Abstract
Research on the taxonomy of European amphibians and reptiles has increased noticeably over the last few decades, indicating the need for recognition of new species and the cancellation of others. This paper provides a critical review of recent changes and draws up a tentative species list.References
Akın, Ç., Bilgin, M., Bilgin, C.C. (2010) Discordance between ventral colour and mtDNA haplotype in the water frog Rana (ridibunda) caralitana, 1988 Arıkan. Amphibia-Reptilia, 31, 9–20.
Albert, E.M. & Fernández, A. (2009) Evidence of cryptic speciation in a fossorial reptile: description of a new species of Blanus (Squamata: Amphisbaenia: Blanidae) from the Iberian Peninsula. Zootaxa, 2234, 56–68.
Albert, E.M., Zardoya, R. & García-París, M. (2007) Phylogeographical and speciation patterns in subterranean worm lizards of the genus Blanus (Amphisbaenia: Blanidae). Molecular Ecology, 16(7), 1519–1531.
Alström, P., Rasmussen, P.C., Olsson, U. & Sundberg, P. (2008) Species delimitation based on multiple criteria: the Spotted Bush Warbler Bradypterus thoracicus complex (Aves: Megaluridae). Zoological Journal of the Linnean Society, 154, 291–307.
Arnold, E.N., Arribas, O. & Carranza, S. (2007) Systematics of the Palaearctic and Oriental lizard tribe Lacertini (Squamata: Lacertidae: Lacertinae) with descriptions of eight new genera. Zootaxa, 1430, 1–86.
Arnold, E.N., Burton, J.A. & Ovenden, D.W. (1978) A field guide to the reptiles and amphibians of Britain and Europe. Collins, London, pp. 272.
Arnold, E.N. & Ovenden, D.W. (2002) A Field Guide to the Reptiles and Amphibians of Britain & Europe. Harper Collins Publishers, London, pp. 288.
Arntzen, J.W., Espregueira Themudo, G. & Wielstra, B. (2007) The phylogeny of crested newts (Triturus cristatus superspecies): nuclear and mitochondrial genetic characters suggest a hard polytomy, in line with the paleogeography of the centre of origin. Contributions to Zoology, 76(4), 261–278.
Arribas, O.J. (2009) Morphological variability of the Cantabro-Pyrenean populations of Zootoca vivipara (JACQUIN, 1787) with description of a new subspecies. Herpetozoa, 21(3/4), 123–146.
Babik, W. , Branicki, W., Crnobrnja-Isailovic, J., Cogalniceanu, D. , Sas, I., Olgun, K., Poyarkov, N.A., Garcia-Paris, M. & Arntzen, J.W. (2005) Phylogeography of two European newt species—discordance between mtDNA and morphology. Molecular Ecology, 14, 2475–2491.
Baha el Din, S. (2006) A Guide to the Reptiles and Amphibians of Egypt. The American University in Cairo Press, Cairo, Egypt, 320 pp.Balletto, E., Bologna, M.A.B. & Giacoma, C. (2007) Bufo viridis Laurenti, 1768 complex. In: Lanza, B., Andreone, F., Bologna, M.A., Corti, C. & Razzetti, E. (Eds), Amphibia. Collana “Fauna d’Italia”. Vol. XLII. Calderini, Bologna, pp. 296–305.
Barbanera, F., Zuffi, M.A.L., Guerrini, M., Gentilli, A., Tofanelli, S., Fasola, M. & Dini, F. (2009) Molecular phylogeography of the asp viper Vipera aspis (Linnaeus, 1758) in Italy: evidence for introgressive hybridisation and mitochondrial DNA capture. Molecular Phylogenetics and Evolution, 52, 103–114.
Bauer, A.M., Jackman, T.R., Greenbaum, E. & Gamble, T. (2008) Phylogenetic relationships of the Italian gekkotan fauna. In: Corti, C. (Ed), Herpetologia Sardiniae. Societas Herpetologica Italica/Edizioni Belvedere, Latina, “le scienze”, 8, pp. 59–62.
Beerli, P. (1994) Genetic isolation and calibration of an average protein clock in western Palearctic water frogs of the Aegean region. PhD dissertation, University Zurich, Zurich, 92 pp.
Beerli, P., Hotz, H., Tunner, H.G., Heppich, S. & Uzzell, T. (1994) Two new water frog species from the Aegean Islands Crete and Karpathos. Notulae Naturae, 470, 1–9.
Bocage, J.V.B. (1863) Liste des mammifères et Reptiles observés en Portugal. Revue et Magasin de Zoologie, 15, 329–333.
Bonaparte, C.L. (1840) Amphibia Europaea ad systema nostrum vertebratorum ordinata. Memorie della Reale Accademia delle Scienze di Torino, 2, 385–456.
Bonato, L. & Steinfartz, S. (2005) Evolution of the melanistic colour in the Alpine salamander Salamandra atra as revealed by a new subspecies from the Venetian Prealps. Italian Journal of Zoology, 72, 253–260.
Böttger, O. (1869) Beitrag zur Kenntnis der Reptilien Spaniens und Portugals. Jahresbericht des Offenbacher Vereins für Naturkunde, 10, 50–59.
Bour, R., Cheylan, M., Crochet, P.-A., Geniez, P., Guyetant, R., Haffner, P., Ineich, I., Naulleau, G., Ohler, A.-M. & Lescure, J. (2008) Liste taxinomique actualisée des Amphibiens et Reptiles de France. Bulletin de la Société Herpétologique de France, 126, 37–43.
Bour, R. & Dubois, A. (1985) Nomenclature ordinale et familiale des Tortues (Reptilia). Studia geologica salmanticensia, vol. especial 1, Studia palaeocheloniologica, I, Salamanca, Ediciones Universidad, 77–86.
Camerano, L. (1884) Monografia degli Anfibi Anuri italiani. Memorie della Reale Accademia delle Scienze di Torino, 35, 187–284.
Canestrelli, D., Caputo, F.P., Bagnoli, C. & Nascetti, G. (2006a) Integrating genetic, demographic and ecological issues for the conservation of the Alpine newt in central Italy. Annales Zoologici Fennici, 43, 322–334.
Canestrelli, D., Cimmaruta, R. & Nascetti, G. (2007a) Phylogeography and historical demography of the Italian treefrog, Hyla intermedia, reveals multiple refugia, population expansions and secondary contacts within peninsular Italy. Molecular Ecology, 16, 4808–4821.
Canestrelli, D. & Nascetti, G. (2008) Phylogeography of the pool frog Rana (Pelophylax) lessonae in the Italian peninsula and Sicily: multiple refugia, glacial expansions and nuclear-mitochondrial discordance. Journal of Biogeography, 35, 1923–1936.
Canestrelli, D., Verardi, A. & Nascetti, G. (2007b) Genetic differentiation and history of populations of the Italian treefrog Hyla intermedia: lack of concordance between mitochondrial and nuclear markers. Genetica, 130, 241–255.
Canestrelli, D., Zangari, F. & Nascetti, G. (2006b) Genetic evidence for two distinct species within the Italian endemic Salamandrina terdigitata (Bonnaterre, 1789) (Amphibia: Urodela: Salamandridae). Herpetological Journal, 16, 221–227.
Carranza, S., Romano, A., Arnold, E.N. & Sotgiu, G. (2008) Biogeography and evolution of European cave salamanders, Hydromantes (Urodela: Plethodontidae), inferred from mtDNA sequences. Journal of Biogeography, 35, 724–738.
Carranza, S., Arnold, E.N., Geniez, P., Roca, J.L. & Mateo, J.A. (2008) Radiation, multiple dispersal and parallelism in Moroccan skinks, Chalcides and Sphenops (Squamata: Scincidae), with comments on Scincus and Scincopus and the age of the Sahara Desert. Molecular Phylogenetics and Evolution, 46, 1071–1094.
Carretero, M.A. (2008) An integrated assessment of a group with complex systematics: the Iberomaghrebian lizard genus Podarcis (Squamata, Lacertidae). Integrative Zoology, 4, 247–266.
Carretero, M.A., Ayllón, E. & Llorente, G. (Eds) (2009) Lista patron de los anfibios y reptiles de España (actualizada a enero de 2009). AHE, Barcelona, 28 pp.
Castellano, S., Cuatto, B., Rinella, R., Rosso, A. & Giacoma, C. (2002) The advertisement call of the European treefrogs (Hyla arborea): a multilevel study of variation. Ethology, 108, 75–89.
Červenka, J., Kratochvíl, L. & Frynta, D. (2008) Phylogeny and taxonomy of the Middle Eastern geckos of the genus Cyrtopodion and their selected relatives. Zootaxa, 1931, 25–36.
Crochet, P.-A. (2007) Nomenclature of European Plethodontid salamanders: Speleomantes Dubois, 1984 has precedence over Atylodes Gistel, 1868. Amphibia–Reptilia, 28(1), 170–172.
Crochet, P.-A. & Dubois, A. (2004) Recent changes in the taxonomy of European amphibians and reptiles. In: Gasc, J.P., Cabela, A., Crnobrnja-Isailovic, J., Dolmen, D., Grossenbacher, K., Haffner, P., Lescure, J., Martens, H., Martínez Rica, J.P., Maurin, H., Oliveira, M.E., Sofianidou, T.S., Veith, M. & Zuiderwijk, A. (Eds), Atlas of the amphibians and reptiles in Europe (second edition). Societas Europaea Herpetologica & Muséum National d'Histoire Naturelle, Paris, pp. 495–516.
Danflous, S., Geniez, Ph., Arnold, E.N. & Ovenden, D.W. (2004) Le guide herpéto: 199 amphibiens et reptiles d'Europe. Editions Delachaux et Niestlé, pp. 288.
de Betta, E. (1874) Fauna d’Italia. Parte Quarta. Rettili ed Anfibi. Dottor Francesco Vallardi, Tipografo-Editore, Milano, pp. 107.
Donaire Barroso, D. & Bogaerts, S. (2003) A new subspecies of Salamandra algira from northern Morocco. Podarcis, 4(3), 84–100.
Donaire Barroso, D., González de la Vega, J.P. & Barnestein, J.A.M. (2009) Aportación sobre los patrones de diseño pigmentario en Salamandra longirostris Joger & Steinfartz, 1994, y nueva nomenclatura taxonómica. Butlletí de la Societat Catalana d’Herpetologia, 18, 10–17.
Dubois, A. (1984) La nomenclature supragénérique des Amphibiens Anoures. Mémoires du Muséum national d’Histoire naturelle, 131, 1–64.
Dubois, A. (1985) Miscellanea nomenclatorica batrachologica (VII). Alytes, 4(2), 61–78.
Dubois, A. (1988) The genus in zoology: a contribution to the theory of evolutionary systematics. Mémoires du Muséum national d’Histoire naturelle, 140, 1–123.
Dubois, A. (1998) Lists of European species of amphibians and reptiles: will we soon be reaching “stability”? Amphibia–Reptilia, 19, 1–28.
Dubois, A. (2004) The higher nomenclature of recent amphibians. Alytes, 22 (1–2), 1–14.
Dubois, A. (2005) Proposals for the incorporation of nomina of higher-ranked taxa into the Code. Bulletin of zoological Nomenclature, 62, 200–209.
Dubois, A. (2008) Authors of zoological publications and nomina are signatures, not persons. Zootaxa, 1771, 63–68.
Dubois, A. & Bour, R. (in press) The nomenclatural status of the nomina of amphibians and reptiles created by Garsault (1764), with a parsimonious solution to an old nomenclatural problem regarding the genus Bufo (Amphibia, Anura) and comments on some nomina created by Laurenti (1768). Zootaxa.
Dubois, A. & Dinesh, K. P. (2007) The status of the nomen Rana (Paa) dhakuriensis Ray, 1997 (Anura, Ranidae), and comments on the Amphibia reported from the Nanda Devi Biosphere Reserve (Uttar Pradesh, India). Alytes, 25 (1–2), 75–82.
Dubois, A. & Günther, R. (1982) Klepton and Synklepton: two new evolutionary systematic cat- egories in zoology. Zoologische Jahrbücher (Systematik), 109, 290–305.
Dubois, A. & Ohler, A. (1995a) Frogs of the subgenus Pelophylax (Amphibia, Anura, genus Rana): a catalogue of available and valid scientific names, with comments on name-bearing types, complete synonymies, proposed common names, and maps showing all type localities. Zoologica Poloniae, “1994”, 39 (3–4), 139–204.
Dubois, A. & Ohler, A. (1995b) Catalogue of names of frogs of the subgenus Pelophylax (Amphibia, Anura, genus Rana): a few additions and corrections. Zoologica Poloniae, “1994”, 39 (3–4), 205–208.
Dubois, A. & Raffaëlli, J. (2009) A new ergotaxonomy of the family Salamandridae Goldfuss, 1820 (Amphibia, Urodela). Alytes, 26(1–4), 1–85.
Duda, M. (2008) First record of a natural male hybrid of Bufo (Pseudepidalea) viridis LAURENTI, 1768 and Bufo (Bufo) bufo LINNEUS, 1758 from Austria. Herpetozoa, 20(3/4), 184–186.
Engelmann, W.E., Fritsche, J., Günther, R. & Obst, F.J. (1993) Lurche und Kriechtiere Europas. Neumann Verlag GmbH, Radebeul, pp. 440.
Espregueira Themudo, G. & Arntzen, J.W. (2007) Molecular identification of marbled newts and a justification of the Triturus marmoratus and T. pygmaeus species status. Herpetological Journal, 17, 24–30.
Espregueira Themudo, G., Wielstra, B. & Arntzen, J.W. (2009) Multiple nuclear and mitochondrial genes resolve the branching order of a rapid radiation of crested newts (Triturus, Salamandridae). Molecular Phylogenetics and Evolution, 52(2), 321–328.
Fonseca, M.M., Brito, J.C., Paulo, O.S., Carretero, M.A. & Harris, D.J. (2009) Systematic and phylogeographical assessment of the Acanthodactylus erythrurus group (Reptilia: Lacertidae) based on phylogenetic analyses of mitochondrial and nuclear DNA. Molecular Phylogenetics and Evolution, 51, 131–142.
Fritz, U., Harris, D.J., Fahd, S., Rouag, R., Graciá Martínez, E., Giménez Casalduero, A., Široký, P., Kalboussi, M., Jdeidi, T.B. & Hundsdörfer, A.K. (2009) Mitochondrial phylogeography of Testudo graeca in the Western Mediterranean: old complex divergence in North Africa and recent arrival in Europe. Amphibia–Reptilia, 30(1), 63–80.
Fritz, U. & Havaš, P. (2007) Checklist of Chelonians of the World. Vertebrate Zoology, 57(2), 149–368.
Frost, D.R. (2009) Amphibian Species of the World: an Online Reference. Version 5.3 (12 February, 2009), American Museum of Natural History, New York, USA. Available from: http://research.amnh.org/herpetology/amphibia/ (December 15th, 2009).
Frost, D.R., Grant, T., Faivovich, J., Bain, R.H., Haas, A., Haddad, C.F.B., de Sa, R.O., Channing, A., Wilkinson, M., Donnellan, S.C., Raxworthy, C.J., Campbell, J.A., Blotto, B.L., Moler, P., Drewes, R.C., Nussbaum, R.A.. Lynch, J.D., Green, D.M. & Wheeler, W.C. (2006) The amphibian tree of life. Bulletin of the American Museum of Natural History, 297, 1–370.
Frost, D.R., Grant, T., Faivovich, J., Bain, R.H., Haas, A., Haddad, C.F.B., de Sa, R.O., Channing, A., Wilkinson, M., Donnellan, S.C., Raxworthy, C.J., Campbell, J.A., Blotto, B.L., Moler, P., Drewes, R.C., Nussbaum, R.A.. Lynch, J.D., Green, D.M. & Wheeler, W.C. (2008) Is The Amphibian Tree of Life really fatally flawed? Cladistics, 24, 385–395.
Frost, D.R., McDiarmid, R.W. & Mendelson, J.R. III (2009) Response to the Point of View of Gregory B. Pauly, David M. Hillis, and David C. Cannatella, by the anuran subcommittee of the SSAR/HL/ASIH Scientific and Standard English Name List. Herpetologica, 65(2), 136–153.
Gamble, T., Bauer, A.M., Greenbaum, E. & Jackman, T.R. (2008a) Out of the blue: a novel, trans-Atlantic clade of geckos (Gekkota, Squamata). Zoologica Scripta, 37(4), 355–366.
Gamble, T., Bauer, A.M., Greenbaum, E. & Jackman, T.R. (2008b) Evidence for Gondwanan vicariance in an ancient clade of gecko lizards. Journal of Biogeography, 35, 88–104.
Gans, C. (2005) Checklist and bibliography of the Amphisbaenia of the world. Bulletin of the American Museum of Natural History, 289, 1–130.
García-París, M., Alcobendas, M. & Alberch, P. (1998) Influence of the Guadalquivir river basin on the mitochondrial DNA evolution of Salamandra salamandra (Caudata: Salamandridae) from Southern Spain. Copeia, 1998, 173–176.
García-París, M., Alcobendas, M., Buckley, D. & Wake, D.B. (2003) Dispersal of viviparity across contact zones in Iberian populations of fire salamanders (Salamandra) inferred from discordance of genetic and morphological traits. Evolution, 57(1), 129–143.
Giovannotti, M., Nisi-Cerioni, P., Kalboussi, M., Aprea, G. & Caputo, V. (2007) Phylogeographic inferences from the mtDNA variation of the three-toed skink, Chalcides chalcides (Reptilia: Scincidae). Journal of Experimental Zoology Part B: Molecular and Developmental Evolution, 308B, 297–307.
Geniez, P., Cluchier, A., Sá-Sousa, P., Guillaume, C.P., Crochet, P.-A. (2007) Systematics of the Podarcis hispanicus complex (Sauria, Lacertidae). I. Redefinition, morphology and distribution of the nominotypical taxon. Herpetological Journal, 17, 69–89.
Golay, P., Monney, J.-C., Conelli, A., Durand, T., Thiery, G., Zuffi, M.A.L. & Ursenbacher, S. (2008) Systematics of the Swiss asp vipers: some implications for the European Vipera aspis (Linnaeus, 1758) complex (Serpentes: Viperidae) — a tribute to Kramer. Amphibia–Reptilia, 29(2), 71–83.
Gonçalves, H., Martínez-Solano, I., Pereira, R.J., Carvalho, B., García-París, M. & Ferrand, N. (2009) High levels of population subdivision in a morphologically conserved Mediterranean toad (Alytes cisternasii) result from recent, multiple refugia: evidence from mtDNA, microsatellite and nuclear genealogies. Molecular Ecology, 18(24), 5143–5160.
Gourbière, S. & Mallet, J. (2010) Are species real? The shape of the species boundary with exponential failure, reinforcement, and the "missing snowball". Evolution, 64(1), 1–24.
Guex, G.D. & Grossenbacher, K. (2003) Salamandra atra. In: Grossenbacher, K., Thiesmeier, B. (Eds), Handbuch der Reptilien und Amphibien Europas, Band 4/IIB: Schwanzlurche Urodela. Aula-Verlag, Wiesbaden, pp. 975–1028.
Guicking, D., Joger, U. & Wink, M. (2008) Molecular phylogeography of the viperine snake (Natrix maura). Organisms, Diversity and Evolution, 8, 130–145.
Guicking, D., Joger, U. & Wink, M. (2009) Cryptic diversity in a Eurasian water snake (Natrix tessellata, Serpentes: Colubridae): evidence from mitochondrial sequence data and nuclear ISSR-PCR fingerprinting. Organisms, Diversity and Evolution, 9, 201–214.
Guicking, D., Lawson, R., Joger, U. & Wink, M. (2006) Evolution and phylogeny of the genus Natrix (Serpentes: Colubridae). Biological Journal of the Linnean Society, 87, 127–143.
Guillaume, C.P. (1987) Les petits Lacertidés du basin méditerranéen occidental (Genera Podarcis et Archaeolacerta essentiellement). Sur quelques problémes d’ordre systématique et biogéographique. PhD dissertation, University Montpellier, Montpellier, 463 pp.
Guillaume, C.P. & Lanza, B. (1982) Comparaison electrophorétique de quelques espèces de Lacertidés Méditerranéens, Genera Podarcis et "Archaeolacerta". Amphibia-Reptilia, 4, 361–375.
Gvoždík, V., Jandzik, D., Lymberakis, P., Jablonski, D. & Moravec, J. (2010) Slow worm, Anguis fragilis (Reptilia: Anguidae) as a species complex: genetic structure reveals deep divergences. Molecular Phylogenetics and Evolution, 55(2), 460–472.
Gvoždík, V., Moravec, J. & Kratochvíl, L. (2008) Geographic morphological variation in parapatric Western Palearctic tree frogs, Hyla arborea and Hyla savignyi: are related species similarly affected by climatic conditions? Biological Journal of the Linnean Society, 95, 539–556.
Harris, D.J., Batista, V., Lymberakis, P. & Carretero, M.A. (2004) Complex estimates of evolutionary relationships in Tarentola mauritanica (Reptilia: Gekkonidae) derived from mitochondrial DNA sequences. Molecular Phylogenetics and Evolution, 30(3), 855–859.
Harris, D.J., Carretero, M.A., Corti, C. & Lo Cascio, P. (2009) Genetic affinities of Tarentola mauritanica (Reptilia : Gekkonidae) from Lampedusa and Conigli islet (SW Italy). North–Western Journal of Zoology, 5(1), 197–205.
Herrero, P. (1991) Polytypic chromosomal variation in Triturus boscai (Urodela: Salamandridae). Genetics Selection Evolution, 23(4), 263–272.
Hofman, S., Spolsky, C., Uzzell, T., Cogalniceanu, D., Babik, W. & Szymura, J. (2007) Phylogeography of the fire-bellied toads Bombina: independent Pleistocene histories inferred from mitochondrial genomes. Molecular Ecology, 16, 2301–2316.
International Commission on Zoological Nomenclature (1999) International Code of Zoological Nomenclature. International Trust for Zoological Nomenclature, London, xxix + 306 pp.
International Commission on Zoological Nomenclature (2005) Opinion 2104 (Case 3226) Lacepède, B.G.E. de la V., 1788, Histoire naturelle des Quadrupèdes Ovipares: rejected as a non-binominal work. Bulletin of Zoological Nomenclature, 62(1), 55.
Iraola, A. & García-París, M. (2004) Evidencia molecular de transporte fraudulento y comercio de Salamandra salamandra (Caudata: Salamandridae). Munibe, 55, 217–222.
Joger, U., Fritz, U., Guicking, D., Kalyabina-Hauf, S., Nagy, Z.T. & Wink, M. (2007) Phylogeography of western Palaearctic reptiles — spatial and temporal speciation patterns. Zoologischer Anzeiger, 246, 293–313.
Kalezić, M.L. (1983) Geographical aspects of genetic variability in the smooth newt (Triturus vulgaris). Genetika, 15(1), 93–103.
Kalezić, M.L. & Tucić, N. (1984) Genic diversity and population genetic structure of Triturus vulgaris (Urodela, Salamandridae). Evolution, 38, 389–401.
Kasapidis, P., Magoulas, A., Mylonas, M. & Zouros, E. (2005) The phylogeography of the gecko Cyrtopodion kotschyi (Reptilia: Gekkonidae) in the Aegean archipelago. Molecular Phylogenetics and Evolution, 35, 612–623.
Kalyabina-Hauf, S., Schweiger, S., Joger, U., Mayer, W., Orlov, N. & Wink, M. (2004) Phylogeny and systematics of adders (Vipera berus complex). Mertensiella, 15, 7–16.
Kearney, M. (2003) Systematics of the Amphisbaenia (Lepidosauria: Squamata) based on morphological evidence from recent and fossil forms. Herpetological Monographs, 17, 1–74.
Kearney, M. & Stuart, B.L. (2004) Repeated evolution of limblessness and digging heads in worm lizards revealed by DNA from old bones. Proceedings of the Royal Society of London B, 271, 1677–1683.
Kelly, C.M.R., Barker N.P., Villet, M.H., Broadley D.G. & Branch W.R. (2008) The snake family Psammophiidae (Reptilia: Serpentes): phylogenetics and species delimitation in the African sand snakes (Psammophis Boie, 1825) and allied genera. Molecular Phylogenetics and Evolution, 47(3), 1045–1060.
Kelly, C.M.R., Barker, N.P., Villet, M.H. & Broadley, D.G. (2009) Phylogeny, biogeography and classification of the snake superfamily Elapoidea: a rapid radiation in the late Eocene. Cladistics, 25, 38–63.
Klewen, R. (1988) Die Landsalamander Europas, Teil I, Die Gattungen Salamandra und Mertensiella. A. Ziemsen Verlag, Wittenberg, Lutherstadt, 184 pp.
Kornilios, P., Kyriazi, P., Poulakakis, N., Kumlutaş, Y., Ilgaz, Ç., Mylonas, M. & Lymberakis, P. (2010). Phylogeography of the ocellated skink Chalcides ocellatus (Squamata, Scincidae), with the use of mtDNA sequences: a hitch-hiker’s guide to the Mediterranean. Molecular Phylogenetics and Evolution, 54(2), 445–456.
Lanza, B. (1983) Ipotesi sulle origini del popolamento erpetologico della Sardegna. Lavori Società italiana di Biogeografia, 8, 723–744.
Lanza, B., Andreone, F., Bologna, M.A., Corti, C. & Razzetti E. (2007) Fauna d'Italia, vol. XLII, Amphibia. Calderini, Bologna, 537 pp.
Lanza, B., Pastorelli, C., Laghi, P. & Cimmaruta, R., (2005) A review of systematics, taxonomy, genetics, biogeography and natural history of the genus Speleomantes Dubois, 1984 (Amphibia Caudata Plethodontidae). Atti del Museo Civico di Storia Naturale di Trieste (Suppl.), 52, 5–135.
Lanza, B. & Vanni, S. (1981) On the biogeography of plethodontid salamanders (Amphibia, Caudata), with a description of a new genus. Monitore zoologico italiano, 15, 117–121.
Lapini, L., Dall’Asta, A. & Scaravelli, D. (1992) First record on the occurrence of Triturus v. vulgaris (Linné, 1758) in north-eastern Italy (Amphibia, Caudata, Salamandridae). Gortania–Atti del Museo Friulano di Storia Naturale, 13, 195–201.
Lescure, J. (2008) Note explicative à la liste taxinomique actualisée des Amphibiens et Reptiles de France. Bulletin de la Société Herpétologique de France, 126, 25–36.
Lymberakis, P., Poulakakis, N., Manthalou, G., Tsigenopoulos, C.S., Magoulas, A. & Mylonas, M. (2007) Mitochondrial phylogeography of Rana (Pelophylax) populations in the Eastern Mediterranean region. Molecular Phylogenetics and Evolution, 44, 115–125.
Lymberakis, P., Poulakakis, N., Kaliontzopoulou, A., Valakos, E. & Mylonas, M. (2008) Two new species of Podarcis (Squamata, Lacertidae) from Greece. Systematics and Biodiversity, 6, 307–318.
Mallet, J. (1995) A species definition for the modern synthesis. Trends in Ecology and Evolution, 10, 294–299.
Mallet, J. (2005) Hybridization as an invasion of the genome. Trends in Ecology and Evolution, 20, 229–237.
Mallet, J. (2008) Hybridization, ecological races, and the nature of species: empirical evidence for the ease of speciation. Philosophical Transactions of the Royal Society B-Biological Sciences, 363, 2971–2986.
Martínez-Solano, I., Alcobendas, M., Buckley, D. & García-París, M. (2005) Molecular characterisation of the endangered Salamandra salamandra almanzoris. Annales Zoologici Fennici, 42, 57–68.
Martínez-Solano, I., Teixeira, J., Buckley, D. & García-París, M. (2006) Mitochondrial DNA phylogeography of Lissotriton boscai (Caudata, Salamandridae): evidence for old, multiple refugia in an Iberian endemic. Molecular Ecology, 15, 3375–3388.
Mattoccia, M., Romano, A. & Sbordoni V. (2005) Mitochondrial DNA sequence analysis of the spectacled salamander, Salamandrina terdigitata (Urodela: Salamandridae), supports the existence of two distinct species. Zootaxa, 995, 1–19.
Mertens, R. & Müller, L. (1928) Liste der Amphibien und Reptilien Europas. Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft, 41, 1–62.
Mertens, R. & Wermuth, H. (1960) Die Amphibien und Reptilien Europas. Verlag W. Kramer, Frankfurt, pp. 264.
Montori, A., Llorente, G.A., Alonso-Zarazaga, M.A., Arribas, O., Ayllón, E., Bosch, J., Carranza, S., Carretero, M.A., Galán, P., García-París, M., Harris, D.J., Lluch, J., Márquez, R., Mateo, J.A., Navarro, P., Ortiz, M., Pérez Mellado, V., Pleguezuelos, J.M., Roca, V., Santos, X. & Tejedo, M. (2005) Lista patrón actualizada de la herpetofauna española—Conclusiones de nomenclatura y taxonomía para las especies de anfibios y reptiles de España. AHE, Barcelona, 48 pp.
Nagy, Z.T., Joger, U., Guicking, D. & Wink, M. (2002) Phylogeography of the European Whip Snake Coluber (Hierophis) viridiflavus as inferred from nucleotide sequences of the mitochondrial cytochrome b gene and ISSR genome fingerprinting. Biota, 3(1–2), 109–118.
Nagy, Z.T., Joger, U., Wink, M., Glaw, F. & Vences, M. (2003): Multiple colonization of Madagascar and Socotra by colubrid snakes: evidence from nuclear and mitochondrial gene phylogenies. Proceedings of the Royal Society of London B, 270, 2613–2621.
Nascetti G., Cimmaruta R., Lanza B. & Bullini L. (1996) Molecular taxonomy of European plethodontid salamanders (genus Hydromantes). Journal of Herpetology, 30(2), 161–183.
Noonan, B.P. & Chippindale, P.T. (2006) Dispersal and vicariance: the complex evolutionary history of boid snake. Molecular Phylogenetics and Evolution, 40, 347–358.
Pauly, G.B., Hillis, D.M.; Cannatella, D.C. (2009) Taxonomic freedom and the role of official lists of species names. Herpetologica, 65(2), 115–128.
Paulo, O.S., Pinheiro, J., Miraldo, A., Bruford, M.W., Jordan, C. & Nichols, R.A. (2008) The role of vicariance vs. dispersal in shaping genetic patterns in ocellated lizard species in the western Mediterranean. Molecular Ecology, 17(6), 1535–1551.
Pavlicev, M. & Mayer, W. (2009) Fast radiation of the subfamily Lacertinae (Reptilia: Lacertidae): history or methodological artefact? Molecular Phylogenetics and Evolution, 52, 727–734.
Pellarini, P. & Lapini, L. (2000) Differenze etologiche e morfologiche tra Triturus vulgaris vulgaris (Linnaeus, 1758) e Triturus vulgaris meridionalis (Boulenger, 1882). In: Giacoma, C. (Ed), Atti I Congresso Nazionale Societas Herpetologica Italica, Torino (1996). Museo Regionale Di Scienze Naturali, Torino, pp. 347–351.
Perera, A. & Harris, D.J. (2008) Genetic diversity in the gecko Tarentola mauritanica within the Iberian Peninsula. Amphibia-Reptilia, 29, 583–588.
Perera, A., Vasconcelos, R., Harris, D.J., Brown, R.P., Carretero, M.A. & Pérez-Mellado, V. (2007) Complex patterns of morphological and mtDNA variation in Lacerta perspicillata (Reptilia, Lacertidae). Biological Journal of the Linnean Society, 90, 479–490.
Pinho, C., Harris, D.J. & Ferrand, N. (2008) Non-equilibrium estimates of gene flow inferred from nuclear genealogies suggest that Iberian and North African wall lizards (Podarcis spp.) are an assemblage of incipient species. BMC Evolutionary Biology, 8, 63.
Plötner J. (2005) Die westpaläarktischen Wasserfrösche—Von Märtyrern der Wissenschaft zur biologischen Sensation. Laurenti Verlag, Bielefeld, 160 pp.
Pramuk, J.B., Robertson, T., Sites, J.W.Jr. & Noonan, B.P. (2008) Around the world in 10 million years: biogeography of the nearly cosmopolitan true toads (Anura: Bufonidae). Global Ecology and Biogeography, 17, 72–83.
Rato, C., Zuffi, M.A.L., Corti, C., Fornasiero, S., Gentilli, A., Razzetti, E., Scali, S., Carretero, M.A. & Harris, D.J. (2009) Phylogeography of the European Whip Snake, Hierophis viridiflavus (Colubridae), using mtDNA and nuclear DNA sequences. Amphibia–Reptilia, 30, 283–289.
Raxworthy, C. J. (1990) A review of the smooth newt (Triturus vulgaris) subspecies, including an identification key. Herpetological Journal, 1, 481–492.
Razzetti, E. (2008) Taxonomic remarks on Bufo lineatus and Bufo siculus. In: Corti, C. (Ed), Herpetologia Sardiniae. 7° Congresso Nazionale Societas Herpetologica Italica. Societas Herpetologica Italica / Edizioni Belvedere, Latina, “le scienze”, 8, pp. 420–424.
Renoult, J.P., Geniez, P., Bacquet, P., Benoit, L. & Crochet, P.-A. (2009) Morphology and nuclear markers reveal extensive mitochondrial introgressions in the Iberian Wall Lizard species complex. Molecular Ecology, 18, 4298–4315.
Rhodin, A.G.J., van Dijk, P.P. & Parham, J.F. (2008) Turtles of the world: annotated checklist of taxonomy and synonymy. Chelonian Research Monographs, 5, 1–38.
Ribéron, A., Miaud, C., Grossenbacher, K. & Taberlet, P. (2001) Phylogeography of the Alpine salamander, Salamandra atra (Salamandridae), and the influence of the Pleistocene climatic oscillations on population divergence. Molecular Ecology, 10, 2555–2560.
Ribéron, A., Miaud, C., Guyetant, R. & Taberlet, P. (2004) Genetic variation in an endemic salamander, Salamandra atra, using amplified fragment length polymorphism. Molecular Phylogenetics and Evolution, 31, 910–914.
Romano, A., Mattoccia, M., Marta, S., Bogaerts, S., Pasmans, F. & Sbordoni, V. (2009) Distribution and morphological characterization of the endemic Italian salamanders Salamandrina perspicillata (Savi, 1821) and S. terdigitata (Bonnaterre, 1789) (Caudata: Salamandridae). Italian Journal of Zoology, 76(4), 422–432.
Rosso, A., Castellano, S. & Giacoma, C. (2004) The advertisement call of Hyla intermedia and H. sarda. Italian Journal of Zoology, 71(S2), 169–173.
Rosso, A., Rinella, R., Castellano, S. & Giacoma, C. (2001) Variazione geografica della caratteristiche morfometriche di Hyla sarda: descrizione del pattern ed analisi delle cause. In: Barbieri, F., Bernini, F., Fasola, M. (Eds), Atti 3rd Congresso Nazionale Societas Herpetologica Italica, Pavia (2000). Pianura, Cremona, pp. 265–268.
Salvi, D., Capula, M., Bombi, P. & Bologna, M.A. (2009a) How many Archaeolacerta inhabit the Corso-Sardinian Plate? Allozyme variation and differentiation in Archaeolacerta bedriagae (Camerano, 1885). Amphibia–Reptilia, 30, 463–470.
Salvi, D., Capula, M., Bombi, P. & Bologna, M.A. (2009b) Genetic variation and its evolutionary implications in a Mediterranean island endemic lizard. Biological Journal of the Linnean Society, 98, 661–676.
Salvi, D., Harris, D.J., Bombi, P., Carretero, M.A. & Bologna, M.A. (in press) Mitochondrial phylogeography of the Bedriaga’s rock lizard, Archaeolacerta bedriagae (Reptilia: Lacertidae) endemic to Corsica and Sardinia. Molecular Phylogenetics and Evolution.
Santos, X., Roca, J., Pleguezuelos, J.M., Donaire, J. & Carranza, S. (2008) Biogeography and evolution of the Smooth snake Coronella austriaca (Serpentes: Colubridae) in the Iberian Peninsula: evidence for Messinian refuges and Pleistocenic range expansions. Amphibia-Reptilia, 29, 35–47.
Santucci, F., Nascetti, G. & Bullini, L. (1996) Hybrid zones between two genetically differentiated forms of the pond frog Rana lessonae in southern Italy. Journal of Evolutionary Biology, 9, 429–450.
Schmidtler, J.F. (2004) Der Teichmolch (Triturus vulgaris (L.)), ein Musterbeispiel für systematische Verwechslungen und eine Flut von Namen in der frühen Erforschungsgeschichte. Sekretär, 4(2), 10–28.
Schmidtler, J.F. (2009) Ichthyosaura, der neue Gattungsname für den Bergmolch—ein Lehrbeispiel in Sachen Nomenklatur. Zeitschrift für Feldherpetologie, 16, 245–250.
Schmidtler, J.F. & Frantzen, M. (2004). Triturus vulgaris (Linnaeus, 1758)—Teichmolch. In: Böhme, W. (Ed), Handbuch der Reptilien und Amphibien Europas, Band 4/IIB: Schwanzlurche Urodela. Aula Verlag, Wiesbaden, pp. 847–967.
Schneider, H. (1974) Structure of the mating calls and relationships of the European tree frogs (Hylidae, Anura). Oecologia, 14, 99–110.
Schneider, H. (2002) Bioacoustic demonstration of the Tree Frog, Hyla arborea (Amphibia: Anura), in western Turkey. Zoology in the Middle East, 21, 77–85.
Schneider, H., Sinsch, U. & Sofianidou, T.S. (1993) The water frogs of Greece. Bioacoustic evidence for a new species. Zeitschrift für Zoologische Systematik und Evolutionsforschung, 31, 47–63.
Schreiber, E. (1875) Herpetologia Europaea. Friedrich Vieweg und Sohn, Braunschweig, Germany, pp. 670.
Sites, J.W. & Marshall, J.C. (2004) Operational criteria for delimiting species. Annual Review of Ecology, Evolution, and Systematics, 35, 199–227.
Sotiropoulos, K., Eleftherakos, K., Džukić, G., Kalezić, M.L., Legakis, A. & Polymeni, R.M. (2007) Phylogeny and biogeography of the alpine newt Mesotriton alpestris (Salamandridae, Caudata), inferred from mtDNA sequences. Molecular Phylogenetics and Evolution, 45(1), 211–226.
Speybroeck, J. & Crochet, P.-A. (2007) Species list of the European herpetofauna—a tentative update. Podarcis, 8(1/2): 8–34.
Spinks, P.Q. & Shaffer, H.B. (2009) Conflicting mitochondrial and nuclear phylogenies for the widely disjunct Emys (Testudines: Emydidae) species complex, and what they tell us about biogeography and hybridization. Systematic Biology, 58(1), 1–20.
Steinfartz, S., Veith, M. & Tautz, D. (2000) Mitochondrial sequence analysis of Salamandra taxa suggests old splits of major lineages and postglacial recolonizations of Central Europe from distinct source populations of Salamandra salamandra. Molecular Ecology, 9, 397–410.
Stöck, M., Moritz, C., Hickerson, M., Frynta, D., Dujsebayeva, T., Eremchencko, V., Macey, J.R., Papenfuss, T.J. & Wake, D.B. (2006) Evolution of mitochondrial relationships and biogeography of Palearctic green toads (Bufo viridis subgroup) with insights in their genomic plasticity. Molecular Phylogenetics and Evolution, 41, 663–689.
Stöck, M., Sicilia, A., Belfiore, N., Buckley, D., Lo Brutto, S., Lo Valvo, M. & Arculeo, M. (2008a) Post-Messinian evolutionary relationships across the Sicilian channel: mitochondrial and nuclear markers link a new green toad from Sicily to African relatives. BMC Evolutionary Biology, 8, 56.
Stöck, M., Dubey, S., Klütsch, C., Litvinchuk, S.N., Scheidt, U. & Perrin, N. (2008b) Mitochondrial and nuclear phylogeny of circum-Mediterranean tree frogs from the Hyla arborea group. Molecular Phylogenetics and Evolution, 49(3), 1019–1024.
Tomović, L. (2006) Systematics of the nose-horned viper (Vipera ammodytes, Linnaeus, 1758). Herpetological Journal, 16, 191–201.
Uetz, P. & Hallermann, J. (2009) The TIGR Reptile Database, JCVI, Rockville. Available from: http://www.reptile-database.org/ (December 15th, 2009).
Ursenbacher, S., Conelli, A., Golay, P., Monney, J.-C., Zuffi, M. A. L., Thierry, G., Durand, T. & Fumagalli, L. (2006) Phylogeography of the asp viper (Vipera aspis) inferred from mitochondrial DNA: evidence for multiple Mediterranean refugial areas. Molecular Phylogenetics and Evolution, 38, 546–552.
Ursenbacher, S., Schweiger, S., Tomović, L., Crnobrnja-Isailović, J., Fumagalli, L. & Mayer, W. (2008) Molecular phylogeography of the nose-horned viper (Vipera ammodytes, Linnaeus (1758)): Evidence for high genetic diversity and multiple refugia in the Balkan peninsula. Molecular Phylogenetics and Evolution, 46(3), 1116–1128.
Vasconcelos ”Vaconcelos”, R., Carretero, M.A. & Harris, D.J. (2006) Phylogeography of the genus Blanus (worm lizards) in Iberia and Morocco based on mitochondrial and nuclear markers—preliminary analysis. Amphibia–Reptilia, 27(3), 339–346.
Van Bocxlaer, I., Biju, S.D., Loader, S.P. & Bossuyt, F. (2008) Toad radiation reveals into-India dispersal as a source of endemism in the Western Ghats-Sri Lanka biodiversity hotspot. BMC Evolutionary Biology, 8, 131.
van der Meijden, A., Chiari, Y., Mucedda, M., Carranza, S., Corti, C. & Veith, M. (2009) Phylogenetic relationships of Sardinian cave salamanders, genus Hydromantes, based on mitochondrial and nuclear DNA sequence data. Molecular Phylogenetics and Evolution, 51(2), 399–404.
Velo-Antón, G., Martínez-Solano, I. & García-París, M. (2008) β-fibrinogen intron 7 variation in Discoglossus (Anura: Discoglossidae): implications for the taxonomic assessment of morphologically cryptic species. Amphibia–Reptilia, 29, 523–533.
Vences, M. (2007) The amphibian tree of life: Ideologie, Chaos oder biologische Realität? Zeitschrift für Feldherpetologie, 14, 153–162.
Venchi, A. & Bologna, M.A. (1996) Lacerta parva Boulenger, a new lizard species for the European fauna. Amphibia-Reptilia, 17, 89–90.
Vidal, N., Azvolinsky, A., Cruaud, C. & Hedges, S.B. (2008) Origin of tropical American burrowing reptiles by transatlantic rafting. Biology Letters, 4, 115–118.
Vidal, N. & Hedges, S.B. (2002) Higher-level relationships of caenophidian snakes inferred from four nuclear and mitochondrial genes. Comptes Rendus Biologies, 325, 987–995.
Vieites, D.R., Min, M.-S. & Wake, D.B. (2007) Rapid diversification and dispersal during periods of global warming by plethodontid salamanders. Proceedings of the National Academy of Sciences, 104, 19903–19907.
Wake, D.B., Salvador, A. & Alonso-Zarazaga, M.A. (2005) Taxonomy of the plethodontid salamander genus Hydromantes (Caudata: Plethodontidae). Amphibia-Reptilia, 26, 543–548.
Wiens, J.J. (2007) Review: The Amphibian Tree of Life. Quarterly Review of Biology, 82, 55–56.
Zaher, H., Grazziotin, F.G., Cadle, J.E., Murphy, R.W., de Moura-Leite, J.C. & Bonatto, S.L. (2009) Molecular phylogeny of advanced snakes (Serpentes, Caenophidia) with an emphasis on South American Xenodontines: a revised classification and descriptions of new taxa. Papéis Avulsos de Zoologia, 49(11), 115–153.
Zangari, F., Cimmaruta, R. & Nascetti, G. (2006) Genetic relationships of the western Mediterranean painted frogs based on allozymes and mitochondrial markers: evolutionary and taxonomic inferences (Amphibia, Anura, Discoglossidae). Biological Journal of the Linnean Society, 87(4), 515–536.
Zheng, Y., Fu, J. & Li, S. (2009) Toward understanding the distribution of Laurasian frogs: a test of Savage’s biogeographical hypothesis using the genus Bombina. Molecular Phylogenetics and Evolution, 52, 70–83.
Zinenko, O., Ţurcanu, V. & Strugariu, A. (2010) Distribution and morphological variation of Vipera berus nikolskii Vedmederja, Grubant et Rudaeva, 1986 in Western Ukraine, The Republic of Moldova and Romania. Amphibia-Reptilia, 31(1), 51–67.