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Published: 2003-09-19

World Catalog of Bee Flies (Diptera: Bombyliidae): Corrigenda and Addenda

J. Linsley Gressitt Center for Research in Entomology, Bishop Museum, 1525 Bernice Street, Honolulu, Hawai‘i 96817-2704, USA
NERC, Centre for Population Biology, Imperial College London, Silwood Park Campus, Ascot, Berkshire SL5 7PY, UK
Diptera Bombyliidae catalog taxonomy distribution


A list of corrections and additions to the world catalog of bee flies, published in 1999 is presented. New taxonomic information presented in this paper includes the following 42 new species-group synonymies: Geron paramonovi Greathead (junior synonym of G. asiaticus Zaitzev), syn. n., Anastoechus olivaceus Paramonov, Anastoechus olivaceus var. corsikanus Paramonov, Anastoechus olivaceus var. corsicanus Zaitzev (all junior synonyms of A. nitidulus Fabricius), syn. n., Anastoechus leucochroicus Hesse (junior synonym of A. varipecten Bezzi, 1921), syn. n., Bombylius punctatus Fabricius, Bombylius sticticus Boisduval, Bombylius subluna Walker (all junior synonyms of Bombomyia vertebralis (Dufour)), syn. n., Bombylius flavescens Palm, Bombylius minor var. ochraceus Paramonov, Bombylius luteolus Evenhuis, Bombylius thapsinoides Evenhuis (all junior synonyms of B. minor Linnaeus), syn. n., Bombylius rhodius Loew (junior synonym of B. trichurus Pallas), syn. n., Bombylius aurulentus Wiedemann (junior synonym of Systoechus ctenopterus (Mikan)), syn. n., Bombylius lucidus Loew, Systoechus tesquorum Becker (all junior synonyms of Systoechusgradatus (Wiedemann)), syn. n., Argyramoeba maculosa Sack, Anthrax ipiriensis Dils & Weyer, Anthrax tajikistanus Evenhuis & Greathead (all junior synonyms of Anthrax chionostigma Tsacas), syn. n., Spogostylum nigrum Austen (junior synonym of Anthrax lucidus (Becker)), syn. n., Argyromoeba velox Loew, Anthrax hellenicus François (all junior synonyms of Anthrax virgo Egger), syn. n., Exoprosopa dispar var. interrupta Paramonov, Exoprosopa dispar ssp. paramonovi Evenhuis (all junior synonyms of E. dispar Loew), syn. n., Thyridanthrax albicingulus Austen, Thyridanthrax amoenus Austen (all junior synonyms of Exhyalanthrax melanchlaenus (Loew)), syn. n., Thyridanthrax innocens Austen (junior synonym of Exhyalanthrax muscarius (Pallas)), syn. n., Anthrax venusta Meigen, Anthrax margaritifer Dufour, Anthrax dolosa Jaennicke, Anthrax turbidus Loew (all junior synonyms of Villa fasciata Meigen), syn. n., Villa orientalis Zaitzev (junior synonym of Villa fasciculata Becker), syn. n., Anthrax humilis Ruthe, Anthrax mucida Zeller, Anthrax pygarga Loew, Anthrax albulus Loew, Villa quinta Becker (all junior synonyms of Villa ixion (Fabricius)), syn. n., Anthrax faustina Osten Sacken, Anthrax alternata var. fulvipes Coquillett, Villa lateralis ssp. semifulvipes Painter in Painter & Painter (all junior synonyms of Villa lateralis (Say)), syn. n., Petrorossia chraminensis Zaitzev, Petrorossia dobrogica Dusa (all junior synonyms of Petrorossia hespera (Rossi)), syn. n.; the following new synonymy in genus-group names: Gazlia Zaitzev (junior synonym of Xeramoeba Hesse), syn. n.; and the following 11 new combinations: Apolysis barri (Tabet) (from Oligodranes), comb. n., Apolysis nauseosus(Tabet) (from Oligodranes), comb. n., Parageron loewi (Becker) (from Usia), comb. n., Chalcochiton carmelitensis (Becker) (from Cytherea), comb. n., Exoprosopa rivularis Meigen (from Heteralonia), comb. n. Micomitra shelkovnikovi (Paramonov) (from Pterobates), comb. n., Xeramoeba albula (Zaitzev) (from Gazlia), comb. n., Xeramoeba kinereti (Zaitzev) (from Chiasmella), comb. n., Xeramoeba stackelbergi (Zaitzev) (from Chiasmella), comb. n., Xeramoeba turkmenica (Zaitzev) (from Chiasmella), comb. n. Two species are returned to their original genus: Exoprosopa dispar Loew (returned to Exoprosopa from Heteralonia), comb. rev., and Spogostylum nitidum Austen (returned to Spogostylum from Anthrax), comb. rev. Three species in Pipunculopsis were invalid when originally proposed by Zaitzev in 2001. They are hereby validated with the criteria fulfilling Article 16.4.2 and the original authorship of Zaitzev should be kept since he provided the information to validate the names herein.


  1. El-Hawagry, M.S. (1999) Two new species of genus Anthrax Scopoli (Bombyliidae-Diptera) from Egypt. Bulletin of the Entomological Society of Egypt, 76[1998], 107–114. (31 December 1999+)

    El-Hawagry, M.S. (2001) Revision of the genus Xeramoeba Hesse (Bombyliidae, Diptera) from Egypt, with description of a new species. Studia Dipterologica, 8, 153–159. (15 August 2001)

    El Hawagry, M.S. (2002) Three new species of anthracine flies (Diptera: Bombyliidae) from Egypt. Zootaxa, 111, 1–8. (26 November 2002)

    El Hawagry, M.S., El Moursy, A.A., Gilbert, F. & Zalat, S. (2000) The tribe Anthracini Latreille (Bombyliidae, Diptera) from Egypt. Egyptian Journal of Biology, 2, 97–117. (31 December 2000+)

    Greathead, D.J. (2000) The family Bombyliidae (Diptera) in Namibia, with descriptions of six new species and an annotated checklist. Cimbebasia, 16, 55–93. (21 March 2000)

    Greathead, D.J. (2001a) A study of the Exoprosopa busiris (Jaennicke) group of Bezzi (1924) in tropical Africa (Diptera: Bombyliidae). Journal of Natural History, 35(1), 127–147. (January 2001)

    Greathead, D.J. (2001b) Notes on the Geron gibbosus Olivier, 1789 and G. halteralis Wiedemann, 1820 species groups (Diptera: Bombyliidae). II - additional species and records from Europe and Asia. Studia Dipterologica, 8(1), 161–173. (15 August 2001)

    Greathead, D.J. (2001c) Exoprosopa pandora (Fabricius, 1805) (Diptera: Bombyliidae) and related species in Europe and the Mediterranean Basin. Insect Systematics and Evolution, 32, 279–284. (October 2001)

    Greathead, D.J. (2003) Notes on Anthrax dentata (Becker, 1906), A. trifasciatus Meigen, 1804, and related species of Bombyliidae (Diptera) in Africa and Eurasia. Deutsche Entomologische Zeitschrift, 50(1), 89–94. (30 June 2003)

    Greathead, D.J. & Evenhuis, N.L. (2001a) Bombylioidea (Diptera: Bombyliidae; Mythicomyiidae) from the island of Sokotra. Zootaxa 14, 11 pp. (26 November 2001)

    Greathead, D.J. & Evenhuis, N.L. (2001b) Annotated keys to the Bombylioidea (Diptera: Bombyliidae; Mythicomyiidae) of Africa. African Invertebrates, 47, 105–244. (15 December 2001)

    Hall, J.C. & Evenhuis, N.L. (2003) Review of the genus Geron (Geron) Meigen in the Nearctic Region (Diptera: Bombyliidae: Toxophorinae). Zootaxa, 181, 72 pp. (16 April 2003)

    Hasbenli, A. & Zaitzev, V.F. (2000) Two new species of Bombylius Linnaeus, 1758 (Diptera, Bombyliidae) from Turkey. Deutsche Entomologisches Zeitschrift, 47, 105–118. (26 June 2000)

    Lamas, C.J.E. & Couri, M.S. (1999) Revisião de Euprepina Hull (Diptera, Bombyliidae, Bombyliinae). Revista Brasileira de Zoología, 16(2), 461–482. (June 1999)

    Lamas, C.J.E., Evenhuis, N.L. & Couri,M.S. (2001) Review of the southern African genus Apatomyza Wiedemann (Diptera, Bombyliidae), with descriptions of new species. Studia Dipterologica, 8(1), 175–186. (15 August 2001)

    Lamas, C.J.E., Evenhuis, N.L. & Couri, M.S. (2003) New species and identifications keys of southern African Crocidiinae (Diptera, Bombyliidae). Deutsche Entomologisches Zeitschrift, 50(1), 95–110. (30 June 2003)

    Nagatomi, A., Tamaki, N. & Evenhuis, N.L. (2000) A new Systropus from Taiwan and Japan (Diptera, Bombyliidae). South Pacific Study, 21(1), 15–18. (31 October 2000)

    Sánchez-Terrón, A. & Roldán Bravo, A.V. (2000) El género Thyridanthrax Osten Sacken, 1886 en el área ibero-balear, con la descripción de una nueva especie (Diptera, Bombyliidae). Boletín de la Asociación Española de Entomología, 24(1-2), 65–84. (20 June 2000)

    Tabet, A.B. [2001a]. A new species of Toxophora from Idaho (Bombyliidae: Diptera). Journal of Basic and Applied Sciences (Libya), 6[1998], 44. (15 June 2001) [The printed date has Ò1998Ó, but correspondence with the author indicated that the journal issue with this article was published on 15 June 2001.]

    Tabet, A.B. [2001b]. New North American Bombyliidae (Diptera). Journal of Basic and Applied Sciences (Libya), 6[1998], 45–47. (15 June 2001) [See Tabet (2001a) annotation for details on dating.]

    Tabet, A.B. [2001c]. Three new species of Geron from Idaho (Diptera: Bombyliidae). Journal of Basic and Applied Sciences (Libya), 6[1998], 48–52. (15 June 2001) [See Tabet (2001a) annotation for details on dating.]

    Tabet, A.B. [2001d]. Description of new species of North American beeflies (Diptera: Bombyliidae. Journal of Basic and Applied Sciences (Libya), 6[1998], 53–57. (15 June 2001) [See Tabet (2001a) annotation for details on dating.]

    Zaitzev, V.F. (1999b) A new genus and new species of flies of the family Bombyliidae (Diptera) from the Palaearctic. Entomologicheskoe Obozrenie, 78(2), 457–463. (22 September 1999) [Date of receipt at ZIS. In Russian; English translation, 2000, in Entomol. Rev. 79(4): 428–433.]

    Zaitzev, V.F. (1999c) On the fauna of flies of the family Bombyliidae (Diptera) of Israel. V. Entomologicheskoe Obozrenie, 78(3), 703–718. (31 December 1999) [Date received by the Russian Entomological Society. In Russian; English translation, 2000a, in Entomol. Rev. 79(6): 640–653.]

    Zaitzev, V.F. (2000b) New species of the genus Petrorossia Bezzi (Diptera, Bombyliidae) from Middle Asia. Entomologicheskoe Obozrenie, 79(2), 482–486. (8 August 2000) [Date of receipt at ZIS. In Russian; English translation, 2001b, in Entomol. Rev. 80(2): 209–212.]

    Zaitzev, V.F. [2001a]. Revision of the genus Pipunculopsis Bezzi, 1925 (Diptera, Bombyliidae). Entomologicheskoe Obozrenie, 79(4)[2000], 911–919. (24 January 2001) [Date of receipt at ZIS. In Russian; English translation 2001c, in Entomol. Rev. 80(5): 578–585.]

    Zaitzev, V.F. [2002a]. Three new species of the family Bombyliidae (Diptera from Israel. Entomologicheskoe Obozrenie, 80(4), 896–900. (1 February 2002) [Date of receipt at ZIS. In Russian; English translation, 2002b, in Entomol. Rev. 81(4): 426–429.]

    Zaitzev, V.F. (2002a) New species of the family Bombyliidae (Diptera) from Palaearctic. Entomologicheskoe Obozrenie, 81(2), 439–444. (11 September 2002) [Date of receipt at ZIS.]

How to Cite

EVENHUIS, N. L., & GREATHEAD, D. J. (2003). World Catalog of Bee Flies (Diptera: Bombyliidae): Corrigenda and Addenda. Zootaxa, 300(1), 1–64.