs. 1–3, pl. 32, pl. 35 figs. 1–5, 11–18.
Medina-Bermúdez, C.I. & Arias-Olmos, D.I. (2007) Diversidad de moluscos terrestres del Santuario de flora y fauna de Iguaque (SFFI) (Boyacá), Colombia. Available at http://www.humboldt.org.co/chmcolombia/servicios/jsp/proyectos/Listados.jsp?desde=0&hasta=10&id=455&conector=AND& (accessed 21 April 2010).
Miller, K. (1878) Die Binnenmollusken von Ecuador. Malakozoologische Blätter, 25, 153–199, pls. 7–8.
Mittermeier, R.A., Myers, N., Thomsen, J.B., Fonseca, G.A.B. da & Olivieri, S. (1998) Biodiversity hotspots and major tropical wilderness areas: Approaches to setting conservation priorities. Conservation Biology, 12, 516–520.
Mousson, A. (1873) Zweite Notiz über einige von Herrn Gustav Wallis aus dem nördlichen Süd-Amerika zurückgebrachte Mollusken. Malakozoologische Blätter, 21, 1–19.
National Geographic (2008) Terrestrial ecoregions of the World. Available at http://www.nationalgeographic.com/wildworld/terrestrial.html (accessed 20 February 2008).
Neubert, E. & Janssen, R. (2004) Die Typen und Typoide des Natur-Museums Senckenberg, 84: Mollusca: Gastropoda: Pulmonata: Orthalicoidea: Bulimulidae (2), Orthalicidae, Placostylidae. Archiv für Molluskenkunde, 133, 193–297.
Nyst, H. (1843) Notice sur deux coquilles colombiennes du genre Bulimus. Mémoires du Societé royale de Sciences, Liège, 1, 261–262, pl. 7.
Nyst, H. (1845) Description de deux Bulimes nouveaux de la Colombie. Bulletin de l’Académie royale des Sciences, des Lettres et des Beaux-Arts de Belgique, 12(1), 227–229, pl.
Parkinson, B., Hemmen, J. & Groh, K. (1987) Tropical landshells of the world. Christa Hemmen, Wiesbaden, 279 pp.
Parodiz, J.J. (1962) New and little-known species of South and Central American land snails (Bulimulidae). Proceedings of the United States National Museum, 113(3462), 429–456, pls. 1–2.
Pedraza-Peñalosa, P. (2008) Three new species of Disterigma (Ericaceae: Vaccinieae) from Western Colombia, with comments on morphological terminology. Brittonia, 60, 1–10.
Petit, R.E. (2009) George Brettingham Sowerby I, II & III: their conchological publications and Molluscan taxa. Zootaxa, 2189, 1–218.
Petit de la Saussaye, S.A.A. (1840) Description de quelques coquilles nouvelles, provenant de l’Amérique méridionale. Revue Zoologique par la Société Cuvierienne, 3, 74–75.
Pfeiffer, L. (1846) Description of thirty new species of Helicea belonging to the collection of H. Cuming Esq. Proceedings of the Zoological Society of London, 1846, 28–34.
Pfeiffer, L. (1847) Diagnosen neuer Heliceen. Zeitschrift für Malakozoologie, 4, 65–71.
Pfeiffer, L. (1848a) Descriptions of nineteen new species of Helicea, from the collection of H. Cuming, Esq. Proceedings of the Zoological Society of London, 1847, 228–232.
Pfeiffer, L. (1848b) Monographia heliceorum viventium..., vol. 2. F.A. Brockhaus, Lipsiae, 594 pp.
Pfeiffer, L. (1853a) Monographia heliceorum viventium..., vol. 3. F.A. Brockhaus, Lipsiae, viii, 711 pp.
Pfeiffer, L. (1853b) Description of fifty-four new species of Helicea, from the collection of Hugh Cuming, Esq. Proceedings of the Zoological Society of London, 1851, 252–263.
Pfeiffer, L. (1853c) In: Küster, H.C. & Pfeiffer, L. (1840–1865), Systematisches Conchylien-Cabinet von Martini und Chemnitz, I, 13(1). Die Gattungen Bulimus, Partula, Achatinella, Achatina und Azeca. Bauer und Raspe, Nürnberg, Lieferung 124, pp. 49–96, pls. 19–24.
Pfeiffer, L. (1854─1860). Novitates conchologicae..., vol. 1. Th. Fischer, Cassel, 138 pp., 36 pls.
Pfeiffer, L. (1855a) Descriptions of nine new species of land-shells, in the collection of H. Cuming, Esq. Proceedings of the Zoological Society of London, 1855, 7–9.
Pfeiffer, L. (1855b) Descriptions of thirty-eight new species of Helicea, from the collection of H. Cuming, Esq. Proceedings of the Zoological Society of London, 1855, 111–119.
Pfeiffer, L. (1855c) Descriptions of fifty-seven new species of Helicea, from Mr. Cuming’s collection. Proceedings of the Zoological Society of London, 1855, 286–298.
Pfeiffer, L. & Clessin, S. (1879–1881) Nomenclator heliceorum viventium qui continentur nomina omnium hujus familae generum specierum hodie cognitarum disposita ex affinitate naturali. Theodori Fischeri, Casselis, 617 pp.
Philippi, R.A. (1842–1844) Abbildungen und Beschreibungen neuer oder wenig gekannter Conchylien, vol. 1. Th. Fischer, Cassel, [14] + 204 pp., 48 pls.
Pilsbry, H.A. (1895–1896) Manual of Conchology, (2) 10. American Bulimi and Bululimuli. Strophocheilus, Plekocheilus, Auris, Bulimulus. Academy of Natural Sciences of Philadelphia, Philadelphia, iv + 213 pp, 51 pls.
Pilsbry, H.A. (1901–1902) Manual of Conchology (2) 14. Oriental bulimoid Helicidae; Odontostominae; Cerionidae. Academy of Natural Sciences, Philadelphia, 1–302, pls. 1–62.
Pilsbry, H.A. (1910) Land Mollusca of the Panama Canal Zone. Proceedings of the Academy of Natural Sciencies, Philadelphia, 62, 502–509, pl. 37.
Pilsbry, H.A. (1935) South American land and freshwater mollusks. 9. Colombian species. Proceedings of the Academy of Natural Sciencies, Philadelphia, 87, 83–88, pl. 6.
Pilsbry, H.A. (1939) South American land and freshwater mollusks, X. Species of Colombia and Ecuador. Notulae Naturae, 19, 1–6.
Ramírez, R., Paredes, C. & Arenas, J. (2003) Moluscos del Perú. Revista de Biología Tropical, 51 (Supplement 3), 225–284.
Ramos-Tafur, G. (2006) Dos nuevas especies de cangrejos de agua dulce (Brachyura: Pseudothelphusidae) de la Serranía de Los Paraguas, Cordillera Occidental de Colombia. Revista de Biología Tropical, 54, 1047–1056.
Reeve, L.A. (1848–1850) Conchologia iconica: or illustrations of the shells of molluscous animals, 5. Monograph of the genus Bulimus. Reeve, Benham, and Reeve, London, 89 pls + legend., i–xi.
Richardson, C.L. (1995) Bulimulidae: catalog of species. Tryonia, 28, 1–458.
Rojas-Alvarado, A.F. (2009a) Novelties in Danaea moritziana complex (Marattiaceae) from Colombia. Métodos en Ecología y Sistemática, 4, 8–19.
Rojas-Alvarado, A.F. (2009b) Novelties in Trichomanes L. from Colombia. Caldasia, 31, 9–11.
Schileyko, A.A. (1999) Treatise on Recent terrestrial pulmonate molluscs, 3. Partulidae, Aillyidae, Bulimulidae, Orthalicidae, Megaspiridae, Urocoptidae. Ruthenica, Supplement 2, 263–436.
Shaw, H.O.N. (1909) On the dates of issue of Sowerby’s “Conchological illustrations,” from the copy preserved in the Radcliffe Library, Oxford. Proceedings of the Malacological Society of London, 8, 333–340.
Simmons, F.A.A. (1981) On the Sierra Nevada de Santa Marta and its watershed. Proceedings of the Royal Geographical Society, 12, 717–718.
Simone, L.R.L. (2006) Land and freshwater molluscs of Brazil. EGB/Fapesp, Sao Paulo, 390 pp.
Smith, E.A. (1877) Descriptions of new species of South American Helicidae in the British Museum. Proceedings of the Zoological Society of London, 1877, 361–365.
Sowerby, G.B. I (1832–1841) The conchological illustrations. G.B. Sowerby, London [see Petit 2009 for bibliographic information].
Sowerby, G.B. III (1895) Descriptions of nine new species of shells. Proceedings of the Malacological Society of London, 1, 214–217, pl. 13.
Tello, J. (1975) Catálogo de la Fauna Venezolana. VIII. Mollusca. Published by the author, Caracas, 599 pp.
Thompson, F.G. (2008). An annotated checklist and bibliography of the land and freshwater snails of Mexico and Central America. Florida Museum of Natural History, Gainesville, pp. 903. Available at http://www.flmnh.ufl.edu/malacology/mexico-central_america_snail_checklist.pdf. (accessed 20 April 2010).
Todd, W.E.C. & Carriker, M.A. jr. (1922) The birds of the Santa Marta region of Colombia: A study in altitudinal distribution. Annals of the Carnegie Museum, 14, 3–583, pls. 1–9.
Vásquez-V., V.H. (ed) (2005) Reservas forestales protectoras de Colombia, Atlas básico. Conservation International, Colombia, Ministerio de Ambiente, Vivienda y Desarrollo Territorial; Embajada Real de los Países Bajos, Instituto Geográfico Agustín Codazzi, Bogotá, 127 pp.
Weyrauch, W.K. (1967) Descripciones y notas sobre gastropodos terrestres de Venezuela, Colombia, Ecuador, Brazil y Peru. Acta Zoológica Lilloana, 21, 457–499.
Wood, H. & Gallichan, J. (2008) The new molluscan names of César-Marie-Felix Ancey including illustrated type material from the National Museum of Wales. Studies in Biodiversity and Systematics of Terrestrial Organisms from the National Museum of Wales, Biotir Reports, 3, i–vi, 1–162.