Abstract
A morphological phylogenetic analysis is conducted of Australasian harvestmen previously included in the family Monoscutidae. Monophyly of Monoscutidae is not supported, and the subfamilies Monoscutinae and Megalopsalidinae are synonymised with the South American subfamily Enantiobuninae. Monoscutidae is re-synonymised with the family Neopilionidae. The analysis also demonstrates the polyphyly of species previously assigned to the genus Megalopsalis. Megalopsalis epizephyros new species, M. eremiotis new species, M. leptekes new species and M. pilliga new species are described and M. serritarsus and M. hoggi are redescribed, all from Australia. Hypomegalopsalis tanisphyros new genus and species is described from Western Australia. Megalopsalis linnaei is transferred to Tercentenarium new genus. Forsteropsalis new genus is established to include species from New Zealand (including Auckland Island): Macropsalis chiltoni (type species), Pantopsalis distincta, Macropsalis fabulosa, Pantopsalis grayi, Megalopsalis grimmetti, Megalopsalis inconstans, Megalopsalis marplesi, Megalopsalis nigra and Pantopsalis wattsi.References
Banks, N. (1891) A new genus of Phalangiidae. Proceedings of the Entomological Society of Washington, 2, 249–250.
Boyer, S.L. & Giribet, G. (2007) A new model Gondwanan taxon: systematics and biogeography of the harvestman family Pettalidae (Arachnida, Opiliones, Cyphophthalmi), with a taxonomic revision of genera from Australia and New Zealand. Cladistics, 23, 337–361.
Canestrini, G. (1871) Nuove specie di Opilionidi Italiani. Bullettino della Società Entomologica Italiana, 3(4), 381–385.
Cokendolpher, J.C. (2007a) Neopilionidae Lawrence, 1931. In: Pinto-da-Rocha, R., Machado, G. & Giribet, G. (eds), Harvestmen: the Biology of Opiliones. Harvard University Press, Cambridge (Massachusetts) and London, pp. 121–123.
Cokendolpher, J.C. (2007b) Protolophidae Banks, 1893. In: Pinto-da-Rocha, R., Machado, G. & Giribet, G. (eds), Harvestmen: the Biology of Opiliones. Harvard University Press, Cambridge (Massachusetts) and London, pp. 127–128.
Cokendolpher, J.C. & Lanfranco L., D. (1985) Opiliones from the Cape Horn Archipelago: new southern records for harvestmen. Journal of Arachnology, 13(3), 311–319.
Cokendolpher, J.C. & Taylor, C.K. (2007) Monoscutidae. In: Pinto-da-Rocha, R., Machado, G. & Giribet, G. (eds), Harvestmen: the Biology of Opiliones. Harvard University Press, Cambridge (Massachusetts) and London, pp. 118–121.
Cokendolpher, J.C., Tsurusaki, N., Tourinho, A.L., Taylor, C.K., Gruber, J. & Pinto-da-Rocha, R. (2007) Eupnoi: Historical systematic synopsis. In: Pinto-da-Rocha, R., Machado, G. & Giribet, G. (eds), Harvestmen: the Biology of Opiliones. Harvard University Press, Cambridge (Massachusetts) and London, pp. 108–114.
Contreras-Medina, R. & Luna Vega, I. (2002) On the distribution of gymnosperm genera, their areas of endemism and systematic biogeography. Australian Systematic Botany, 15, 193–203.
Cox, C.B. (2001) The biogeographic regions reconsidered. Journal of Biogeography, 28(4), 511–523.
Crawford, R.L. (1992) Catalogue of the genera and type species of the harvestman superfamily Phalangioidea (Arachnida). Burke Museum Contributions in Anthropology and Natural History, 8, 1–60.
Crosby, T.K., Dugdale, J.S. & Watt, J.C. (1998) Area codes for recording specimen localities in the New Zealand subregion. New Zealand Journal of Zoology, 25, 175–183.
Curtis, D.J. & Machado, G. (2007) Ecology. In: Pinto-da-Rocha, R., Machado, G. & Giribet, G. (eds.), Harvestmen: the Biology of Opiliones. Harvard University Press, Cambridge (Massachusetts) and London, pp. 280–308.
Cuvier, G. (1795) Description de deux espèces nouvelles d’insectes. Magazin Encyclopédique, nouvelle série, 1, 205–207, pl. 2.
De Geer, C. (1778) Mémoires pour servir à l’histoire des insectes, Tome 7. Pierre Hesselberg, Stockholm, xii + 950 pp., 49 pls.
Dunlop, J.A. (2006) Baltic amber harvestman types (Arachnida: Opiliones: Eupnoi and Dyspnoi). Fossil Record, 9(2), 167–182.
Dunlop, J.A. (2007) Paleontology. In: Pinto-da-Rocha, R., Machado, G. & Giribet, G. (eds), Harvestmen: the Biology of Opiliones. Harvard University Press, Cambridge (Massachusetts) and London, pp. 247–265.
Edgecombe, G.D. (2001) Revision of Paralamyctes (Chilopoda: Lithobiomorpha: Henicopidae), with six new species from eastern Australia. Records of the Australian Museum, 53, 201–241.
Forster, R.R. (1944) The genus Megalopsalis Roewer in New Zealand with keys to the New Zealand genera of Opiliones. Records of the Dominion Museum, 1(1), 183–192.
Forster, R.R. (1948a) A new sub-family and species of New Zealand Opiliones. Records of the Auckland Institute and Museum, 3, 313–318.
Forster, R.R. (1948b) A new genus and species of the family Acropsopilionidae (Opiliones) from New Zealand. Transactions of the Royal Society of New Zealand, 77(1), 139–141.
Forster, R.R. (1949) Australian Opiliones. Memoirs of the National Museum of Victoria, 16, 59–89.
Forster, R.R. (1955) Further Australian harvestmen (Arachnida: Opiliones). Australian Journal of Zoology, 3(3), 354–411.
Forster, R.R. (1964) The Araneae and Opiliones of the subantarctic islands of New Zealand. In: Gressitt, J.L. (ed.), Insects of Campbell Island. Pacific Insects Monograph, 7. B.P. Bishop Museum, Honolulu, pp. 58–115.
Gardner, S.L. & Campbell, M.L. (1992) Parasites as probes for biodiversity. Journal of Parasitology, 78(4), 596–600.
Giribet, G., Edgecombe, G.D., Wheeler, W.C. & Babbitt, C. (2002) Phylogeny and systematic position of Opiliones: a combined analysis of chelicerate relationships using morphological and molecular data. Cladistics, 18, 5–70.
Giribet, G. & Kury, A.B. (2007) Phylogeny and biogeography. In: Pinto-da-Rocha, R., Machado, G. & Giribet, G. (eds), Harvestmen: the Biology of Opiliones. Harvard University Press, Cambridge (Massachusetts) and London, pp. 62–87.
Giribet, G., Rambla, M., Carranza, S., Baguñà, J., Riutort, M. & Ribera, C. (1999) Phylogeny of the arachnid order Opiliones (Arthropoda) inferred from a combined approach of complete 18S and partial 28S ribosomal DNA sequences and morphology. Molecular Phylogenetics and Evolution, 11, 296–307.
Giribet, G., Vogt, L., Pérez González, A., Sharma, P. & Kury, A.B. (2010) A multilocus approach to harvestman (Arachnida: Opiliones) phylogeny with emphasis on biogeography and the systematics of Laniatores. Cladistics, 25, 1–30.
Giribet, G. & Wheeler, W.C. (1999) On gaps. Molecular Phylogenetics and Evolution, 13, 132–143.
Goloboff, P.A., Farris, J.S. & Nixon, K.C. (2008) TNT, a free program for phylogenetic analysis. Cladistics, 24, 774–786.
Gruber, J. (1974) Bemerkungen zur Morphologie und systematischen Stellung von Caddo, Acropsopilio und verwandter Formen (Opiliones, Arachnida). Annalen des Naturhistorischen Museums in Wien, 78, 237–259.
Gruber, J. & Hunt, G.S. (1973) Nelima doriae (Canestrini), a south European harvestman in Australia and New Zealand (Arachnida, Opiliones, Phalangiidae). Records of the Australian Museum, 28, 383–392.
Hickman, V.V. (1957) Some Tasmanian harvestmen of the sub-order Palpatores. Papers and Proceedings of the Royal Society of Tasmania, 91, 65–79.
Hogg, H.R. (1910) Some New Zealand and Tasmanian Arachnidae. Transactions and Proceedings of the New Zealand Institute, new series, 42, 273–283.
Hogg, H.R. (1920) Some Australian Opiliones. Proceedings of the Zoological Society of London, 1920(1), 31–48, pls 1–3.
Huang, D., Selden, P.A. & Dunlop, J.A. (2009) Harvestmen (Arachnida: Opiliones) from the Middle Jurassic of China. Naturwissenschaften, 96(8), 955–962.
Hunt, G.S. (1990) Taxonomic value of spiracle microstructure in the Megalopsalididae (Opiliones, Phalangioidea). In: Koponen, S., Lehtinen, P.T. & Rinne, V. (eds), Proceedings of the XI International Congress of Arachnology, Turku, Finland, 7–12 August 1989. Acta Zoologica Fennica, 190, 187–194.
Hunt, G.S. & Cokendolpher, J.C. (1991) Ballarrinae, a new subfamily of harvestmen from the Southern Hemisphere. Records of the Australian Museum, 43, 131–169.
International Commission on Zoological Nomenclature (1999) International Code of Zoological Nomenclature, fourth edition. International Trust for Zoological Nomenclature, London, 306 pp.
Kauri, H. (1954) Report from Professor T. Gislén’s expedition to Australia in 1951-1952. 9. Harvest-spiders from S. W. Australia. Lunds Universitets Årsskrift Ny Fjöld, Avdelning 2, 50(11), 1–10.
Kauri, H. (1961) Opiliones. In: Hanström, B., Brinck, P. & Rudebeck, G. (eds), South African Animal Life: Results of the Lund University Expedition in 1950-1951, vol. 8. Almquist & Wiksell, Uppsala, pp. 9–197.
Koch, C.L. (1835) [Arachniden], Heft 128. In: Herrich-Schäffer, G.A.W. (ed.), Deutschlands Insecten, Hefte 111–190, 1829–1844. F. Pustet, Regensburg, 24 pls on unnumbered pages.
Koch, C.L. (1839) Übersicht des Arachnidensystems, Zweites Heft. C. H. Zeh’schen Buchhandlung, Nürnberg, 38 pp., 6 pls.
Kury, A.B. (2003) Annotated catalogue of the Laniatores of the New World (Arachida, Opiliones). Revista Ibérica de Aracnología, special monographic volume, 1, 1–337.
Lawrence, R.F. (1931) The harvest-spiders (Opiliones) of South Africa. Annals of the South African Museum, 29(2), 341–508.
Linder, H.P. & Crisp, M.D. (1995) Nothofagus and Pacific biogeography. Cladistics, 11, 5–32.
Linnaeus, C. (1758) Systema naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Editio decima, reformata. Tomus 1. Laurentius Salvius, Stockholm, iv + 824 pp.
Marples, B.J. (1944) A new species of harvestman of the genus Megalopsalis. Transactions of the Royal Society of New Zealand, 73(4), 313–314.
Martens, J. (1976) Genitalmorphologie, System und Phylogenie der Weberknechte (Arachnida: Opiliones). Entomologica Germanica, 3(1–2), 51–68.
Martens, J. (1978) Spinnentiere, Arachnida: Weberknechte, Opiliones. In: Senglaub, K., Hannemann, H.-J. & Schumann, H. (eds), Die Tierwelt Deutschlands, vol. 64. Gustav Fischer Verlag, Jena, 464 pp.
Martens, J. (1986) Die Grossgliederung der Opiliones und die Evolution der Ordnung (Arachnida). In: Barrientos, J.A. (ed.), Actas X Congreso Internacional de Aracnologia (Jaca, Spain, September 1986), vol. 1. Juvenil, Barcelona, pp. 289–310.
Martens, J. (1987) Opiliones aus dem Nepal-Himalaya VI. Gagrellinae (Arachnida: Phalangiidae). Courier Forschungsinstitut Senckenberg, 93, 87–202.
McInnes, S.J. & Pugh, P.J.A. (1998) Biogeography of limno-terrestrial Tardigrada, with particular reference to the Antarctic fauna. Journal of Biogeography, 25, 31–36.
McLoughlin, S. (2001) The breakup history of Gondwana and its impact on pre-Cenozoic floristic provincialism. Australian Journal of Botany, 49, 271–300.
Mello-Leitão, C.F. de (1931) Notas sobre Arachnideos Argentinos. Annaes da Academia Brasileira de Sciencias, 3(2), 83–97, 2 pls.
Meyer-Rochow, V.B. & Liddle, A.R. (1988) Structure and function of the eyes of two species of opilionid from New Zealand glow-worm caves (Megalopsalis tumida: Palpatores, and Hendea myersi cavernicola: Laniatores). Proceedings of the Royal Society of London Series B—Biological Sciences, 233, 293–319.
Michener, C.D. (1979) Biogeography of the bees. Annals of the Missouri Botanical Garden, 66(3), 277–347.
Muona, J. (1991) The Eucnemidae of south-east Asia and the western Pacific—a biogeographical study. Australian Systematic Botany, 4, 165–182.
Nation, J.L. (1983) A new method using hexamethyldisilazane for preparation of soft insect tissues for scanning electron microscopy. Stain Technology, 58(6), 347–351.
Phillipps, W.J. & Grimmett, R.E.R. (1932) Some new Opiliones from New Zealand. Proceedings of the Zoological Society of London, 1932, 731–740.
Pinto-da-Rocha, R. & Gruber, J. (2007) Caddidae Banks, 1893. In: Pinto-da-Rocha, R., Machado, G. & Giribet, G. (eds), Harvestmen: the Biology of Opiliones. Harvard University Press, Cambridge (Massachusetts) and London, pp. 115–118.
Pocock, R.I. (1903a) On some new harvest-spiders of the order Opiliones from the southern continents. Proceedings of the Zoological Society of London, 1902(2), 392–413.
Pocock, R.I. (1903b) Fifteen new species and two new genera of tropical and southern Opiliones. Annals and Magazine of Natural History, series 7, 11(65), 433–450, pls 11–12.
Reid, A.L. (1996) Review of the Peripatopsidae (Onychophora) in Australia, with comments on peripatopsid relationships. Invertebrate Taxonomy, 10, 663–936.
Roewer, C.F. (1910) Revision der Opiliones Plagiostethi (=Opiliones, Palpatores) I. Teil: Familie der Phalangiidae. (Subfamilien: Gagrellini, Leiobunini, Leptobunini). Abhandlungen aus dem Gebiete der Naturwissenschaften, herausgegeben vom Naturwissenschaftlichen Verein in Hamburg, 19(4), 1–294, pls 1–6.
Roewer, C.F. (1911) Übersicht der Genera der Subfamilie der Phalangiini der Opiliones Palpatores nebst Beschreibung einiger neuer Gattungen und Arten. Archiv für Naturgeschichte, part I, 77(suppl. 2), 1–106, pls 1–3.
Roewer, C.F. (1912) Revision der Opiliones Palpatores (=Opiliones Plagiostethi). II. Teil: Familie der Phalangiidae. (Subfamilien: Sclerosomini, Oligolophini, Phalangiini). Abhandlungen aus dem Gebiete der Naturwissenschaften, herausgegeben von Naturwissenschaftlichen Verein in Hamburg, 20(1), 1–295, pls 1–4.
Roewer, C.F. (1923) Die Weberknechte der Erde: Systematisches Bearbeitung der bisher bekannten Opiliones. Gustav Fischer, Jena, v + 1116 pp.
Roewer, C.F. (1956) Über Phalangiinae (Phalangiidae, Opiliones Palpatores). (Weitere Weberknechte XIX). Senckenbergiana biologica, 37(3–4), 247–318.
Sclater, P.L. (1866) Notes upon the American Caprimulgidae. Proceedings of the Zoological Society of London, 1866, 123–145.
Shear, W.A. (1982) Opiliones. In: Parker, S.P. (ed.), Synopsis and Classification of Living Organisms, vol. 2. McGraw-Hill Book Company, New York, pp. 104–110.
Shultz, J.W. (1998) Phylogeny of the Opiliones (Arachnida): An assessment of the “Cyphopalpatores” concept. Journal of Arachnology, 26, 257–272.
Shultz, J.W. & Regier, J.C. (2001) Phylogenetic analysis of Phalangida (Arachnida, Opiliones) using two nuclear protein-coding genes supports monophyly of Palpatores. Journal of Arachnology, 29, 189–200.
Šilhavý, V. (1970) Nouvelles recherches sur la famille des Neopilionidae Lawrence. Bulletin du Muséum National d’Histoire Naturelle, series 2, 41(suppl. 1), 171–175.
Simon, E. (1879) Descriptions d’Opiliones nouveaux. Annales de la Société Entomologique de Belgique, 22(Comptes-Rendus), lxx–lxxv.
Simon, E. (1884) Arachnides recueillis par la mission du Cap Horn en 1882–1883. Bulletin de la Société Zoologique de France, 9, 117–144, pl. 3.
Soares, B.A.M. & Soares, H.E.M. (1947) Alótipos e formas novas de Opiliões Paranaenses (Opiliones—Gonyleptidae, Phalangiidae). Papéis Avulsos do Departamento de Zoologia, 8(5), 63–84.
Sørensen, W. (1886) Opiliones descripsit W. Sørensen. In: Koch, L. & Keyserling, E. von (eds), Die Arachniden Australiens nach der Natur beschrieben und abgebildet, vol. 2. Bauer und Raspe, Nürnberg, pp. 53–86, pls 5–6.
Staręga, W. (2002) Baltic amber harvestmen (Opiliones) from Polish collections. Annales Zoologici, 52(4), 601–604.
Suzuki, S. (1958) A remarkable new phalangid, Ischyropsalis yezoensis (Ischyropsalidae, Opiliones), from Hokkaido. Annotationes Zoologicae Japonenses, 31(3), 167–170.
Swenson, U., Backlund, A., McLoughlin, S. & Hill, R.S. (2001) Nothofagus biogeography revisited with special emphasis on the enigmatic distribution of subgenus Brassospora in New Caledonia. Cladistics, 17, 28–47.
Swenson, U. & Bremer, K. (1997) Pacific biogeography of the Asteraceae genus Abrotanella (Senecioneae, Blennospermatinae). Systematic Botany, 22(3), 493–508.
Taylor, C.K. (2004) New Zealand harvestmen of the subfamily Megalopsalidinae (Opiliones: Monoscutidae)—the genus Pantopsalis. Tuhinga, 15, 53–76.
Taylor, C.K. (2008a) A new species of Monoscutinae (Arachnida, Opiliones, Monoscutidae) from New Zealand, with a redescription of Monoscutum titirangiense. Journal of Arachnology, 36, 176–179.
Taylor, C.K. (2008b) A new species of Monoscutidae (Arachnida, Opiliones) from the wheatbelt of Western Australia. Records of the Western Australian Museum, 24(4), 375–380.
Taylor, C.K. (2009a) Revision of the Australian Gagrellinae (Arachnida: Opiliones: Sclerosomatidae), with a description of a new species. Australian Journal of Entomology, 48, 217–222.
Taylor, C.K. (2009b) Australiscutum, a new genus of Monoscutidae (Arachnida: Opiliones) from eastern Australia, with the first record of asymmetrical chelicerae in Opiliones. Insect Systematics and Evolution, 40, 319–332.
Taylor, C.K. & Hunt, G.S. (2009) New genus of Megalopsalidinae (Arachnida: Opiliones: Monoscutidae) from north-eastern Australia. Zootaxa, 2130, 41–59.
Tourinho, A.L. (2007) Sclerosomatidae Simon, 1879. In: Pinto-da-Rocha, R., Machado, G. & Giribet, G. (eds), Harvestmen: the Biology of Opiliones. Harvard University Press, Cambridge (Massachusetts) and London, pp. 128–131.
Vinnersten, A. & Bremer, K. (2001) Age and biogeography of major clades in Liliales. American Journal of Botany, 88(9), 1695–1703.
Weidner, H. (1959) Die entomologischen Sammlungen des Zoologischen Staatsinstituts und Zoologischen Museums Hamburg. 1. Teil, Pararthropoda und Chelicerata, I. Mitteilungen aus dem Hamburgischen Zoologischen Museum und Institut, 57, 89–142.
Weston, P.H. & Crisp, M.D. (1994) Cladistic biogeography of waratahs (Proteaceae: Embothrieae) and their allies across the Pacific. Australian Systematic Botany, 7, 225–249.
White, A. (1849) Descriptions of apparently new species of Aptera from New Zealand. Proceedings of the Zoological Society of London, 17, 3–6 (reprinted 1850, Annals and Magazine of Natural History, series 2, 5, 50–53).