Skip to main content Skip to main navigation menu Skip to site footer
Type: Articles
Published: 2010-06-09
Page range: 1–38
Abstract views: 92
PDF downloaded: 2

Taxonomic revision of the wide-front fiddler crabs of the Uca lactea group (Crustacea: Decapoda: Brachyura: Ocypodidae) in the Indo-West Pacific

Senckenberg Forschungsinstitut, Senckenberganlage 25, D-60325 Frankfurt am Main, Germany
Senckenberg Forschungsinstitut, Senckenberganlage 25, D-60325 Frankfurt am Main, Germany
Formerly Institute of Oceanology Chinese Academy of Sciences, 7 Nanhai Road, Qingdao 266071, China
Crustacea Taxonomy Ocypodidae fiddler crabs Uca Austruca lactea species-group

Abstract

The Indo-West Pacific broad-front fiddler crabs, formerly attributed to the "lactea species-group" are revised. The subgenus Uca (Austruca) Bott, 1973, is here revived for accommodating the informal "lactea species-group". Uca (Austruca) presently covers 7 species, partly with a restricted regional distribution, of which one (Uca cryptica sp. nov. from Indonesia) is new to science.

References

  1. Adiwiryono S., Sukristiyono & Toro V. (1984) The occurrence of crustaceans in the Tanjung Bungin mangrove forest, south Sumatera, Indonesia, pp. 241–257. In: Soepadmo, E., Rao, A.N. & Macintosh, D.J. [Eds.]. Proceedings of the Asian Symposium on Mangrove Environment Research and Management, Kuala Lumpur 25-29 August 1980. University of Malaya & UNESCO, Kuala Lumpur, 828 pp.

    Alcock A. (1892) On the habits of Gelasimus annulipes. Annals and Magazine of Natural History, 6 (10), 415–416.

    Alcock A. (1900) Material for a carcinological fauna of India. No. 6: The Brachyura Catometopa, or Grapsoidea. Journal of the Asiatic Society of Bengal, 69 (2), 279–456.

    Alcock A. & Anderson A.R.S. (1894) List of the shore and shallow water Brachyura collected during the season 1893-94. Natural history notes from H.M. Indian Marine survey steamer “Investigator”. Ser. ii, No. 17. Journal of the Asiatic Society of Bengal, 63, (4), 197–209.

    Alen B.J. & Levinton J.S. (2007) Costs of bearing a sexually selected ornamental weapon in a fiddler crab. Functional Ecology, 21, 154–161.

    Al-Ghais S.M. & Cooper, R.T. (1996) Brachyura (Grapsidae, Ocypodidae, Portunidae, Xanthidae and Leucosiidae) of Umm Al Quwain mangal, United Arab Emirates. Tropical Zoology, 9, 409–430.

    Apel M. (2001) Taxonomie und Zoogeographie der Brachyura, Paguridea und Porcellanidae (Crustacea: Decapoda) des Persisch-Arabischen Golfes. Unpublished Ph.D. Thesis, Johann Wolfgang Goethe-Universität, Frankfurt am Main, 268 pp.

    Apel M. & Türkay M. (1999) Taxonomic composition, distribution and zoogeographic relationships of the grapsid and ocypodid crab fauna of intertidal soft bottoms in the Arabian Gulf. Estuarine Coastal and Shelf Science, 49 (suppl. A), 131–142.

    Balss H. (1922) Ostasiatische Decapoden. iv. Die Brachyuren (Canceridae). Archive Für Naturgeschichte, A (11), 94–166.

    Balss H. (1924) Decapoden des Rotes Meeres iii. Die Parthenopoiden. Cyclo- und Catometopen. Expedition S.M. Schiff ‘Pola’ in das Rote Meer, Nördliche und Südliche Hälfte, 1895/96-1897/98. Zoologische Ergebnise xxxiv. Denckschriften Akademie der Wissenschaften (Wien), 99, 1–18.

    Barnard K.H. (1950) Descriptive Catalogue of South African decapod Crustacea (Crabs and Shrimps). Annals of the South African Museum, 38, 837 pp.

    Barnes R.S.K. (2010) A remarkable case of fiddler crab, Uca spp., alpha diversity of Wallacea. Hydrobiologia, 637, 249–253.

    Beinlich B. & von Hagen H.O. (2006) Materials for a more stable subdivision of the genus Uca Leach. Zoologische Mededelingen, 80 (2), 9–32.

    Boone L. (1934) Scientific Results of the World Cruise of the Yacht ”Alva”, 1931, William K. Vanderbilt, Commanding. Crustacea: Stomatopoda and Brachyura. Bulletin of the Vanderbilt Marine Museum, 5, 210 pp. Borradaile L.A. (1907) Land crustaceans. In: Gardiner, J.S. [Ed.], Fauna and Geography of the Maldive and Laccadive Archipelagoes. Being the Account of the Work carried out on and of the Collections made by an Expedition during the years 1899 and 1900. Volume 1, Cambridge University Press, Cambridge, pp. 65–100.

    Borradaile L.A. (1910) On the Land and Amphibious Decapoda of Aldabra. Transactions of the Linnean Society of London, series 2, Zoology, 13, 405–409.

    Bott R. (1973) Die Verwandtschaftliche Beziehungen de Uca-Arten (Decapoda: Ocypodidae). Senckenbergiana Biologica, 54, 315–325.

    Bouvier E.L. (1915) Décapodes marcheurs (Reptantia) et stomatopodes recueillis à l’île Maurice par M. Paul Carie. Bulletin Scientifique de la France et de la Belgique, 48 (3), 178–318.

    Boyce D.R. (1924) The calling crabs of Durban Bay, Uca annulipes (Milne-Edwards). South African Journal of Natural History, 4, 250–252.

    Barnwell F.H. (1980) Taxonomic differentiation within the lactea-complex of fiddler crabs in the Sunda shelf region. American Zoologist, 20, 957 [abstract].

    Brandis D., Storch V. & Türkay M. (1999) Morphology and function of the copulatory system in freshwater crabs of the genus Potamon. Journal of Morphology, 239(2), 157–166.

    Buitendijk A.M. (1947) Crustacea Anornura and Brachyura. Zoological notes frorn Port Dickson, iii. Zoologische Mededeelingen, 28, 280–284.

    Cano G. (1889) Crostacei brachiuri ed anornuri raccolti nel viaggio della "Vettor Pisani" intorno al globo. Bolettino della Società dei Naturalisti in Napoli, 3, 79–106, 169–269.

    Chace F.A. (1942) Decapod Crustacea. Scientific results of a fourth expedition to forested areas in Eastern Africa. iii. Bulletin of the Museum of Comparative Zoology, Harvard College, 91, 185–233.

    Chhapgar B.F. (1957) On the marine crabs (Decapoda, Brachyura) of Bombay State. Part ii. Journal of the Bombay Natural History Society, 54, 503–549.

    Claude M. (1964) Check list of the Crustacea Brachyura (Crabs) recorded from Mauritius. The Mauritius Institute Bulletin, 5 (1), pp. 1–60.

    Clayton D.A (1986) Ecology of mudflats with particular reference to those of the northern Arabian Gulf. In: Halwagy, R., Clayton, D. & Behbehani, M. (Eds.), Marine Environment and Pollution. Proceedings of the 1st Arabian Gulf Conference on Environment and Pollution, Kuwait, 79 February 1982. University of Kuwait, pp. 83–96.

    Crane J. (1975) Fiddler Crabs of the World. Ocypodidae: Genus Uca. Princeton University Press, Princeton, New Jersey, i–xxiii + 736 pp.

    Crosnier A. (1965) Crustacés Décapodes, Grapsidae et Ocypodidae. Faune de Madagascar, 18, pp 1–143.

    Dai Ai-Yun & Yang S. (1991) Crabs of the China Seas. China Ocean Press Beijing; Springer Berlin, Heidelberg, New York, Tokyo, 682 pp., pls. 1–74 (English edition).

    Dana J.D. (1852) Crustacea, Part i. In: United States Exploring Expedition during the years 1838 to 1842, under command of Charles Wilkes, U.S.N.C. Sherman, Philadelphia, 13 (1), i–viii + 685 pp.

    Davie P.J.F. (2002) Crustacea: Malacostraca: Eucarida (Part 2): Decapoda, Anomura, Brachyura. Zoological Catalogue of Australia. In: Wells, A. & W.W.K. Houston (Eds.), Vol. 19.3B. Melbourne: CSIRO Publishing. 641 pp.

    Dayson M. (2008) Factors affecting mating tactics in the fiddler crab, Uca vocans hesperiae. Ethology, 114, 75–84.

    Doflein F. (1899) Amerikanische Dekapoden der k. bayerischen Staatssammlungen. Sitzungsber. Königlich Bayerischen Akademie der Wissenschaften, 29, 177–95.

    Felgenhauer B.E. & Abele L.G. (1990) Evolution of the foregut in lower Decapoda. In: Felgenhauer, B.E., Waltung, L. & Thistle, A.B. [Eds.], Functional morphology of feeding and grooming in Crustacea. Crustacean Issues, 6, pp. 205–219.

    Forest J. & Guinot D. (1961) Crustacés décapodes brachyoures de Tahiti et des Tuamotu. In: Expédition Française sur les Récifs Coralliens de la Nouvelle-Calédonie. Volume préliminaire. Fondation Singer-Polignac, Paris, xi + 195 pp., pls. 1–18.

    Frith D.W. (1977) A preliminary list of macrofauna from a mangrove forest and adjacent biotops at Suran Island, western Peninsula Thailand. Phuket Marine Biological Center Research Bulletin, 17, pp. 1–16.

    Gee N. G. (1925) Tentative list of Chinese decapod Crustacea including those represented in the collection of the U.S.N.M. with localities at which collected. Lingnaam Agricultural Review, 3 (2), 156–66.

    George R.W. & Jones D.S. (1982) A revision of the fiddler crabs of Australia (Ocypodidae: Uca). Record of the Western Australian Museum, Suppl. 14, 99 pp.

    Ghory F.S. & Siddiqui F.A. (2006) A comparative study of the first zoeal stage of Uca urvillei and Uca annulipes (Crustacea. Brachyura. Ocypodidae) reared in the laboratory. Turkish Journal of Zoology, 30(2), 161–166.

    Gordon I. (1931) Brachyura from the coasts of China. Journal of the Linnean Society of London, Zoology, 37(254), 525–558.

    Gordon I. (1934) Résultats scientifiques du voyage aux Indes Orientales Néerlandaises de LL. AA. RR. le Prince et la Princesse Léopold de Belgique. Crustacea Brachyura. Memoires du Musée Royal D’Histoire Naturelle de Belgique, Hors Série 3 (15), 3–78.

    Guinot D. (1979) Données nouvelles sur la morphologie, la phylogenèse et la taxonomie des Crustacés Décapodes Brachyoures. Mémoires du Muséum national d'Histoire naturelle, N.S. (A, Zool.) 112, 1–354, figs. 1–70, pls. 1–27, tabs. 1–5.

    Haan W. De (1833–1850) Crustacea. In: Siebold, P.F. de (Ed.): Fauna Japonica. Lugduni-Batavorum, Bd. 4, xvii + xxxi + 244 pp., pls. 1–70.

    Hagen H.O. von & Jones D.S. (1986) The fiddler crabs (Ocypodidae: Uca) of Darwin, Northern Territory, Australia. The Beagle, Records of the Northern Territory Museum of Arts and Science, 6 (1), 55–68.

    Heller C. (1865) Crustaceen. In: Reise der österreichischen Fregatte Novara“ um die Erde, in den Jahren 1857, 1858, 1859 unter den Befehlen des Commodore B. Von Wüllerstrof-Urbair. Zoologischer Theil, 2 (3), 280 pp, pls. 1–25.

    Henderson J. R. (1893) A contribution to Indian carcinology. Transactions of the Linnean Society London, 2 (10), 325–458, pls. 36–40.

    Hilgendorf F. (1869) Crustaceen. In: Baron Carl Claus Von Der Decken’s Reisen in Ost-Afrika. Dritter Band: Wissenschaftliche Ergebnisse. Erste Abtheilung: Säugethiere, Vögel, Amphibien, Crustaceen, Mollusken und Echiodermen. C.F. Winter’sche Verlagshandlung, Leipzig & Heidelberg, 3 (1), 67–116, pls. 1–6.

    Hilgendorf F. (1879) Die von Herrn W. Peters in Mocambique gesammelten Crustaceen, bearbeitet von Hrn. Dr. F. Hilgendorf. Monatsberichte der königlich preussischen Akademie der Wissenschaften zu Berlin, 1878 (1879), 782–850, pls. 1–4.

    Hoffmann C.K. (1874) Crustacés et Echinodermes de Madagascar et de l’Ile de la Réunion. In: Pollen, F.P.L. & Van Dam, D.C., Recherches sur la faune de Madagascar et ses dépendances, 5 (2), 1–58, pls. i–x.

    Hogarth P.J. (1989) The Marine Crustacea of Dhofar, Southern Oman. Journal of Oman Studies, 10, 99–124.

    Hornby R. (1997) A survey of the habitats, invertebrate fauna and environmental sensitivity of the mainland coast of the UAE, with information on status and distribution of Crustaceans. TRIBULUS, Bulletin of the Emirates Natural History Group, 7 (2), 11–17.

    How M.J., Hemmi J.M., Zeil J. & Peters R. (2008) Claw waving display changes with receiver distance in fiddler crabs, Uca perplexa. Animal Behaviour, 75, 1015–1022.

    Hywel-Davies A. (1994) A Quantitative Analysis of the Horizontal and Vertical Zonation of Brachyura and Mollusca Asociated with the Qurm Mangal, Muscat, Sultanate of Oman. MasterTthesis, School of Ocean Sciences, University of Wales, Bangor, 61 pp.

    Iguchi R., Tajima M. & Wada K. (1997) Distribution of fiddler crabs, Uca arcuata and Uca lactea in the Yoshino River Estuary and vicinities (Tokushima Prefecture). Bulletin of the Tokushima Prefectural Museum, 7, 69–79.

    Ismail N.S. & Ahmed M.A.E. (1993) Macrobenthic invertebrates of mangrove, Avicennia marina (Forskal), and of intertidal flats of Khor Kalba, U.A.E., Gulf of Oman. In: Lieth, H. & Al Masoom, A. (Eds.), Towards The Rational Use Of High Salinity Tolerant Plants, Vol. 1. Tasks for Vegetation Sciences, 27 (1), 155–161.

    Jones D.A. (1986) A Field Guide to the Sea Shores of Kuwait and the Arabian Gulf. University of Kuwait Blandford Press, Poole, Kuwait (1986), 192 pp.

    Jones D.S. & Morton B. (1994) The fiddler crabs (Ocypodidae: Uca) of Hong Kong. Asian Marine Bailogy, 11, 9–40.

    Kamita T. (1935) On the Brachyura of the west Korean waters (Yellow Sea). Zoological Magazine (Tokyo), 47, 232–245.

    Kellogg C.R. (1928) Crustacea of Fukien Province. Lingnaan Science Journal, 5 (4), 351–356.

    Kemp S. (1915) Fauna of the Chilka Lake No. 3. Crustacea Decapoda. Memoirs of the Indian Museum, 5, 199–325.

    Kemp S. (1918) Zoological results of a tour in the Far East. Crustacea Decapoda and Stomatopoda. Memoirs of the Asiatic Society of Bengal, 6, 218–297.

    Khan A.M. (1978) Development of asymmetry in the claws of the fiddler crab Uca lactea. Pakistan Journal of Zoology, 10(1), 49–54.

    Kim W. & Kim S.H. (1995) The brachyuran crabs (Crustacea: Decapoda) of Chindo Island, Korea. Korean Journal of Systematic Zoology, 11(4), 497–508.

    Kingsley J.S. (1880) Carcinological notes. Proceedings of the Academy of Natural Sciences of Philadelphia. For 1879: 34-37; ii. Revision of the Gelasimi, 135–152.

    Kossmann R. (1877) Malacostraca (1. Teil: Brachyura). Zoologische Ergebnisse einer im Auftrage der Königlichen Academie der Wissenschaften u Berlin ausgeführten Reise in die Küstengebiete des Roten Meeres, 3 (1), 66 pp, pls 1–3.

    Krauss F. (1843) Die Südafrikanischen Crustaceen. Eine Zusammenstellung aller bekannten, Malacostraca. Bemerkungen über deren Lebensweise und geographische Verbreitung, nebst Beschreibung und Abbildung mehrerer neuen Arten. Schweizerbart’sche Verlagsbuchhandlung, Stuttgart, 68 pp.

    Kwok W.P.W. & Tang W.-S. (2006) Fiddler crabs in Hong Kong—an overview. Hong Kong Biodiversity, 12, 1–7.

    Lailvaux S.P., Reaney L.T. & Backwell P.R.Y. (2008) Dishonest signalling of fighting ability and multiple performance traits in the fiddler crab Uca mjoebergi. Functional Ecology, 23 (2), 359–366.

    Lanchester W.F. (1900a) On a collection of crustaceans made at Singapore and Malacca. Part 1. Crustacea Brachyura. Proceedings of the Zoological Society of London, 3, 719–770.

    Lanchester W.F. (1900b) On some malacostracous crustaceans from Malaysia in the collection of the Sarawak Museum. Annals and Magazine of Natural History, (7) 6, 249–264.

    Lanchester W.F. (1901) On the Crustacea collected during the “Skeat” Expedition to the Malay Peninsula, together with a note on the genus Actaeopsis. Part i. Brachyura, Stomatopoda, and Macrura. Proceedings of the General Meetings for Scientific Business of the Zoological Society of London, 1901 (2), 534–574.

    Laurie R.D. (1915) Reports on the marine biology of the Sudanese Red Sea, XXI. On the Brachyura. Journal of the Linnean Society of London, Zoology, 31, 407–475, figs. 1–5, pls. 42–45.

    Lee P.S. & Lim, S.S.L. (2003) Distributional patterns and species density of a fiddler crab community (Uca annulipes and U. vocans) at Telok Assam, Bako National Park, Sarawak. Sarawak Museum Journal, 58(79), 183–198.

    Lenz H. (1905) Ostafrikanische Dekapoden und Stomatopoden. Gesammelt von Herrn Prof. Dr. A. Voeltzkow. In: Voeltzkow, A.: Wissenschaftliche Ergebnisse der Reisen in Madagaskar und Ostafrika in den Jahren 1889-95. Abhandlungen der Senckenbergischen naturforschenden Gesellschaft, 27 (4), 341–392.

    Lenz H. (1910) Crustaceen von Madagaskar, Östafrika und Ceylon. In: Voeltzkow, A. 1910. Reise in Östafrika. Stutgart, vol. 2, pp. 539–576.

    Lenz H. & Richters F. (1881) Beitrag zur Crustaceenfauna von Madagascar. Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft, 12, 421–428.

    Lewinsohn C. (1977) Die Ocypodidae des Roten Meeres (Crustacea, Decapoda, Brachyura). Zoologische Verhandelingen, 152, 45–84.

    Lim S.S.L. (2004) A comparative study of some mouthpart adaptations of Uca annulipes (H. Milne Edwards, 1837) and U. vocans (Linnaeus, 1758) (Brachyura, Ocypodidae) in relation to their habitats. Crustaceana, 77(10), 1245–1251.

    Lim S.S.L., Lee P.S. & Diong C.H. (2005) Influence of biotope characteristics on the distribution of Uca annulipes (H. Milne Edwards, 1837) and U. vocans (Linnaeus, 1758) (Crustacea, Brachyura, Ocypodidae) on Pulau Hantu Besar, Singapore. Raffles Bulletin of Zoology, 53(1), 111–114.

    Lin C.-C. (1949) A catalogue of brachyurous Crustacea of Taiwan. Quarterly Journal of the Taiwan Museum, 2, 10–33.

    Litulo C. (2004) Fecundity of the pantropical fiddler crab Uca annulipes (H. Milne Edwards, 1837) (Brachyura. Ocypodidae) at Costa do Sol mangrove, Maputo Bay, southern Mozambique. Western Indian Ocean Journal of Marine Science, 3(1), 87–91.

    Litulo C. (2005) Population biology of the fiddler crab Uca annulipes (Brachyura: Ocypodidae) in a tropical East African mangrove (Mozambique). Estuarine, Coastal and Shelf Science, 62, 283–290.

    Maccagno T. (1928) Crostacei decapodi. Le specie del genre Uca Leach conservate nel Regio Museo Zoologico di Torino. Bolletino dei Musei di Zoologia e di Anatomia Comparata della R. Università di Torino, 41 (11), 1–52.

    Macia A., Quincardete, I. & Paula, J. (2001) A comparison of alternative methods for estimating population density of the fiddler crab Uca annulipes at Saco Mangrove, Inhaca Island (Mozambique). Hydrobiologia, 449, 213–219.

    Man J.G. De (1880) On some species of Gelasimus and Macrophthalmus. Notes from the Leyden Museum, 2, 67–72.

    Man J.G. De (1887–1888a) Report on the podophthalmous Crustacea of the Mergui Archipelago, collected for the Trustees of the Indian Museum, Calcutta, by Dr. John Aanderson, F.R.S., Superintendent of the Museum. Journal of the Linnean Society of London, Zoology, 22: 1–64, pls. 1–3 (pt. 1, Nov. 1887), 65–128, pls. 4–8 (pt. 2, Dec. 1887), 129–176, pls. 9–12 (pt. 3, Jan. 1888), 177–240, pls. 13–15 (pt. 4, Apr. 1888), 241–312, pls. 16–19 (pt. 5, Jul. 1888).

    Man J.G. De (1887–1888b) Bericht über die im indischen Archipel von Dr. J. Brock gesammelten Decapoden und Stomatopoden. Archiv für Naturgeschichte, 53 (1): 215–288, pls. 7–10 (1887), 289–600, pls. 11–22 (1888).

    Man J.G. De (1891) Carcinological studies in the Leyden Museum, No 5. Notes from the Leyden Museum, 13, pp. 1–61.

    Man J.G. De (1895) Bericht über die von Herrn Schiffskapitän Storm zu Atjeh, an den westlichen Küsten von Malakka, Borneo und Celebes sowie in der Java-See gesammelten Decapoden und Stomatopoden. Zoologische Jahrbücher, Abtheilung für Systematik, Geographie und Biologie der Thiere, 8 (4), 485–609.

    Man J.G. De (1902) Die von Herrn Prof. Kükenthal im Indischen Archipel gesammelten Dekapoden und Stomatopoden. (Ergebnisse einer zoologischen Forschungsreise in den Molukken und Borneo). Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft, 25 (3), 467–929.

    Miers E.J. (1879) On a collection of Crustacea made by Capt. H.C. St. John, R.N., in the Korean and Japanese seas. Part i. Podophthalmia. Proceedings of the Zoological Society of London, 1879, 17–61, 3 pls.

    Miers E.J. (1880) On a collection of Crustacea from the Malaysian region. Part iii. Crustacea Anomura and Macrura (except Penaeidea). The Annals and Magazine of Natural History, series 5, 370–384.

    Miers E.J. (1886) Report on the Brachyura collected by H.M.S. Challenger during the years 1873–1876. In: Murray, J. (ed.) Zoology. Report on the Scientific Results of the Voyage of H.M.S. Challenger During the Years 1873–76, vol. 17, 362 pp.

    Milne Edwards H. (1837) Histoire naturelle des Crustacés, comprenant l’Anatomie, la Physiologie et la classification de ces animaux. ii. Paris, 1837, 531 pp.

    Milne Edwards H. (1852) Observation sur les affinités zoologiques et la classification naturelle des crustacés. Annales des Sciences Naturelles (Zoologie), (3) 18, 109–166.

    Miyake S. (1936) A list of known species of Brachyura from Ishigakishima. Annotationes Zoologicæ Japonenses, 15(4), 506–513.

    Miyake S. (1938) Notes on decapod crustaceans collected by Prof. Teiso Esaki frorn Micronesia. Annotationes Zoologicæ Japonenses, 17, 107–112.

    Miyake S. (1939) Notes on Crustacea Brachyura collected by Professor Teiso Esaki's Micronesia Expeditions 1937–1938 together with a check list of Micronesian Brachyura. Records of Oceanographic Works in Japan, 10 (2), 168–247.

    Miyake S. (1983) Japanese crustacean decapods and stomatopods in color. 1. Macrura, Anomura and Stomatopoda. Osaka: Hoikusha. 261 pp.

    Mokhtari M., Savari A., Rezai H., Kochanian P. & Bitaab A. (2007) Population ecology of fiddler crab, Uca lactea annupiles (Decapoda: Ocypodidae) in Sirik mangrove estuary, Iran. Estuarine, Coastal and Shelf Science, 76, 273–281.

    Morgan G.J. (1990) A collection of Thalassinidea, Anomura and Brachyura (Crustacea: Decapoda) from the Kimberley Region of northwestern Australia. Zoologische Verhandelingen, Leiden, 265, 90 pp, figs. 1–6.

    Murai M., Goshima S. & Henmi Y. (1987) Analysis of the mating system of the fiddler crab, Uca lactea. Animal Behaviour, 35(5), 1334–1342.

    Musgrave A. (1929) Life in a mangrove swamp. Australian Museum Magazine, 3 (10), 341–347.

    Naiyanetr P. (1980) Crustacean Fauna of Thailand (Decapoda and Stomatopoda). Department of Biology, Faculty of Science, Chulalongkorn University of Bangkok, Thailand, 73 pp.

    Naiyanetr P. (2007) Checklist of crustacean fauna in Thailand (Decapoda, Stomatopoda, Anostraca, Myodocopa and Isopoda). Open Biodiversity Series, 19, 197 pp.

    Ng P.K.L, Wang C.-H., Ho P.-Ho. & Shih H.-T. (2001) An annotated checklist of brachyuran crabs from Taiwan (Crustacea: Decapoda). National Taiwan Museum Special Publication Series, Number 11, 86 pp, pl. 8.

    Ng P.K.L. & Davie P.J.F. (2002) A checklist of the Bbrchyuran crabs of Phuket and Western Thailand. Phuket Marine Biological Center Special Publication, 23(2), 369–384.

    Ng P.K.L., Guinot D. & Davie P.J.F. (2008) Systema brachyurorum: Part I. An annotated checklist of extant brachyuran crabs of the world. Raffles Bulletin of Zoology, Suppl. 17, 286 pp.

    Nobili G. (1899a) Contribuzioni alla conscenza della Fauna carcinologia della Papuasia, delle Malucche e dell’Austrialia. Annali del Museo civico di Storia naturale Genova, 20, 230–282.

    Nobili G. (1899b) Decapodi e Stomatopodi Indo-Malesi. Annali del Museo civico di Storia naturale Genova, 40, 473–523.

    Nobili G. (1901) Note intorno ad una collezione di crostacei di Sarawak (Borneo). Bolletino di Zoologia ed Anatomia comparata. Della R. Universita di Torino, 16 (397), 1–14.

    Nobili G. (1903) Crostacei di Pondichery, Mahe, Bombay etc. Bolletino di Zoologia ed Anatomia comparata, della R. Universita di Torino, 18 (452), 24 pp.

    Nobili G. (1906a) Crustacés décapodes et stomatopodes. In: Mission J. Bonnier et Ch. Perèz (Golfe Persique, 1901). Bulletin Scientifique de la France et de la Belgique, 40, 13–159, pls. 2–7.

    Nobili G. (1906b) Faune carcinologique de la Mer Rouge. Décapodes et Stomatopodes. Annales des Sciences naturelles, (Zoologie), (9) 4, 1–347, figs. 1–12, pls. 1–11.

    Ono Y. (1965) On the ecological distribution of ocypodoid crabs in the estuary. Memoirs of the Faculty of Science of Kyushu University, Series E, 4 (1), pp. 1–60.

    Ortmann A.E. (1894a) Die Decapoden-Krebse des Strassburger Museums. viii. Theil. Abteilung: Brachyura (Brachyura genuina Boas) iii. Unterabteilung: Cancroidea, 2. Section: Cancrinea, 2. Gruppe: Catometopa. Zoologische Jahrbücher, Abtheilung für Systematik, Geographie und Biologie der Thiere, 7, 683–772, pl. 23.

    Ortmann A.E. (1894b) Crustaceen. In: Semon (ed.) Zoologische Forschungsreisen in Australien und dem malayischen Archipel. Denkschriften der Medicinisch-Naturwissenschaftlichen Geselschaft zu Jena, 8, 3–80.

    Ortmann A.E. (1897) Carcinologische Studien. Zoologische Jahrbücher. Abteilung für Systematik, Geographie und Biologie der Thiere, 10(3), 258–372.

    Panikkar N.K. & Aiyar R.G. (1937) The brackish-water fauna of Madras. Proceeding of Indian Academic of Science, 6 (B5), 284–337.

    Pesta O. (1911) Crustacea. Botanische und Zoologische Ergebnisse einer Wissenschaftlichen Forschunsreise nach den Samoa Inseln, Dem Neuguinea-Arcihpel und den Salomonsinseln, von März bis Dezember 1905. Kaiserlich-Königlichen Hof und Staatsdruckerei, Wien, 30 pp, figs. 5, pl. 3.

    Pillay K.K. & Nair N.B. (1971a) The annual reproductive cycles of Uca annulipes, Portunus pelagicus and Metapenaeus affinis (Decapoda. Crustacea) from the south-west coast of India. Marine Biology, Berlin, 11(2), 152–166.

    Pillay K.K. & Nair N.B. (1971b) The reproductive cycle of three decapod crustaceans from the south-west coast of India. Current Science, 40(7), 161–162.

    Pillay K.K. & Nair N.B. (1973) Observations on the breeding biology of some crabs from the southwest coast of India. Journal of the Marine. Biological Association of India, 15(2), 754–770.

    Pretzmann G. (1971) Ergebnisse einiger Sammelreisen nach Vorderasien. 2. Teil: Marine Brachyura. Annalen des Naturhistorischen Museums Wien, 75, 477–487.

    Rajendran A. (1972) Distribution of Uca annulipes (Latreille) in the intertidal mudflats of the Vellar estuary. Proceedings of the Indian National Science Academy, 38(5–6), 378–379.

    Rathbun M.J. (1910) The Danish Expedition to Siam 1899–1900. v. Brachyura. Kongelige Danske Videnskabernes Selskab, Naturvidenskabelige Matematiske Afhandlinger, 5(4), 301–367.

    Rathbun M.G. (1924) Results of Dr. E. Mjoberg’s Swedish scientific expeditions to Australia 1910–1913. 37. Brachyura, Albuneidae and Porcellanidae. Arkiv för Zoologi, 16 (23), 1–33.

    Richters F. (1880) Decapoda. In: Möbius, K. (ed.) Beiträge zur Meeresfauna der Insel Mauritius und der Seychellen bearbeitet von K. Möbius, F. Richters und E. von Martens nach Sammlungen, angelegt auf einer Reise nach Mauritius von K. Möbius. Berlin: Verlag der Gutmannischen Buchhandlung, 139–178.

    Roberts C.M., McLean C.J., Veron J.E.N., Hawkins J.P., Allen G.R., McAllister D.E., Mittermeier C.G., Schueler F.W., Spalding M., Wells F., Vynne C. & Werner, T.B. (2002) Marine Biodiversity Hotspots and Conservation Priorities for Tropical Reefs. Science, 295, 1280–1284.

    Rosenberg M.S. (2001) The systematics and taxonomy of fiddler crabs: a phylogeny of the genus Uca. Journal of Crustacean Biology, 21, 839–869.

    Roux J. (1917) Crustacés (Expedition de 1903). Nova Guinea, 5, 589–621.

    Sakai T. (1934) Brachyura from the coast of Kyushu, Japan. Science Reports of the Tokyo Bunrika Daigaku Section, 1 (25), 281–330.

    Sakai T. (1936) Report on the Brachyura collected by Mr. F. Hiro at Palao (Pelew) Islands. Science Reports of the Tokyo Bunrika Daigaku, (B) 2, 155–177.

    Sakai, T. (1939) Studies on the Crabs of Japan. Brachygnatha, Brachyrhyncha, IV. Yokendo, Tokyo, 365–741 + 1–23 [index] pp., pls. 42–111.

    Sakai T. (1940) Biogeographic review on the distribution of crabs in Japanese waters. Records of Oceanographic Works in Japan, 11 (1), 27–63.

    Sakai, T. (1976) Crabs of Japan and the Adjacent Seas. Kodansha, Tokyo, vol. 1 [English text], xxix + 773 pp., figs. 1–379, maps 1–3; vol. 2 [Japanese text], 461 pp., figs. 1, 2; vol. 3 [plates], 16 pp. + pls. 1–251.

    Sakai K., Türkay M. & Yang S.L. (2006) Revision of the Helice/Chasmagnathus complex (Crustacea: Decapoda: Brachyura). Abhandlungen der Senckenbergischen Naturforschenden Geselschaft, 565, 1–76.

    Sankarankutty C. (1961) On Decapoda Brachyura from the Andaman and Nicobar Islands. 1. Families Portunidae, Ocypodidae, Grapsidae and Mictyridae. Journal of the Marine Biological Association of India, India, 3 (1, 2), 101–119.

    Schenkel E. (1902) Beitrag zur Kentnis der Dekapodenfauna von Celebes. Verhandlungen der Schweizerischen Naturforschenden Geselschaft (Basel, 13, 485–585. pls. 7–13.

    Sewell R.B. (1913) Notes on the biological work of the R.I.M.S.S. “Investigator” during survey seasons 1910–1911 and 1911-1912. Journal of the Asiatic Society of Bengal, 9, 329–390.

    Sheppard C.R.C., Price A.R.G. & Roberts C. (1992) Marine Ecology of the Arabian Region Patterns and Processes in Extremetropical Environments. Academic Press, London, 359 pp.

    Shih H.-T, Mok H.-K, Chang H.-W & Lee S.-C. (1999) Morphology of Uca formosensis Rathbun, 1921 (Crustacea: Decapoda: Ocypodidae), an endemic fiddler crab from Taiwan, with notes on its ecology. Zoological Studies, 38(2), 164–177.

    Shih H.-T, Kamrani E., Davie P.J.F & Liu M.Y. (2009) Genetic evidence for the recognition of two fiddler crabs, Uca iranica and U. albimana (Crustacea: Brachyura: Ocypodidae), from the northwestern Indian Ocean, with notes on the U. lactea species-complex. Hydrobiologia, 635, 373–382.

    Shinn E.A. (1976) Coral reef recovery in Florida and the Persian Gulf. Environmental Geology, 1, 241–254.

    Skov M.W., Vannini M., Shunula J.P., Hartnoll R.G. & Cannicci S. (2002) Quantifying the density of mangrove crabs. Ocypodidae and Grapsidae. Marine Biology (Berlin), 141(4), 725–732.

    Stebbing T.R.R. (1917) The Malacostraca of Natal. Annals of the Durban Museum, 2, 1–33.

    Stephensen, K. (1946) The Brachyura of the Iranian Gulf. With an appendix: The male pleopoda of the Brachyura. In: Jessen, K. & Spärck, R. (Eds.), Danish Scientific Investigations in Iran, part 4 [1945], 1–237.

    Stimpson W. (1858) Prodromus descriptionis animalium evertebratorum, quae in Expeditione ad Oceanum Pacificum Septentrionalem, a Republica Federata missa, Cadwaladaro Ringgold et Johanne Rodgers Ducibus, observavit et descripsit. Pars V. Crustacea Ocypodoidea. Proceedings of the Academy of Natural Sciences of Philadelphia, 10, 93–110.

    Suvatti C. (1938) A check-list of aquatic fauna in Siam (excluding fishes). Bureau of Fisheries (Bangkok), pp. 116.

    Syomons C.T. (1920) Notes on certain shore crabs. Spolia Zeylanica, 306–313.

    Tanaka H., Shibagaki K., Ikezawa H., Kanesawa A. & Wada K. (2004) Occurrence of two species of fiddler crab (Decapoda, Ocypodidae) on the tidal flat of the Aono River, Izu Peninsula, Shizuoka Prefecture, Japan. Japanese Journal of Benthology, 59, 8–12.

    Tesch J.J. (1918) The Decapoda Brachyura of the Siboga Expedition. II. Goneplacidae and Pinnotheridae. Siboga Expeditie Monographie, 39c1 (84), 149–295, pls. 7–18.

    Tirmizi N.M. & Ghani N. (1996) Marine Fauna of Pakistan: 5: Crustacea: Brachyura, Brachyrhyncha, Part 1: Xanthidae, Goneplacidae, Pinnotheridae, Ocypodidae, Grapsidae. Center of Excellence, University of Karachi, 1–188.

    Türkay M. (1975) Statement: Die Bedeutung des Gonopodenaufbaus für die Aufklärung von Verwandtschaftsverhältnissen bei dekapoden Crustaceen. Aufsetzen und Reden senckenbergische naturforschung Geselschaft (Frankfurt a. M.), 27, 114–115.

    Tweedie M.W.F. (1937) On the crabs of the family Ocypodidae in the collection of the Raffles Museum. Bulletin of the Raffles Museum, 13, 140–170.

    Tweedie M.W.F. (1950) Grapsoid crabs from Labuan and Sarawak. Sarawak Museum Journal, 5 (2), 356–367.

    Vannini M. & Valmori P. (1981) Researches on the coast of Somalia. The shore and the dune of Sar Uanle. 31. Ocypodidae and Gecarcinidae. Monitore Zoologico Italiano, n.s., suppl. 14, 199–226.

    Vatova,A. (1943) Decapoda della Somalia. Thalassia, 6 (2), 1–37.

    White A. (1847) List of the specimens of Crustacea in the Collection of the British Museum. London, I–VIII + 143 pp.

    Yamaguchi T. (1970) Ecology of a fiddler crab (Uca lactea). Calanus, 2, 5–30.

    Yamaguchi T. (1973) Asymmetry and dimorphism of chelipeds in the fiddler crab, Uca lactea De Haan. Zoological Magazine (Tokyo), 82(3), 154–158.

    Yamaguchi T. (1977) Studies on the handedness of the fiddler crab, Uca lactea. Biological Bulletin (Woods Hole), 152(3), 424–436.

    Yamaguchi T. (1978) Studies on the decision of handedness in the fiddler crab Uca lactea. Calanus, 6, 29–51.

    Yamaguchi T. (1983) Representative shore animals in the Amakusa Islands (1). The fiddler crab, Uca lactea lactea (de Haan). Calanus, 8, 25–32.

    Yamaguchi T. (2001a) The breeding period of the fiddler crab, Uca lactea (Decapoda, Brachyura, Ocypodidae) in Japan. Crustaceana, 74(3), 285–293.

    Yamaguchi T. (2001b) The mating system of the fiddler crab, Uca lactea (Decapoda, Brachyura, Ocypodidae). Crustaceana, 74(4), 389–399.

    Yamaguchi T. & Ogata R. (2000) Studies of the first and second maxillipeds of the fiddler crab, Uca lactea. Crustacean Research, 29, 133–142.

    Yang S.L. (1986) The diagnostic value of gasctric mill’s construction as taxonomic charcter on the classification of crabs (Brachyura: Crustacea). Acta Zootaxonomica Sinica, 11(2), 151–159.

    Zehnter L. (1894) Crustacés de 1’Archipel malais-voyage de M.M. Bedot en Ch. Pichet dans 1’Archipel malais. Revue suisse de Zoologie, 2 (1), 135–214.

    Zeil J., Hemmi J.M. & Bachwell P.R. (2006) Fiddler crabs. Current Biology, 16, 40–41.