Abstract
The recent representatives of the subfamily Cyclocypridinae Kaufmann, 1900 are revised here, based on some newly collected Australian material, as well as an extensive study of type material of already described species deposited in various museums. The following two new genera are proposed: Kempfcyclocypris gen. nov. and Keysercypria gen. nov. The genus Kempfcyclocypris is erected to include single new species from subterranean waters of New South Wales, Australia. Kempfcyclocypris australis gen. et sp. nov. can be distinguished from other members of the subfamily by the following characters: 6-segmented antennula, absence of the sexual bristles on antenna, weakly asymmetrical prehensile palps, absence of the basal seta on the second thoracopod, and the long distal seta on the penultimate segment of the third thoracopod. Keysercypria is erected to include some South American species previously described in the genera Physocypria Vávra, 1897 or Cypria Zenker, 1854. The main characters of this genus are: rather globular carapace, with or without marginal tubercles; very short setae on endopodal segments of the third thoracopod; unequally long setae “h1” and “h2” on the terminal segment of the same appendage; and the presence of the basal seta on the second thoracopod. Keysercypria affinis (Klie, 1933) comb. nov. is chosen as the type species and, together with K. deformis (Klie, 1940) comb. nov., K. longiseta (Klie, 1930) comb. nov., K. obtusa (Klie, 1940) comb. nov., and K. pellucida (Sars, 1901), redescribed in the present paper and lectotype and paralectotype are designated. After examining and redescribing the type species of the genus Physocypria Vávra, 1897, P. bullata Vávra, 1897, the genus Mecynocypria Rome, 1962 is synonymised with Physocypria. Lectotype of P. bullata is here designated. For each valid genus a diagnosis, a key to species and a distribution map are provided. A list of six genera and 87 species, currently belonging to the subfamily Cyclocypridinae, is given at the end of the paper, along with their synonyms. Twenty four species are not included in the keys due to lack of data; these are also listed at the end of the paper.References
Baltanás, A. & Geiger, W. (1998) Intraspecific morphological variability: morphometry of valve outlines. In: Martens, K. (Ed.), Sex and Parthenogenesis , Evolutionary Ecology of Reproductive Modes in Non-Marine Ostracods. Backhuys, Leiden: 127–142.
Battish, S.K. (1985) A new species of Cypria (Ostracoda: Crustacea) from India. Research Bulletin (Science) of the Punjab University, 36, 155–156.
Brady, G.S. (1904) On Entomostraca collected in Natal by Mr. James Gibson. Proceedings of the Zoological Society of London, 2, 121–128.
Brady, G.S. (1913) On Freshwater Entomostraca from various parts of South Africa. Annals of the Natal Museum, 2, 459–474.
Brady, G.S. (1886) Notes on Entomostraca collected by Mr. A. Haly in Ceylon. Journal of the Linnean Society of London, Zoology, 19, 293–317.
Brady, G.S. & Norman, A. M. (1889) A monograph of the marine and freshwater Ostracoda of the North Atlantic and of Northwestern Europe. Section I: Podocopa. Scientific Transactions of the Royal Dublin Society, 4, 63–270.
Bronstein, Z.S. (1928) K poznaniyu Ostracoda Kavkaza I Persii. Raboty Severo-Kavkazkoy Hidrobiologicheskoy Stantsii, 2, 67–119.
Broodbakker, N.W. & Danielopol, D.L. (1982) The chaetotaxy of Cypridacea (Crustacea, Ostracoda) limbs: proposal for a descriptive model. Bijdragen tot de Dierkunde, 52, 103–120.
Cole, M.E. (1965) Seven new species of ostracods from Tennessee. Journal of the Tennessee Academy of Science, 40, 132–142.
Daday, E. (1905) Untersuchungen über die Süsswasser-Mikrofauna Paraguays. Mit einem Anhang von W. Michaelsen. Zoologica, 18, 1–374.
Daday, J. (1910) Die Süsswasser-Mikrofauna Deutsch-Ost-Afrikas. Zoologica, 59, 1–314.
Danielopol, D.L. (1969) Recherches sur la morphologie de l’organe copulateur mâle chez quelques ostracodes du genre Candona Baird (fam. Cyprididae Baird). In: Neale, J. W. (Ed.), The taxonomy, morphology and ecology of Recent Ostracoda. Oliver & Boyd Ltd, Edinburgh, 136–153.
Dobbin, C.N. (1941) Fresh-water Ostracoda from Washington and other western localities. University of Washington Publications in Biology, 4, 175–246.
Dobbin-Evenson, C.N. (1969) Designation of lectotypes of fresh-water species described by Dobbin, 1941 (Ostracoda, Crustacea). In: Neale, J.W. (Ed.), The taxonomy, morphology and ecology of recent ostracoda. Oliver and Boyd Edinburgh, 491–494.
Farkas, H.K. (1958) Über einige Süsswasser-Ostracoden von Südamerika. Annales Historico Naturales Musei Nationalis Hungarici, 9, 307–312.
Fischer, S. (1855) Beitrag zur Kenntnis der Ostracoden. Abhandlungen der Mathematisch-Physicalischen Classe der Königlich Bayerischen Akademie der Wissenschaften, 7, 635–666.
Ferguson, E., Jr. (1966) Some freshwater ostracods from the western United States. Transactions of the American Microscopical Society, 85, 313–318.
Frogley, M.R., Griffiths, H.I. & Martens, K. (2002) Modern and Fossil Ostracods from Ancient Lakes. In: Holmes, J.A. & Chivas, A.R. (Eds), The Ostracoda, application in Quaternary Research. Geophysical Monograph, 131, 167–184.
Fuhrmann, R. & Pietrzeniuk, E. (1990) Die Ostrakodenfauna des Interglazials von Gröbern (Kreis Gräfenhainichen). In Eismann, L. (Ed.), Die Eemwarmzeit und die frühe Weichseleiszeit im Saale-Elbe-Gebiet. Geologie, Paläontologie, Palökologie. Altenburger Naturwissenschaftliche Forschungen, 5, 168–193.
Furtos, N.C. (1933) The Ostracoda of Ohio. Ohio Biological Survey, 5, 524 pp.
Furtos, N.C. (1935) Fresh-water Ostracoda from Massachusetts. Journal of the Washington Academy of Science, 25, 529–544.
Furtos, N.C. (1936a) On the Ostracoda from the cenotes of Yucatan and vicinity. Carnegie Institution of Washington, 457, 89–115.
Furtos, N.C. (1936b) Fresh-water Ostracoda from Florida and North Carolina. The American Midland Naturalist, 117, 491–522.
Green, J. (1962) Zooplankton of the River Sokoto. The Crustacea. Proceedings of the Zoological Society of London, 138, 415–453.
Gurney, R. (1916) On some fresh-water Entomostraca from Ceylon. Proceedings of the Zoological Society of London, 23, 333–342.
Henry, M. (1923) A monograph of the freshwater Entomostraca of New South Wales. Part III. Ostracoda. The Proceedings of the Linnean Society of New South Wales, 48, 267–285.
Hartmann, G. (1959) Beitrag zur Kenntnis des Nicaragua-Sees unter besonderer Berücksichtigung seiner Ostracoden (mit Beschreibung von 5 neuen Arten). Zoologischer Anzeiger, 162, 269–294.
Hartmann, G. (1964) Asiatische Ostracoden. Systematische und zoogeographische Untersuchungen. Internationale Revue der gesamten Hydrobiologie, Systematische Beihefte, 3, 155 pp.
Hartmann, G. (1989) Ostracoda. In: Bronn, H.G. (Ed.), Klassen und Ordnungen des Tierreichs. VEB Gustav Fischer Verlag, Jena, 2 (4), 787–1067.
Hartmann, G. & Puri, H.S. (1974) Summary of neontological and paleontological classification of Ostracoda. Mitteilungen aus dem Hamburgischen Zoologischen Museum und Institut, 70, 7–73.
Hoff, C.C. (1942) The ostracods of Illinois. Their biology and taxonomy. Illinois Biological Monographs, 19, 196 pp.
Jančařík, A. (1949) Prispevek k poznani rodu Cyclocypris (Ostracoda) s popisem noveho druhu. Prirodovedecky Sbornik Ostravykeho Kraje, 10, 1–26.
Jurine, L. (1820) Histoire des Monocles, qui se trouvent aux environs de Genève. I-XVI, 1–260.
Karanovic, I. (2000) On Pseudocypridopsis petkovskii sp. nov., a stygobiont freshwater ostracod (Crustacea, Ostracoda, Cypridopsinae) from Montenegro (SE Europe). Mitteilungen aus dem Hamburgischen Zoologischen Museum und Institut, 97, 59–66.
Karanovic, I. (2007) Candoninae (Ostracoda) from the Pilbara region in Western Australia. Crustaceana Monographs, 7, 433 pp.
Kaufmann, A. (1900) Cypriden und Darwinuliden der Schweiz. Revue Suisse de Zoologie, 8, 209–423.
Kempf, E. (1980) Index and Bibliography of Nonmarine Ostracoda. 1. Index A. Geologisches Institut der Universität zu Köln, Sonderveröffentlichungen, 35, 1–188.
Kempf, E. (1997) Index and Bibliography of Nonmarine Ostracoda. 6. Index A, Supplement 1. Geologisches Institut der Universität zu Köln, Sonderveröffentlichungen, 109, 1–142.
Klie, W. (1923) Beitrag zur Kenntnis der Süßwasserostracoden Russlands. Arbeiten der Biologischen Wolga-Station, 7, 1–10.
Klie, W. (1930) Ostracoden aus dem paraguayischen Teile des Gran-Chaco. Archiv für Hydrobiologie, 22, 221–258.
Klie, W. (1933) Süß- und Brackwasser Ostracoden von Bonaire, Curaçao und Aruba. Zoologische Jahrbücher, 64, 289–508.
Klie, W. (1936) Zwei neue Süßwasser-Ostracoden der Unterfamilie Candocyprinae von der Insel Korfu. Zoologischer Anzeiger, 113, 325–331.
Klie, W. (1939) Studien über Ostracoden aus dem Ohridsee: I. Candocyprinae. Archiv für Hydrobiologie, 35, 28–45.
Klie, W. (1940) Süßwasserostracoden aus Nordbrasilien, VII. Ilyocyprinae und Candocyprinae. Zoologischer Anzeiger, 130, 219–229.
Klie, W. (1941) Süßwasserostracoden aus Südosteuropa. Zoologischer Anzeiger, 133, 233–244.
Klie, W. (1944) Ostracoda. Exploration du Parc National Albert, Mission H. Dumas (1935-1936), 12, 1–62.
Klugh, A.B. (1923) A new Cyclocypris from eastern Canada. Transactions of the Royal Canadian Institute, 14, 337–342.
Kovalenko, A.L. (1982) Novye vidy roda Cypria (Crustacea, Ostracoda). Izvestiya Akademii Nauk Moldavskoy SSR, Seriya Biologicheskikh i Khimicheskikh Nauk, 3, 38–42.
Kovalenko, A.L. (1987) Bentocypria – Novyy rod nadsemeystva Cypridacea (Crustacea, Ostracoda); Bentocypria – a new genus of the superfamily Cypridacea (Crustacea, Ostracoda). In: Nevesskaya L.A. (Ed.) Stratigrafiya verkhnego fanerozoya Moldavii. (Sbornik Nauchnykh Trudov), Kishinev, 99–105.
Kovalenko, A.L., Nikolaeva, I.A, Pavlovskaya, V.I. & Karmischina, G.I. (1989) Nadsemeyestvo Cypridacea Baird, 1845 (Superfamily Cypridacea Baird, 1845). In: Kovalenko, A.L., Nikolaeva, I.A, Pavlovskaya, V.I., Karmischina, G.I. & Aladin, N.V. (Eds) Ostrakody Kaynozoya (Cenozoic Ostracoda). Prakticheskoe Rukovodsvto po Mikrofaune SSSR (Practical Manual of Microfauna of the USSR). Leningrad (“Nedra”), 3, 92–119.
Krstić, N. (1995) A new Cyclocypris subgenus: Laevicypris. In: Riha, J. (Ed.), Ostracoda and Biostratigraphy. Proceedings of the 12. International Symposium on Ostracoda, Prague, 1994. Balkema, Rotterdam, 37–42.
Krstić, N. (2006) Pliocene ostracodes of the Paludinian Beds in the Pannonian Plain, Serbian Part. Herald of the Natural History Museum, Belgrade, Special publications, 409 pp.
Krstić, N. & Keyser, D. (2008) The tribe Cypriini Kovalenko (Ostracoda, Crustacea) from the Upper Pliocene of Metohia (Kosovo, Serbia). Bulletin de l’Academie serbe des sciences et des arts, Classe des Sciences mathématiques et naturelles, Sciences naturelles, 44, 57–83.
Külöylüoğlu, O. (2008) Cyclocypris vinyardi sp. n. (Ostracoda) described from a rheocrene spring in Nevada (U.S.A.). Crustaceana, 81, 385–402.
Maddocks, R.F. (1992) Anchialine Cyprididae (Ostracoda) from the Galapagos Islands, with a review of the subfamily Paracypridinae. Zoological Journal of the Linnean Society, 104, 1–29.
Maddocks, R.F. (2000) The antennula in podocopid Ostracoda: Chaetotaxy, ontogeny, and morphometries. Micropaleontology, 2, 1–37.
Margalef, R. (1961) La vida en los charcos de agua dulce de Nueva Esparta (Venezuela). Memoria de la Sociedad de Ciencias Naturales La Salle, 21, 75–110.
Marmonier P., Meisch C. & Danielopol D.L. (1989) A review of genus Cavernocypris Hartmann (Ostracoda, Cypridopsinae): systematics, ecology and biogeography. Bulletin de la Société des naturalistes luxembourgeois, 89, 221–278.
Martens, K. (1982) On a small collection of freshwater ostracods (Crustacea Ostracoda) from Somalia, with description of two new species. Italian Journal of Zoology, 5, 149–170.
Martens, K. (1987) Homology and functional morphology of the sexual dimorphism in the antenna of Sclerocypris Sars, 1924 (Crustacea, Ostracoda, Megalocypridinae). Bijdragen tot de Dierkunde, 57, 183–190.
Martens, K. (1992) On Namibcypris costata n. gen., n. sp. (Crustacea, Ostracoda, Candoninae) from a spring in northern Namibia, with the description of a new tribe and a discussion on the classification of the Podocopina. Stygologia, 7, 27–42.
Martens, K. (1997) Speciation in ancient lakes. Tree, 12, 177–182.
Martens, K & Schön, I. (1999) Crustacean biodiversity in ancient lakes: a review. Crustaceana, 72, 899–910.
Martens, K., Horne, D.G. & Griffiths, H.I. (1998) Age and diversity of non-marine ostracods. In: Martens, K. (Ed.), Sex and parthenogenesis: evolutionary ecology of reproductive modes in non-marine ostracods. Backhuys Publications Leiden, 37–55.
Matzke-Karasz, R. (1995) Aktuelle Gattungs- und Artmerkmale bei Scottia, Cyclocypris, Psychrodromus und Mesocypris (Ostracoda). Geologisches Institut der Universität zu Köln, Sonderveröffentlichungen, 97, 285 pp.
Matzke-Karasz, R., Smith, R.J. & Homma, M. (2004) Cyclocypris diebeli Absolon, 1973 (Ostracoda, Crustacea), extinct in Europe, extant in Japan. Journal of Natural History, 38, 1635–1663.
McKenzie, K. (1971) Species list of South African Freshwater Ostracoda with an appendix listing museum collections and some further determinations. Annals of the South African Museum, 57, 157–213.
McKenzie, K.G. (1972) Check list of Ostracoda recorded from the Indian Subcontinent and Ceylon (1840-1971), Melbourn, 1-3, 1–59.
McKenzie, K.G. (1979) Tasmancypris, a new marine ostracod genus: and a review of the family Paracyprididae (Crustacea: Ostracoda). Papers and Proceedings of the Royal Society of Tasmania, 113, 29–37.
McKenzie, K.G. (1980) A new subfamily from Gorong, Seram, Molucas, with description of Renaudcypris new genus (Crustacea, Ostracoda). Bulletin Muséum national d'Histoire naturelle, 4, 507–515.
Meisch, C. (1980) Contribution to the taxonomy of Physocypria kraepelini and Cypricercus alacer (Crustacea, Ostracoda). Archives de L’Institut Grand Ducal de Luxembourg Section des Sciences naturelles, physiques et mathématiques, 39, 43–53.
Meisch, C. (1987) Ostracodes d'eau douce récoltés dans le sud-ouest de la France. Bulletin de la Société des naturalistes luxembourgeois, 87, 89–118.
Meisch, C. (1996) Contribution to the taxonomy of Pseudocandona and four related genera, with the description of Schellencandona nov. gen., a list of the Candoninae genera, and a key to the European genera of the subfamily (Crustacea, Ostracoda). Bulletin de la Société des Naturalistes luxembourgeois, 97, 211–237.
Meisch, C. (2000) Freshwater Ostracoda of Western and Central Europe. In: Schwoerbel, J & Zwick, P (Eds), Süßwasserfauna von Mitteleuropa. Spektrum Akademischer Verlag, Heidelberg, Berlin, 8/3, 1–522.
Meisch, C. (2007) On the origin of the putative furca of the Ostracoda (Crustacea). Hydrobiologia, 585, 181–200.
Minati K., Cabral, M.C., Pipik, R., Danielopol, D.L., Linhart, J. & Neubauer, W. (2008) Morphological variability among European populations of Vestalenula cylindrica (Straub) (Crustacea, Ostracoda). Palaeogeography, Palaeoclimatology, Palaeoecology, 264, 296–305.
Müller, G.W. (1898) Die Ostracoden. Abhandlungen der Senckenbergischen naturforschenden Gesellschaft, 21, 257–290.
Müller, G.W. (1906) Ostracoda aus Java. Mitteilungen aus dem Naturhistorischen Museum in Hamburg, 23, 138–142.
Neil, J.V. (2000) Factors influencing intraspecific variation and polymorphism in marine podocopid Ostracoda, with particular reference to Tertiary species from southern Australia. Hydrobiologia, 419, 161–180.
Okubo, I. (1990) Seven new species of freshwater ostracoda from Japan (Freshwater Ostracoda from Japan XIV). Research on Crustacea, 19, 1–12.
Petkovski, T.K. (1976) Zwei neue und eine seltene Ostracoden-Art der Gattung Cypria Zenker aus Jugoslawien. Acta Musei Macedonici Scientiarum Naturalium, 7, 173–192.
Petkovski, T.K. & Meisch, C. (1994) Cypria bicolor n. sp., a new crenobiont freshwater ostracod (Crustacea, Ostracoda) from Slovenia. Bulletin de la Société des Naturalistes luxembourgeois, 95, 229–236.
Redeke, H.C. (1936) Ostracoda. Flora en Fauna der Zuiderzee, Supplement, 1936, 97–112.
Roberts, J.H., Holmes, J.A. & Swan, A.R.H. (2002) Ecophenotypy in Limnocythere inopinata (Ostracoda) from the late Holocene of Kajemarum Oasis (north-eastern Nigeria). Palaeogeography, Palaeoclimatology, Palaeoecology, 185, 41–52.
Rome, R. (1962) Ostracodes. - Exploration Hydrobiologique du Lac Tanganyika (1946-1947). Résultats Scientifiques 3, 1–305.
Rome, R. (1965) Ostracodes. Parc National de l’Upemba. – Mission G. F. de Witte, 69, 68 pp.
Sars, G.O. (1895) On some South-African Entomostraca raised from dried mud. Skrifter I Videnskabs-Selskabet, I. Mathematisk-Naturvidenskabs Klasse 1895, 8, 56 pp.
Sars, G.O. (1896) On fresh-water Entomostraca from the neighborhood of Sydney, partly raised from dried mud. Archiv for Mathematik og Naturvidenskab, 18, 1–81.
Sars, G.O. (1901) Contribution to the knowledge of the fresh-water Entomostraca of South America. Archiv for Mathematik og Naturvidenskab, 24, 1–52.
Sars, G.O. (1903) Fresh-water Entomostraca from China and Sumatra. Archiv for Mathematik og Naturvidenskab, 25, 1–44.
Sars, G.O. (1910) Zoological results of the third Tanganyika Expedition, conducted by Dr. W.A. Cunnington, F.Z.S., 1904-1905. Reports on the Ostracoda. Proceedings of the Zoological Society of London, 65, 732–760.
Savatenalinton, S. & Martens, K. (2008) Redescription of Hemicypris mizunoi OKUBO, 1990 (Crustacea, Ostracoda) from Thailand, with a reassessment of the validity of the genera Hemicypris and Heterocypris. Bulletin de l'Institut Royal des Sciences Naturelles de Belgique, Biologie, 78, 17–27.
Sharpe, R.W. (1897) Contribution to a knowledge of the North American fresh-water Ostracoda included in the families Cytheridae and Cyprididae. Illinois State Laboratory of Natural History, 4, 414–484.
Smith, R.J. & Janz, H. (2008) Recent species of the Family Candonidae (Ostracoda Crustacea) from the ancient Lake Biwa, Central Japan. Journal of Natural History, 42, 2865–2922.
Smith, R.J. & Kamiya, T. (2006) Six new species of fresh and brackish water ostracods (Crustacea) from Yakushima, Southern Japan. Hydrobiologia, 559, 331–355.
Stock, J.H & Von Vaupel Klein, J.C. (1996) Mounting media revisited: the suitability of Reyne’s fluid for small crustaceans. Crustaceana, 69, 794–798.
Tressler, W.L. (1937) Ostracoda. Internationale Revue der gesamten Hydrobiologie und Hydrographie, 34, 188–207.
Turner, C.H. (1893) Fresh-water Ostracoda of the United States. Geological and Natural History survey of Minnesota, Zoological Series, 2, 277–337.
Vávra, V. (1891) Monographie der Ostracoden Böhmens. Archiv für die naturwissenschaftliche Landesdurchforschung von Böhmen, 8, 1–116.
Vávra, V. (1897) Die Süsswasser-Ostracoden Deutsch-Ost-Afrikas. In: K. Möbius (Ed.) Deutsch Ost-Afrika. Vol. 4: Die Thierwelt Ost-Afrikas und der Nachbargebiete, Wirbellose Thiere, Verlag Dietrich Reimer, Berlin, 1–28.
Victor, R. & Fernando, C.H. (1979) The freshwater ostracods (Crustacea: Ostracoda) of India. Records of the Zoological Survey of India, 74, 147–242.
Victor, R. & Fernando, C.H. (1981) Redescription of three cyclocypridinae freshwater ostracods (Crustacea, Ostracoda) from Southeast Asia. Journal of Natural History, 15, 757–768.
Victor, R. & Michael, R.G. (1975) Nine new species of freshwater ostracoda from Madurai area in South India. Journal of Natural History, 9, 361–376.
Wagenleitner, H. (1990) Morphology and evolution of Cypria cavernae n. sp. (Ostracoda, Crustacea). Bulletin de la Société des naturalistes luxembourgeois, 90, 199–226.
Wouters, K. (1999) Two new species of the genus Phlyctenophora Brady, 1880 (Crustacea, Ostracoda) from the Indo-Pacific Region. Bulletin de l'Institut Royal des Sciences Naturelles de Belgique, Biologie, 69, 83–92.
Würdig, N.L. & Pinto, I.D. (1993) New species of Cyclocypridinae (Ostracoda) from the north coast of the State of Rio Grande do Sul, Brazil. Anais da Academia Brasileira de Ciências, 65, 285–294.
Yin, Y., Geiger, W. & Martens, K. (1999) Effects of genotype and environment on phenotypic variability in Limnocythere inopinata (Crustacea: Ostracoda). Hydrobiologia, 400, 85–114.
Zenker, G.F.W. (1854) Monographie der Ostracoden. Archiv für Naturgeschichte, 20, 1–87.