Skip to main content Skip to main navigation menu Skip to site footer
Type: Articles
Published: 2010-11-03
Page range: 29–50
Abstract views: 51
PDF downloaded: 1

On the identity of Monsieur Dussumier’s Dutch tortoise and the lectotype of Testudo dussumieri Gray, 1831

Seychelles Islands Foundation (SIF), c/o P.O. Box 321, Mahé, Seychelles
Reptilia Aldabra tortoise lectotype prevailing name Seychelles Testudo dussumieri Testudo elephantina Testudo gigantea

Abstract

Although it was forgotten for over a century, the binomen Testudo dussumieri Gray, 1831, is an available name, and the specimen RMNH 3231 deposited in the natural history museum of Leiden – evidently one of the two original syntypes of T. dussumieri – has been designated as the lectotype of this taxon. Recently several authors have actively promoted this as the name-bearing type for the Aldabra tortoise, escalating debates in which this chelonian has been immersed for nearly two decades. This lectotype designation is highly significant to nomenclatural and taxonomic disputes regarding tortoises (Testudinidae), living and extinct, from the western Indian Ocean; and an attempt has been made in this paper to compile all information relevant to the lectotype as well as to better understand the history of the binomen applied to it. Several critical aspects of the history are uncertain and open to speculation. The provenance of RMNH 3231 is unknown and unlikely to be Aldabra Atoll; the specimen was most likely collected in the granitic Seychelles, between 1823 and 1829. The combination of estimated date and locality of collection raises the possibility that the lectotype is not an Aldabra tortoise, but rather an extinct taxon from the granitic Seychelles. It is concluded that RMNH 3231 is not a suitable name-bearing type for the Aldabra tortoise, and the continued use of the name T. dussumieri will cause persistent nomenclatural and taxonomic confusion and unending debate.

References

  1. Adler, K. (2007)* The development of systematic reviews of the turtles of the world. Vertebrate Zoology, 57(2), 139–148.

    Anon. (1823) Naufrage dans l’archipel des Seychelles, en 1808, par M. de C…. Journal des Voyages, Découvertes et Navigations Modernes; ou Archives Géographiques du XIXe siècle, 18, 359–378. [note: the correct title of this publication ends with “… par M. de C ….”, evidently because the author intended to keep his/her name anonymous.]

    Anon. (1875) Biographical notice of the late Dr. John Edward Gray. Annals of the Magazine of Natural History, series 4, 15, 281–285.

    Arnold, E.N. (2009)* Comments on the proposed conservation of usage of Testudo gigantea Schweigger, 1812 (currently Geochelone (Aldabrachelys) gigantea) (Reptilia, Testudines) (Case 3463; see BZN 66: 34–50, 80–87). Bulletin of Zoological Nomenclature, 66(2), 177.

    Arvy, L. (1972) Jean-Jacques Dussumier, master mariner and cetologist (1792–1883). Investigations on Cetacea, 4, 263–269, pls. 1–5.

    Austin, J.J., Arnold, E.N. & Bour, R. (2003) Was there a second adaptive radiation of giant tortoises in the Indian Ocean? Using mitochondrial DNA to investigate speciation and biogeography of Aldabrachelys (Reptilia, Testudinidae). Molecular Ecology, 12, 1415–1424.

    Balmer, O., Ciofi, C., Galbraith, D.A., Swingland, I.R., Zug, G.R. & Caccone, A. (2010) Population genetic structure of Aldabra giant tortoises. Journal of Heredity, doi:10.1093/jhered/esq096

    Barley, A.J., Spinks, P.Q., Thompson, R.C. & Shaffer, H.B. (2010)* Fourteen nuclear genes provide phylogenetic resolution for difficult nodes in the turtle tree of life. Molecular Phylogenetic and Evolution, 55, 1189–1194.

    Boessenkool, S., Star, B., Scofield, R.P., Seddon, P.J. & Waters, J.M. (2010) Lost in translation or deliberate falsification? Genetic analyses reveal erroneous museum data for historic penguin specimens. Proceedings of the Royal Society, B, 227, 1957–1064.

    Boulenger, G.A. (1889)* Catalogue of the Chelonians, Rhynchocephalians, and Crocodiles in the British Museum (Natural History). Taylor and Francis, London, x, 311 pp., 6 pls.

    Bour, R. (1982) Contribution à la connaissance des tortues terrestres des Seychelles: définition du genre endémique et description d’une espèce nouvelle probablement originaire des îles granitiques et au bord de l’extinction. Comptes Rendus Hebdomadaires Séances de l’Académie des Sciences, Paris, 295, Sér. III, 117–118, 121–122, 1 pl.

    Bour, R. (1984a) L’identité de Testudo gigantea Schweigger, 1812 (Reptilia, Chelonii). Bulletin du Muséum National d’Histoire Naturelle, Paris, 4 Sér., 6, section A(1), 159–175.

    Bour, R. (1984b) Taxonomy, history and geography of Seychelles land tortoises and fresh-water turtles. In: Stoddart, D. R. (Ed.). Biogeography and Ecology of the Seychelles Islands. Dr. W. Junk, The Hague, pp. 281–307.

    Bour, R. (1985 [1984]) Les tortues terrestres géantes des îles de l’océan Indien occidental: données géographiques, taxinomiques et phylogénétiques. In: F. de Broin and E. Jiménez-Fuentes (Eds.) Comunicaciones del I Simposium Internacional sobre Quelonios Fósiles, París, Octubre, 1983. Studia Geologica Salmanticensia, Volumen Especial 1 (Studia Palaeocheloniologica I) [1984], pp. 17–76.

    Bour, R. (1988 [1987]) Tortues et insularité: les tortues des Seychelles. Bulletin de la Société Zoologique de France, 112, 401–418.

    Bour, R. (1994) Recherches sur des animaux doublement disparus: les tortues géantes subfossiles de Madagascar. Mémoires et Travaux de l’Institut de Montpellier, 19, 1–254.

    Bour, R. (2006a) An unnamed tortoise from the Seychelles Islands. Emys, 13(3), 24–30.

    Bour, R. (2006b) Identity of Testudo gigantea Schweigger, 1812 and rediscovery of the type specimen. Emys, 13(4), 12–23.

    Bour, R. & Pritchard, P.C.H. (2009)* Comments on the proposed conservation of usage of Testudo gigantea Schweigger, 1812 (currently Geochelone (Aldabrachelys) gigantea) (Reptilia, Testudines) (Case 3463; see BZN 66: 34–50, 80–87). Bulletin of Zoological Nomenclature, 66(2), 169–174.

    Bour, R., Iverson, J.B. & Pritchard, P.C.H. (2010a)* Comments on the proposed conservation of usage of Testudo gigantea Schweigger, 1812 (currently Geochelone (Aldabrachelys) gigantea) (Reptilia, Testudines) (Case 3463; see BZN 66: 34–50, 80–87, 169–186; 274–290; 352–357). Bulletin of Zoological Nomenclature, 67(1), 72–73.

    Bour, R., Pritchard, P.C.H. & Iverson, J.B. (2010b)* Comments on the proposed conservation of usage of Testudo gigantea Schweigger, 1812 (currently Geochelone (Aldabrachelys) gigantea) (Reptilia, Testudines) (Case 3463; see BZN 66: 34–50, 80–87, 169–186; 274–290; 352–357). Bulletin of Zoological Nomenclature, 67(1), 73–77.

    Braithwaite, C.J.R. (1984) Geology of the Seychelles. In: Stoddart, D. R. (Ed.). Biogeography and Ecology of the Seychelles Islands. Dr. W. Junk, The Hague, pp. 17–38.

    Buskirk, J. (2009)* Comments on the proposed conservation of usage of Testudo gigantea Schweigger, 1812 (currently Geochelone (Aldabrachelys) gigantea; Reptilia, Testudines) (Case 3463; see BZN 66: 34–50, 80–87, 169–186). Bulletin of Zoological Nomenclature, 66(3), 286.

    Cheke, A. (2009)* Comments on the proposed conservation of usage of Testudo gigantea Schweigger, 1812 (currently Geochelone (Aldabrachelys) gigantea) (Reptilia, Testudines) (Case 3463; see BZN 66: 34–50, 80–87). Bulletin of Zoological Nomenclature, 66(2), 174–176.

    Cheke, A. (2010)* Comments on the proposed conservation of usage of Testudo gigantea Schweigger, 1812 (currently Geochelone (Aldabrachelys) gigantea) (Reptilia, Testudines) (Case 3463; see BZN 66: 34–50, 80–87, 169–186; 274–290; 352–357). Bulletin of Zoological Nomenclature, 67(1), 79–81.

    Coote, J. (2001) The history of western herpetoculture before the 20th century. In: Proceedings of the International Herpetological Symposium on Captive Propagation and Husbandry, 25, 19–47.

    Cowan, C.F. (1969) Notes on Griffith’s Animal Kingdom of Cuvier (1825–1835). Journal of the Society for the Bibliography of Natural History, 5(2), 137–140.

    Crumly, C.R. (1988)* A nomenclatural history of tortoises (family Testudinidae). Smithsonian Herpetological Information Service, 75, 1–17.

    Dubois, A. (2010a)* Nomenclatural rules in zoology as a potential threat against natural history museums. Organisms Diversity & Evolution, 10, 81–90.

    Dubois, A. (2010b)* Zoological nomenclature in the century of extinctions: priority vs. ‘usage’. Organisms Diversity & Evolution,10, 259–274.

    Dubois, A., Ohler, A. & Brygoo, E.-R. (2010)* Comments on the proposed conservation of usage of Testudo gigantea Schweigger, 1812 (currently Geochelone (Aldabrachelys) gigantea) (Reptilia, Testudines) (Case 3463; see BZN 66: 34–50, 80–87, 169–186; 274–290; 352–357). Bulletin of Zoological Nomenclature, 67(1), 82–89.

    Duméril, A.M.C. & Bibron, G. (1835)* Erpétologie générale ou Histoire naturelle complète des Reptiles. Vol. 2, L’histoire de toutes les espèces de l’ordre des tortues ou chéloniens, et les généralités de celui des lizards ou sauriens. Roret, Paris. ii, 680 pp., pls. xi–xxiv.

    Duméril, A.M.C. & Duméril, A.H.A. (1851) Catalogue Méthodique de la Collection des Reptiles. Muséum d’Histoire Naturelle de Paris/Gide et Baudry, Paris, iv, 224 pp.

    Engelmann, W. (1846)* Bibliotheca Historico-Naturalis. Verzeichniss der Bücher über Naturgeschichte welche in Deutschland, Scandinavien, Holland, England, Frankreich, Italien und Spanien in den Jahren 1700–1846 erschienen sind, Vol. 1. Wilhelm Engelmann, Leipzig, viii, 786 pp.

    Fitzinger, L.J.F.J. (1835 [1836]) Entwurf einer systematischen Anordnung der Schildkröten, nach den Grundsätzen der natürlichen Methode. Annalen des Wiener Museums der Naturgeschichte, 1, 103–128.

    Flower, W.H. (1883) On the characters and divisions of the family Delphinidae. Proceedings of the Zoological Society of London, 1883, 466–513.

    Fontes da Costa, P. (2002)* The culture of curiosity at The Royal Society in the first half of the eighteenth century. Notes and Records of the Royal Society of London, 56(2), 147–166.

    Foster, W. (Ed.) (1905) The Journal of John Jourdain,1608–1617, describing his Experiences in Arabia, India, and the Malay Archipelago. Hakluyt Society, Cambridge, UK, lxxxii, 394 pp.

    Frazier, J. (2006a)* A neotype for the Aldabra tortoise, Testudo gigantea Schweigger, 1812. Herpetological Review, 37(3), 275–280.

    Frazier, J. (2006b) Book Review: “Giant Tortoises of the Indian Ocean. The Genus Dipsochelys Inhabiting the Seychelles Islands and the Extinct Giants of Madagascar and the Mascarenes.” Herpetological Review, 37(3), 368–373.

    Frazier, J. (2009)* Case 3463. Testudo gigantea Schweigger, 1812 (currently Geochelone (Aldabrachelys) gigantea; Reptilia, Testudines): proposed conservation of usage of the specific name by maintenance of a designated neotype, and suppression of Testudo dussumieri Gray, 1831 (currently Dipsochelys dussumieri). Bulletin of Zoological Nomenclature, 66(1), 34–50.

    Fritz, U. & Havaš, P. (2007)* Checklist of chelonians of the world. Vertebrate Zoology, 57, 149–368.

    Fritz, U., Hundsdörfer, A.K., Široký, P., Auer, M., Kami, H., Lehmann, J., Mazanaeva, L.F., Türkozan, O. & Wink, M. (2007)* Phenotypic plasticity leads to incongruence between morphology-based taxonomy and genetic differentiation in western Palaearctic tortoises (Testudo graeca complex; Testudines, Testudinidae). Amphibia-Reptilia, 28, 97–121.

    Gerlach, J. (1997a)* Chelonia and people in Seychelles. Testudo (British Chelonia Group), 4(4), 25–30.

    Gerlach, J. (1997b) Seychelles Red Data Book-1997. The Nature Protection Trust of Seychelles, Victoria, Seychelles, [ii], 130 pp.

    Gerlach, J. (1998a) Famous Tortoises. J. Gerlach, Cambridge, England, [iii], 51 pp.

    Gerlach, J. (1998b) The rediscovery and conservation of Seychelles giant tortoises. International Zoo News, 45(1), 4–10.

    Gerlach, J. (1999a) Distinctive neural bones in Dipsochelys giant tortoises: Structural and taxonomic characters. Journal of Morphology, 240, 33–37.

    Gerlach, J. (1999b) Seychelles reptiles & amphibians. Madagascar and Mascarene Reptiles and Amphibians, 1, 8–10.

    Gerlach, J. (1999c) Feeding behavior and the saddleback shell of Dipsochelys arnoldi. Chelonian Conservation and Biology, 3(3), 496–500.

    Gerlach, J. (1999d) Notes on some Seychelles reptiles. Bulletin of the British Herpetological Society, 67, 53–60.

    Gerlach, J. (1999e) Productivity and energetics of giant tortoises on Aldabra, a reappraisal of existing data. Phelsuma, 7, 74–75.

    Gerlach, J. (2001a) Tortoise phylogeny and the ‘Geochelone’ problem. Phelsuma, 9 (suppl. A), 1–24.

    Gerlach, J. (2001b) Population dynamics of insular vertebrates, examples from the Seychelles islands. Biota, 2(2), 125–135.

    Gerlach, J. (2002) Seychelles Giant Tortoise conservation project patron: Sir David Attenborough. Seychelles Giant Tortoise News (The Nature Protection Trust of Seychelles) No. 13.

    Gerlach, J. (2003a) Ausgestorben geglaubte Riesenschildkröten wieder entdeckt – Herkunft und Erhaltung der Seychellen-Riesenschildkröten. elaphe, Rheinbach, 11(2), 57–61. [translated by Lucian Heichler].

    Gerlach, J. (2003b) Captive breeding of Dipsochelys giant tortoises. Phelsuma, 11, 8–12.

    Gerlach, J. (2004a) Giant Tortoises of the Indian Ocean. The Genus Dipsochelys inhabiting the Seychelles Islands and the extinct giants of Madagascar and the Mascarenes. Chimaira, Frankfurt am Main, 207 pp.

    Gerlach, J. (2004b) Effects of diet on the systematic utility of the tortoise carapace. African Journal of Herpetology, 53(1), 77–85.

    Gerlach, J. (2005) Thermoregulation in captive Indian Ocean giant tortoises. Chelonian Conservation and Biology, 4(4), 937–941.

    Gerlach, J. (2009)* Comments on the proposed conservation of usage of Testudo gigantea Schweigger, 1812 (currently Geochelone (Aldabrachelys) gigantea) (Reptilia, Testudines) (Case 3463; see BZN 66: 34–50, 80–87). Bulletin of Zoological Nomenclature, 66(2), 184–186.

    Gerlach, J. & Bour, R. (2003) Morphology of hatchling Dipsochelys giant tortoises. Radiata English, 12(3), 11–20.

    Gerlach, J. & Canning, K.L. (1995a) The Seychelles giant tortoise, its rediscovery and prospects for conservation. In: Devaux, B. (Ed.) Proceedings of the International Congress on Chelonian Conservation, Gonfaron, France SOPTOM, Gonfaron, pp. 133–135.

    Gerlach, J. & Canning, K.L. (1995b) The Seychelles giant tortoise rediscovered? Oryx, 29(2), 74–75.

    Gerlach, J. & Canning, K.L. (1996) Evolution and history of the giant tortoises of the Aldabra island group. Testudo (British Chelonia Group), 4(3), 33–40.

    Gerlach, J. & Canning, K.L. (1998a) Taxonomy of Indian Ocean giant tortoises (Dipsochelys). Chelonian Conservation and Biology, 3(1), 3–19.

    Gerlach, J. & Canning, K.L. (1998b) Identification of Seychelles Giant Tortoises. Linnaeus Fund Research Report. Chelonian Conservation and Biology, 3(1), 133–135.

    Gerlach, J., Muir, C. & Richmond, M.D. (2006) The first substantiated case of trans-oceanic tortoise dispersal. Journal of Natural History, 40(41–43), 2403–2408.

    Gray, J.E. (1831a) A synopsis of the species of the Class Reptilia. In: Griffith, E. & Pidgeon, E. (Eds.), The animal kingdom arranged in conformity with its organization, by the Baron Cuvier. Vol. 9: The class Reptilia arranged by the Baron Cuvier, with specific descriptions. Whittaker, Treacher, and Co., London, pp. 1–110 [page numbers duplicated].

    Gray, J.E. (1831b)* Synopsis Reptilium; or short descriptions of the species of reptiles. Pt. I. Cataphracta, tortoises, crocodiles, and enaliosaurians. Treuttel, Wurtz and Co., London, viii, 85 pp., 10 pls.

    Gray, J.E. (1844)* Catalogue of the Tortoises, Crocodiles, and Amphisbænians, in the Collection of the British Museum. Trusties of the British Museum, London, viii, 80 pp.

    Gray, J.E. (1855)* Catalogue of Shield Reptiles in the Collection of the British Museum. Part I. Testudinata (Tortoises). Trusties of the British Museum, London, [iii], 79, [ii] pp., 42 pls.

    Gray, J.E. (1869)* Notes on the families and genera of tortoises (Testudinata), and on the characters afforded by a study of their skulls. Proceedings of the Zoological Society of London, 1869, 165–225, pl. 15.

    Gray, J.E. (1870)* Supplement to the Catalogue of Shield Reptiles in the Collection of the British Museum. Part I. Testudinata (Tortoises). Trusties of the British Museum/Taylor and Francis, London, x, 120 pp.

    Gray, J.E. (1872)* Appendix to the Catalogue of Shield Reptiles in the Collection of the British Museum. Part I. Testudinata (Tortoises). Trusties of the British Museum/Taylor and Francis, London, [iv], 28 pp.

    Gray, J.E. (1873)* Hand-list of the Specimens of Shield Reptiles in the British Museum. Trusties of the British Museum, London, iv, 124 pp.

    Gray, J.E. (1875) List of books, memoirs, and miscellaneous papers by John Edward Gray, with a few historic notes. Taylor and Francis, London, 58 pp. [printed for private distribution, completed after Gray’s death by John Saunders].

    Grünewald, F. (2009)* Museumcollecties...RMNH 3231 (Dipsochelys dussumieri) Gray, 1831. Trionyx, 7(5), 136–142.

    Grünewald, F. (2010)* Comments on the proposed conservation of usage of Testudo gigantea Schweigger,1812 (currently Geochelone (Aldabrachelys) gigantea) (Reptilia, Testudines) (Case 3463; see BNZ 66: 34–50, 80–87, 169–186, 274–290, 352–357; 67: 71–90). Bulletin of Zoological Nomenclature, 67(2), 177–178.

    Günther, A.C.L.G. (1877)* The Gigantic Land-tortoises (Living and Extinct) in the Collection of the British Museum. Taylor and Francis, London, v, 96 pp., 54 pls.

    Holthuis, L.B. & Sakai, T. (1970) Ph. F. von Siebold and Fauna japonica: A history of early Japanese zoology. Academic Press of Japan, Tokyo, 323 pp.

    Hoogmoed, M.S. (2009)* Comments on the proposed conservation of usage of Testudo gigantea Schweigger, 1812 (currently Geochelone (Aldabrachelys) gigantea; Reptilia, Testudines) (Case 3463; see BZN 66: 34–50, 80–87, 169–186). Bulletin of Zoological Nomenclature, 66(4), 354–356.

    Hoogmoed, M.S., Gassó Miracle, M.E. & van den Hoek Ostende, L.W. (2010)* Type specimens of recent and fossil Testudines and Crocodylia in the collections of the Netherlands Centre for Biodiversity Naturalis, Leiden, the Netherlands. Zoologische Mededelingen Leiden, 84, in press.

    Hubrecht, A.A.W. (1881)* On certain Tortoises in the collections of the Leyden Museum. Notes from the Leyden Museum, 3(12), 41–50.

    Ibáñez, N., Montserrat, J.M., Soriano, I. & Camarasa, J.M. (2006)* Plant material exchanged between James Petiver (ca. 1663–1718) and Joan Salvador (1683–1725). I. The Balearic plants conserved in the BC-Salvador and BM-Sloane Herbaria. Notes & Records of the Royal Society, 2006, 60, 241–248.

    ICZN [International Commission on Zoological Nomenclature] (1999)* International Code of Zoological Nomenclature. Fourth Edition. International Trust for Zoological Nomenclature, London, XXIX, 306 pp.

    Iverson, J.B. (1992) A Revised Checklist with Distribution Maps of the Turtles of the World. Privately Published, Richmond, Indiana, xiii, 363 pp.

    Iverson, J.B. (2009)* Comments on the proposed conservation of usage of Testudo gigantea Schweigger, 1812 (currently Geochelone (Aldabrachelys) gigantea; Reptilia, Testudines) (Case 3463; see BZN 66: 34–50, 80–87, 169–186). Bulletin of Zoological Nomenclature, 66(3), 284–285.

    James, K.A. (2004) “Humbly dedicated”: Petiver and the audience for natural history in early eighteenth-century Britan. Archives of Natural History, 31(2), 318–329.

    King, F.W. & Burke, R.I. (Eds.). (1989) Crocodilian, Tuatara and Turtle species of the World: A taxonomic and geographic reference. Association of Systematics Collections, Washington D.C., xxii, 216 pp.

    Laissus, Y. (1973) Notes sur les voyages de Jean-Jacques Dussumier (1792–1883). Annales de la Société des Sciences Naturelles de la Charente-Maritime, 5(5–9), 387–406.

    Lamette, C. (1886)* Mouvement géographique; Correspondance & faits géographiques. Bulletin de la Société Normande de Géographie, 8, 170–178.

    Lawrence, T., Pelkey, N. & Soares, S. (2010) Comment on the proposed conservation of usage of Testudo gigantea Schweigger, 1812 (currently Geochelone (Aldabrachelys) gigantea) (Reptilia, Testudines) (Case 3463; see BZN 66: 34–50, 80–87, 169–186, 274–290, 352–357; 67: 71–90, 170–178). Bulletin of Zoological Nomenclature, 67(3), 246–254.

    Le, M., Raxworthy, C.J., McCord, W.P. & Mertz, L. (2006) A molecular phylogeny of tortoises (Testudines: Testudinidae) based on mitochondrial and nuclear genes. Molecular Phylogenetics and Evolution, 40, 517–531.

    Linnæus, C. von (1758)* Systema naturæ per regna tria naturæ, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. (10th ed.) Vol. I. Laurentii Salvii, Holmiae, [1–4], 1–824 pp.

    Lisney, A.A. (1960) A Bibliography of British Lepidoptera 1609–1799. The Chiswick Press, London, xviii, 315 pp.

    Loveridge, A. & Williams, E.E. (1957)* Revision of the African tortoises and turtles of the suborder Cryptodira. Bulletin of the Museum of Comparative Zoology, 115(6), 161–557, 18 pls.

    Matyot, P. (2009)* Comments on the proposed conservation of usage of Testudo gigantea Schweigger, 1812 (currently Geochelone (Aldabrachelys) gigantea; Reptilia, Testudines) (Case 3463; see BZN 66: 34–50, 80–87, 169–186). Bulletin of Zoological Nomenclature, 66(4), 352–354.

    Mertens, R. & Wermuth, H. (1955) Die rezenten Schildkröten, Krokodile und Brückenechsen. Eine kritische Liste der heute lebenden Arten und Rassen. Zoologische Jahrbücher, Abteilung für Systematik, Ökologie und Geographie der Tiere, 83 (5), 323–440.

    Ng, P.K.L. (2009)* Comments on the proposed conservation of usage of Testudo gigantea Schweigger, 1812 (currently Geochelone (Aldabrachelys) gigantea) (Reptilia, Testudines) (Case 3463; see BZN 66: 34–50, 80–87). Bulletin of Zoological Nomenclature, 66(2), 183.

    Palkovacs, E.P., Gerlach, J. & Caccone, A. (2002)* The evolutionary origin of Indian Ocean tortoises (Dipsochelys). Molecular Phylogenetics and Evolution, 24, 216–227.

    Palkovacs, E.P., Marschner, M., Ciofi, C., Gerlach, J. & Caccone, A. (2003)* Are the native giant tortoises from the Seychelles really extinct? A genetic perspective based on mtDNA and microsatellite data. Molecular Ecology, 12, 1403–1413.

    Petiver, J. (1711?)* Gazophylacii Naturæ & Artis Decas Septima & Octava. Bateman, London, 4 pp.

    Petiver, J. (1711)* Catalogus Classicus & Topicus, Omnium Rerum Figuratarum in V Decadibus, Seu Secundo Volumine Gazophylacii Naturæ & Artis Singulis ad proprias Tabulas & Numeros relatis. Bateman, London, pp. 1–4, pls. 51–102.

    Petiver, J. (1764a)* Catalogus Classicus & Topicus, Omnium Rerum Figuratarum in V Decadibus, Seu Secundo Volumine Pp. 1–4 [second section]. In: Opera, Historiam Naturalem Spectantia; or, Gazophylacium. Vol. 1. John Millan, London.

    Petiver, J. (1764b)* Gazophylacii Naturæ & Artis Decas Septima & Octava. Pp. 5–8 [fourth section]. In: Opera, Historiam Naturalem Spectantia; or, Gazophylacium. Vol. 1. John Millan, London.

    Petiver, J. (1764c)* Opera, Historiam Naturalem Spectantia; or, Gazophylacium. Vol. 1. John Millan, London, [ii], pp. 1–4, pp. 1–4, pp. 1–12, pp. 1–10, pls. 1–156, pp. 1–4, pls. 1–22 [# 16 duplicated], pp. i–ii, pp. 1–2.

    Petiver, J. (1767a)* Gazophylacii Naturæ & Artis Decas Septima & Octava. Pp. 5–8 [fourth section]. In: Opera, Historiam Naturalem spectantia (Ed. J. Empson). Vol. 1. John Millian, London.

    Petiver, J. (1767b)* Opera, Historiam Naturalem spectantia (Ed. J. Empson). Vol. 1. John Millian, London, [ii, 1], pp. 1–12, pp. 1–12, pp. 1–10, pp. 1–4, pp. 1–30, pls. 1–156, p. [1].

    Petiver, J. (1767c)* Catalogus Classicus & Topicus, Omnium Rerum Figuratarum in V Decadibus, Seu Secondo Volumine. Pp. 1–4 [first section]. In: Opera, Historiam Naturalem spectantia (Ed. J. Empson).Vol. 2. John Millan, London.

    Plummer, P. (1995) Planet Aldabra. In: Amin, M., Willettts, D. & Skerrett, A. (Eds.) Aldabra: World Heritage Site. Camerapix, Nairobi, pp. 49–70.

    Pritchard, P.C.H. (1986) A reinterpretation of Testudo gigantea Schweigger, 1812. Journal of Herpetology, 20(4), 522–534.

    Rasmussen P.C. & Prys-Jones R.P. (2003) History vs mystery: the reliability of museum specimen data. Bulletin of the British Ornithologists' Club Supplement 123A, 66–94.

    Rothschild, W. (1915)* On the gigantic land tortoises of the Seychelles and Aldabra-Madagascar group with some notes on certain forms of the Mascarene group. Novitates Zoologicae, Tring 22, 418–442, pls. 33–76.

    Schlegel, H. (1837)* Essai sur la physionomie des serpens, Vol. 1, M. H. Schonekat, Libraire-Éditeur, Amsterdam, xxviii, 606 pp.

    Sherborn, C.D. & Jentik, F.A. (1895) On the dates of the parts of Siebold’s ‘Fauna Japonica’ and Giebel’s ‘Allgemeine Zoologie’ (first edition). Proceedings of the Zoological Society of London, 1895, 149–150.

    Siebenrock, F. (1909)* Synopsis der rezenten Schildkröten, mit Berücksichtigung der in historischer Zeit ausgestorbenen Arten. Zoologische Jahrbücher (Systematik), Supplement 10, 427–618.

    Simonton, D.K. (1997) Creative productivity: a predictive and explanatory model of career trajectories. Psychological Review, 104(1), 66–89.

    Simonton, D.K. (1999) Origins of Genius: Darwinian Perspectives on Creativity. Oxford University Press, New York, x, 308 pp.

    Stejneger, L. (1907)* Herpetology of Japan and adjacent territory. Bulletin of the United States National Museum, 58, i–xx, 1–577.

    Stoddart, D.R. & Peake, J.F. (1979) Historical records of Indian Ocean giant tortoise populations. Philosophical Transactions of the Royal Society, London B, 286, 147–161.

    Strauch, A. (1890) Bemerkungen über die Schildkröten-Sammlung im zoologischen Museum der Kaiserlichen Akademie der Wissenschaften zu St. Petersburg. Mémoires de l'Académie Impériale des Sciences de Saint-Pétersbourg, Sciences Naturelles, Sér. 7, 38(2), [2], 1–127, [1], 4 pls.

    Temminck, C.J. & Schlegel, H. (1834)* Reptilia. Les Chéloniens. In: von Siebold, P. F. Fauna Japonica sive descriptio animalium, quae in itinere per Japoniam, jussu et auspiciis superiorum, qui summum in India Batava imperium tenent suscepto annis 1823-30, collegit notis observationibus et adumbrationibus illustravit Ph. Fr. de Siebold. Lugduni Batavorum (1833–1838), pp. 1–80.

    Weiss, H.B. (1927) James Petiver's Gazophylacii. Journal of the New York Entomological Society, 35(4), 411–414.

    Wermuth, H. & Mertens, R. (1961) Schildkröten, Krokodile, Brückenechsen. Gustav Fischer, Jena, XXIX, 422 pp.

    Wermuth, H. & Mertens, R. (1977) Liste der rezenten Amphibien und Reptilien. Testudines, Crocodylia, Rhynchocephalia. Das Tierreich, 100, i–xxvii, 1–174 pp.

    Werneburg, I. & Sánchez-Villagra, M. (2009) Timing of organogenesis support basal position of turtles in the amniote tree of life. BMC Evolutionary Biology, 9(1), 82.

    Williams, E.E. (1950)* Variation and selection in the cervical central articulations of living turtles. Bulletin of the American Museum of Natural History, 94(9), 505–562.

    Williams, E.E. (1952)* A new fossil tortoise from Mona Island, West Indies, and a tentative arrangement of the tortoises of the world. Bulletin of the American Museum of Natural History, 99(9), 545–560.

    Williams, E.E. (1970) Bibliographic notes on Bell’s Monograph of the Testudinata and Tortoises, Terrapins and Turtles drawn from life by Sowerby and Lear. In: J. de C. Sowerby. 1872. Sowerby & Lear's Tortoises, terrapins, and turtles with a new introduction by Ernest E. Williams. Facsimile reprints in herpetology 28. Society for Study of Amphibians and Reptiles, Athens, Ohio, pp. iii–vi.

    Zhao, E.-M. & Adler, K. (1993) Herpetology of China. Society for the Study of Amphibians and Reptiles, Oxford, Ohio, 522 pp.