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Type: Articles
Published: 2009-12-22
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Oribatid mite (Arachnida: Oribatida) coenoses from SW Sardinia

Department of Evolutionary Biology, University of Siena, via A. Moro 2, I-53100 Siena, Italy
Arachnida Oribatidae soil fauna faunistics biogeography population dynamics Italy

Abstract

Oribatid mite communities were investigated at five sites in SW Sardinia (Carbonia-Iglesias and Medio Campidano provinces) in three representative habitat types: holm oak (Quercus ilex) forest, cork oak (Quercus suber) forest and pineland (Pinus radiata and P. pinaster). A total of 1,180 oribatid mite specimens, belonging to 67 different species, were extracted from soil samples. Significant differences in species diversity, abundance and Berger-Parker index of dominance were found among the different vegetation types throughout the sampling period, and non-metric multidimensional scaling (n-MDS) ordination confirmed a clear separation between the three coenoses, showing how oribatid assemblages are significantly affected by habitat evolution. The faunistic knowledge of the oribatid mites of Sardinia is rather poor, the present research represents one of the first contributions for the main Sardinian island. Besides a majority of eurytopic and widespread species representing the main part of the edaphic communities in Mediterranean landscapes, some rare and characteristic species are recorded. They are remnants of the original populations of Sardinia and the western part of the Italian peninsula, resulting from the complex tectonic events that formed the western Mediterranean basin. Among these, Belorchestes gebennicus Grandjean, 1957 (Zetorchestidae), Microppia minus longisetosa Subías & Rodriguez, 1988, and Pluritrichoppia insolita Subías & Arillo, 1989 (Oppiidae) had never been collected in Italy, single individuals of Oxyoppioides and Thamnacarus probably belong to undescribed species, while Brachychthonius hirtus Moritz, 1976 (Brachychthoniidae), Mongaillardia aeoliana (Bernini, 1979) (Amerobelbidae), Berniniella aeoliana (Bernini, 1973), Lauroppia similifallax Subías & Minguez, 1986, Ramusella (Ramusella) gyrata (Mahunka & Paoletti, 1984) (Oppiidae) and Ophidiotrichus oglasae Bernini, 1975 (Oribatellidae) are new to the Sardinian fauna.

References

  1. Avanzati, A.M., Salomone, N., Baratti, M. & Bernini, F. (2003) Taxonomic revision of Amerus troisii (Berlese, 1883) (Acari, Oribatida, Ameridae) using morphological and biochemical characters. Journal of Natural History, 37, 797–819.

    Balogh, J. (1943) Magyarország, Páncélosatkái (Conspectus oribateorum hungariae). Matematekai Természtttudományi Közeményei, Budapest, 202 pp.

    Balogh, J. & Mahunka, S. (1983) Primitive Oribatids of the Palaearctic Region. Elsevier, Amsterdam, Oxford, New York, 372 pp.

    Baratti, M. & Bernini, F. (1994) Taxonomic revision of Carabodes coriaceus C. L. Koch, 1836 and C. arduinii Valle, 1955 (Acarida Oribatida). Acarologia, 35, 247–265.

    Bardgett, R., Wardle, D.A. & Yeates, G.W. (1998) Linking above-ground and below-ground food webs: how plant responses to foliar herbivory influence soil organisms. Soil Biology and Biochemistry, 30, 1867–1878.

    Behan-Pelletier, V.M. (1999) Oribatid mite biodiversity in agroecosystems: role for bioindication. Agriculture Ecosystems and Environment, 74, 411–423.

    Berger, W.H. & Parker, F.L. (1970) Diversity of planktonic Foraminifera in deep-sea sediments. Science, 168, 1345–1347.

    Bergman, B., Stortelder, A. & Westoff, V. (1998) Contenuto di informazione floristica e vegetazionale. In: Bas Pedroli, G.M., Vos, W., Dijkstra, H. & Rossi, R. (Eds), Studio degli effetti ambientali della diga sul Torrente Farma. Una procedura di valutazione di impatto ambientale applicata al progetto di derivazione di acque della Val di Farma (Toscana, Italia). Progetto Toscana. Serie di ambiente, territorio, economia della Regione Toscana, 4. Marsilio editore, Venezia, pp. 307–312.

    Berlese, A. (1883) Acari, Myriopoda et Scorpiones hucusque in Italia reperta. Cryptostigmata, fasc. III, Padova.

    Berlese, A. (1903) Acari nuovi. Manipulus I. Redia, 1, 235–252.

    Berlese, A. (1904) Acari nuovi. Manipulus III. Redia, 2, 10–32.

    Berlese, A. (1905) Acari nuovi. Materiali pel “Manipulus V”. Redia, 2, 231–238.

    Berlese, A. (1908) Elenco di generi e specie nuove di Acari. Redia, 5, 1–15.

    Berlese, A. (1910a) Acari nuovi. Manipulus V, VI. Redia, 6, 199–234.

    Berlese, A. (1910b) Brevi diagnosi di generi e specie nuovi di Acari. Redia, 6, 346–388.

    Berlese, A. (1916) Centuria prima di Acari nuovi. Redia, 12, 19–67.

    Berlese, A. (1923) Centuria sesta di Acari nuovi. Redia, 15, 237–262.

    Bernini, F. (1973) Notulae Oribatologicae VII. Gli Oribatei (Acarida) dell'isolotto di Basiluzzo (Isole Eolie). Lavori della Società Italiana di Biogeografia, 3, 355–480.

    Bernini, F. (1975) Notulae Oribatologicae XIII. La famiglia Oribatellidae (Acarida, Oribatei). Lavori della Società Italiana di Biogeografia, 5, 429–507.

    Bernini, F. (1979a) Notulae Oribatologicae XXIII. A new Amerobelbid mite (Acarida, Oribatida) from Lipari (Aeolian islands). Animalia, 6, 257–265.

    Bernini, F. (1979b) Biogeographic and faunistic data on the Oribatids of the Tuscan Archipelago. In: Rodriguez, J.G. (Ed.), Recent Advances in Acarology Volume II. Academic Press, New York, pp. 559–565.

    Bernini, F. (1983) Notulae Oribatologicae XXVIII. Some Italian Protoplophoridae (Acarida, Oribatida). Redia, 66, 47–86.

    Bernini, F. & Avanzati, A.M. (1989a) Notulae Oribatologicae XLVIII. Taxonomic position of Steganacarus brevipilus (Berlese, 1923) and Tropacarus in the Steganacaridae system (Acarida, Oribatida). International Journal of Acarology, 15, 5–16.

    Bernini, F. & Avanzati, A M. (1989b) Notulae Oribatologicae XLIX. Taxonomic revision of Steganacarus (Steganacarus) applicatus (Sellnick, 1920) and the description of a new West-Mediterranean Steganacarus species (Acarida, Oribatida). International Journal of Acarology, 15, 65–74.

    Bernini, F., & Nannelli, R. (1982) Notulae Oribatologicae XXVI. Contribution to the knowledge of the genus Cepheus (Oribatida, Acarida) in Italy. Redia, 65, 155–188.

    Bernini, F., Castagnoli, M. & Nannelli, R. (1995) Arachnida Acari. In: Minelli, A., Ruffo, S. & La Posta, S. (Eds), Checklist delle specie della fauna italiana, 24. Calderini, Bologna, pp. 1–131.

    Bray, J.R. & Curtis, J.T. (1957) An ordination of the upland forest communities of Southern Wisconsin. Ecological Monographs, 27, 325–349.

    Cancela da Fonseca, J.P. (1991) Ecological diversity and ecological systems complexity: local or global approach? Revue d’Ecologie et de Biologie du Sol, 28, 51–66.

    Canestrini, G & Canestrini, R. (1882) Acari italiani nuovi e poco noti. Atti del Reale Istituto Veneto di Scienze, Lettere e Arti, 5, 1–4.

    Caruso, T. & Migliorini, M. (2006) A new formulation of the geometric series with applications to oribatid (Acari, Oribatida) species assemblages from human-disturbed Mediterranean areas. Ecological Modelling, 95, 402–406.

    Caruso, T., Pigino, G., Bernini, F., Bargagli, R. & Migliorini, M. (2007) The Berger-Parker index as an effective tool for monitoring the biodiversity of disturbed soils: a case study on Mediterranean oribatid (Acari: Oribatida) assemblages. Biodiversity and Conservation, 16, 3277–3285.

    Clarke, K.R. & Warwick, R.M. (1994) Change in marine community: an approach to statistical analysis and interpretation. Plymouth Marine Laboratory, Natural Environment Council, 144 pp.

    Coggi, A. (1898) Descrizione di specie nuove di Oribatidi italiani. Bulletino della Società Entomologica Italiana, 30, 68–83.

    Cole, L., Bradford, M.A., Shaw, P.J.A. & Bardgett, R.D. (2006) The abundance, richness and functional role of soil meso- and macrofauna in temperate grassland – a case study. Applied Soil Ecology, 33, 186–198.

    Cox, T.F. & Cox, A.A. (1994) Multidimensional Scaling. Chapman and Hall, London, 313 pp.

    Forsslund, K.-H. (1942) Schwedische Oribatei (Acari). I. Arkiv för Zoologi, 34, 1–11.

    Forsslund, K.-H. (1963) Notizen über Oribatei (Acari). III. Entomologisk Tidskrift, 84, 282–283.

    Fountain, M.T. & Hopkin, S.P. (2001) Biodiversity of Collembola in urban soils and the use of Folsomia candida to assess soil “quality”. Ecotoxicology, 13, 555–572.

    Fujikawa, T., Fujita, M. & Aoki, J. (1993) Checklist of Oribatid Mites of Japan (Acari: Oribatida). Journal of the Acarological Society of Japan, 2 (Suppl. 1), 1–121.

    Golosova, L., Karppinen, E. & Krivolutsky, D.A. (1983) List of oribatid mites (Acarina, Oribatei) of northern palaearctic region. II. Siberia and the Far East. Acta Entomologica Fennica, 43, 1–14.

    Grandjean, F. (1931) Le genre Licneremaeus Paoli (Acariens). Bulletin de la Societé Zoologique de France, 56, 221–250.

    Grandjean, F. (1932) Observations sur les Oribates (3me série). Bulletin du Muséum National d’Histoire Naturelle, Paris, 4, 292–306.

    Grandjean, F. (1957) Belorchestes gebennicus n.sp. nouvel Oribate sauteur. Bulletin du Muséum National d’Histoire Naturelle, Paris, 29, 148–155.

    Henig-Sever, N., Poliakov, D. & Broza, M. (2001) A new method for estimation of wildfire intensity based on ash pH and soil micro-arthropod community. Pedobiologia, 45, 98–106.

    Hooper, D.U., Bignell, D.E., Brown, V.K., Brussaard, L., Wardle, D.A., Coleman, D.C., Giller, K.E., Lavelle, P., van der Putten, P.W.H., de Ruiter, P.C., Rusek, J., Silver, W.L., Tiedje, J.M. & Wolters, V. (2000) Interactions between aboveground and belowground biodiversity in terrestrial ecosystems: patterns, mechanisms, and feedbacks. Bioscience, 50, 1049–1061.

    Jacot, A.P. (1935) On Galumna alatus Willmann. Annals and Magazine of Natural History, 15, 489–492.

    Karppinen, E. & Krivolutsky, D.A. (1982) List of oribatid mites (Acarina, Oribatei) of northern paleartic region. I. Europe. Acta Entomologica Fennica, 41, 1–18.

    Karppinen, E., Krivolutsky, D.A. & Poltavskaja, M.P. (1986) List of oribatid mites (Acarina, Oribatei) of the northern palearctic region. III. Arid lands. Annales Entomologici Fennici, 52, 81–94.

    Karppinen, E., Krivolutsky, D.A., Tarba, Z.M., Shtanchaeva, U.Y. & Gordeeva, E.W. (1987) List of oribatid mites (Acarina, Oribatei) of northern region. IV. Caucasus and Crimea. Annales Entomologici Fennici, 53, 119–137.

    Ketmaier, V., Giusti, F. & Caccone, A. (2006) Molecular phylogeny and historical biogeography of the land snail genus Solatopupa (Pulmonata) in the peri-Tyrrhenian area. Molecular Phylogenetics and Evolution, 39, 439–451.

    Koch, C.L. (1839) Deutschlands Crustaceen, Myriapoden und Arachniden, 29–30. Regensburg.

    Koch, C.L. (1841) Deutschlands Crustaceen, Myriapoden und Arachniden, 31–32. Regensburg.

    Krisper, G. (1984) Wiederbeschreibung und Verbreiitungsanalyse der bodenbewohnenden Milbe Zetorchestes falzonii Coggi (Acari, Oribatei). Mitteilungen des Naturwissenschaftlichen Vereines für Steiermark, 114, 331–350.

    Krisper, G. (1987) Artengliederung und Verbreitung der Gattung Zetorchestes in Europa (Acari, Oribatide). Zoologische Mededelingen, 61, 15–30.

    Kruskal, J.B. & Wish, M. (1978) Multidimensional Scaling. Sage, Beverly Hills, California, 97 pp.

    Lamoncha, K.L. & Crossley, D.A. jr. (1998) Oribatid mite diversity along an elevation gradient in a southeastern Appalachian forest. Pedobiologia, 42, 43–55.

    Lawton, J.H. (1994) What do species do in ecosystems? Oikos, 72, 367–374.

    Lindo, Z. & Visser, S. (2004) Forest floor microarthropod abundance and oribatid mite (Acari: Oribatida) composition following partial and clear-cut harvesting in the mixed-wood boreal forest. Canadian Journal of Forest Research, 34, 998–1006.

    Linnaeus, C. (1758) Systema Naturae per Regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Ed. 10a. Vol. I. Laurentii Salvii. Holmiae, 2 + 824 pp.

    Lions, J.-C. (1966) Oribates (Acariens) de la Provence: Nellacarus adansoni n.sp. Vie et Milieu, 2C, 795–807.

    Magurran, A.E. (1988) Ecological diversity and its measurement. Croom Helm, London, 179 pp.

    Mahunka, S. & Paoletti, M.G. (1984) Oribatid mites and other mites (Tarsonemida, Anoetidae, Acaridae) from woods and farms monocultivated with corn in the low laying plain (Veneto and Friuli, NE Italy). Redia, 67, 93–128.

    Maraun, M. & Scheu, S. (2000) The structure of oribatid mite communities (Acari, Oribatida): patterns, mechanisms and implication for future research. Ecography, 23, 374–383.

    Marshall, V.G., Reeves R.M. & Norton, R.A. (1987) Catalogue of the Oribatida (Acari) of continental United States and Canada. Memoirs of the Entomological Society of Canada, 139, 1–418.

    Mason, F., Cerretti, P., Nardi, G., Whitmore, D., Birtele, D., Hardersen, S. & Gatti, E. (2006) Aspects of biological diversity in the CONECOFOR plots. IV. The InvertebrateBiodiv pilot project. In: Ferretti, M., Petriccione, B., Bussotti, F. & Fabbio, G. (Eds), Aspects of biodiversity in selected forest ecosystems in Italy: status and changes over the period 1996-2003. Third report of the Task Force on Integrated and Combined (I&C) evaluation of the CONECOFOR programme. Annali dell’Istituto sperimentale per la Selvicoltura, 30 (Suppl. 2), 51–70.

    May, R.M. (1975) Patterns of species abundances and diversity. In: Cody, M.L. & Diamond, J.M. (Eds), Ecology and Evolution of Communities. Belknap Press of Harvard University Press, Cambridge, Massachusetts, pp. 81–120.

    Menke, H.-G. (1964) Revision of Ceratozetidae, 3. Ceratozetes laticuspidatus n.sp. (Arach., Acari, Oribatei). Senckenbergiana Biologia, 45, 635–640.

    Michael, A.D. (1880) A further contribution to the knowledge of British Oribatidae (Part II). Journal of the Royal Microscopical Society, 3, 177–201.

    Michael, A.D. (1882) Further notes on British Oribatidae. Journal of the Royal Microscopical Society, 2, 10.

    Michael, A.D. (1885) New British Oribatidae. Journal of the Royal Microscopical Society, 5, 385–397.

    Michael, A.D. (1888) British Oribatidae, vol. 2. The Ray Society, London, pp. 337–657.

    Migliorini, M, Petrioli, A. & Bernini, F. (2002) Comparative analysis of two edaphic zoocoenoses (Oribatid mites and Carabid beetles) in five habitats of the ‘Pietraporciana’ and ‘Lucciolabella’ Nature Reserves (Orcia Valley, central Italy). Acta Oecologica, 23, 361–374.

    Migliorini, M., Pigino, G., Avanzati, A.M., Salomone, N. & Bernini, F. (2005a) Experimental fires in Mediterranean environment: effects on oribatid mite communities, In: Weigmann, G., Alberti, G., Wohltmann, A. & Ragusa, S. (Eds), Phytophaga XIV. Acarine biodiversity in the natural and human sphere. Proceedings of 5thSymposium of the European Association of Acarologists, Berlin 2004. Luxograph, Palermo, pp. 271–278.

    Migliorini, M., Pigino, G., Caruso, T., Fanciulli, P.P., Leonzio, C. & Bernini, F. (2005b) Soil communities (Acari Oribatida; Hexapoda Collembola) in a clay pigeon shooting range. Pedobiologia, 49, 1–13.

    Mihelčič, F. (1955) Oribatiden der Iberischen Halbinsel II. Zoologicher Anzeiger, 155, 306–309.

    Mihelčič, F. (1956) Oribatiden Südeuropas V. Zoologicher Anzeiger, 157, 154–174.

    Minguez, M.E, Ruiz, E. & Subías L.S. (1985) El género Quadroppia Jacot, 1939 (Acari, Oribatida, Oppiidae). Boletín de la Asociación Española de Entomología, 9, 95–118.

    Moore, J.C., Walter, D.E. & Hunt, H.W. (1988) Arthropod regulation of micro- and mesobiota in below-ground detrital food webs. Annual Review of Entomology, 33, 419–439.

    Moritz, M. (1976) Revision der Europäischen gattungen und arten der famiglie Brachychthoniidae (Acari, Oribatei). Mitteilungen aus dem Zoologischen Museum in Berlin, 52, 227–319.

    Nicolet, H. (1855) Histoire Naturelle des Acariens qui se trouvent aux environs de Paris. Archives du Muséum National d’Histoire Naturelle, Paris, 7, 381–482.

    Niedbala, W. (1992) Phthiracaroidea (Acari, Oribatida) Systematic Studies. Elsevier Science Publishers, Amsterdam, Oxford, New York, Tokyo and PWN, Polish Scientific Publishers, Warszawa, 612 pp.

    Niedbala, W. (2004) Fauna Europaea: Acari, Oribatida. In: Magowski, W. (Ed.), Fauna Europaea: Acari: Acariformes. Fauna Europaea version 1.1, available at http://www.faunaeur.org [accessed September 2009 as version 1.3 of April 19th 2007.]

    Norton, R.A. (1985) Aspects of the biology and systematics of soil arachnids, particularly saprophagous and mycophagus mites. Quaestiones Entomologicae, 21, 523–541.

    Norton, R.A. (1990) Acarina: Oribatida. In: Dindal, D.L. (Ed.), Soil Biology Guide. John Wiley & Sons Inc., New York, pp. 779–803.

    Noti, M.I., André, H.M., Ducarme, X. & Lebrun, P. (2003) Diversity of soil oribatid mites (Acari: Oribatida) from High Katanga (Democratic Republic of Congo): multiscale and multifactor approach. Biodiversity and Conservation, 12, 767–785.

    Oudemans, A.C. (1900) New list of Dutch Acari, 1st part. Tijdschrift voor Entomologie, 43, 150–171.

    Oudemans, A.C. (1902a) Acarologische Aanteekeningen. Entomologische Berichten, 1, 36–39.

    Oudemans, A.C. (1902b) New list of Dutch Acari, Second Part. With remarks on known and description of a new genera and species. Tijdschrift voor Entomologie, 45, 50–64.

    Oudemans, A.C. (1914). Acarologisches aus Maulwurfsnestern. Archiv fur Naturgeschicthe, 79, 24–45.

    Paoli, G. (1908) Monografia del genere Dameosoma Berl. e generi affini. Redia, 5, 31–91.

    Pavlitshenko, P.G. (1994) A guide to the Ceratozetoid Mites (Oribatei, Ceratozetoidea) of Ukraine. Natsionalnoi Akademii Nauk Ukrainy, Kiev, 142 pp.

    Pérez-Iñigo, C. (1970) Acaros oribatidos de suelos de España peninsular e islas Baleares (Acari, Oribatei) (parte II). Eos, 45, 241–317.

    Pérez-Iñigo, C. (1987) Oribatidos de las islas Azores (I) (Acari: Oribatei). Eos, 63, 197–228.

    Pérez-Iñigo, C. (1993) Acari. Oribatei, Poronota. In: Ramos, M.A. et al. (Eds), Fauna Iberica, vol. 3. Museo Nacional de Ciencias Naturales, CSIC, Madrid, 320 pp.

    Pérez-Iñigo, C. (1997) Acari. Oribatei, Gymnonota I. In: Ramos, M.A. et al. (Eds) Fauna Iberica, vol. 9. Museo Nacional de Ciencias Naturales, CSIC, Madrid, 374 pp.

    Putten, W.H. (van der), Vet, L.E.M., Harver, J.A. & Wäckers, F.L. (2001) Linking above- and belowground multitrophic interactions of plants, herbivores, pathogens and their antagonists. Trends in Ecology and Evolution, 16, 547–554.

    Salomone, N, Avanzati, A.M., Baratti, M. & Bernini, F. (2003) Genetic and morphological analysis of some European species of the “coriaceus group” of Carabodes (Acari, Oribatida, Carabodidae) and description of C. tyrrhenicus sp. nov. Zoologischer Anzeiger, 242, 121–136.

    Straalen, N.M. (van) (1998) Evaluation of bioindicator systems derived from soil arthropod communities. Applied Soil Ecology, 9, 429–437.

    Subías, L.S. (2004) Listado sistemático, sinonímico y biogeográfico de los ácaros oribátidos (Acariformes, Oribatida) del mundo. Graellsia, 60 (número extraordinario), 3–305.

    Subías, L.S. & Arillo, A. (1989) Pluritrichoppia insolita gen. nov., sp. nov. (Acari, Oribatida, Oppiidae) del sur de la España. Redia, 72, 251–257.

    Subías, L.S. & Arillo, A. (2001) Acari. Oribatei, Gymnonota II. In: Ramos, M.A., Tercedor, J.A., Ros, X.B., Noguera, J.G., Sierra, A.G., Macpherson Mayol, E., Piera, F.M., Marino, J.S. & Gonzáles, J.T. (Eds), Fauna Iberica, vol. 15. Museo Nacional de Ciencias Naturales, CSIC, Madrid, 289 pp.

    Subías, L.S. & Gil-Martin, J. (1997) Systematic and biogeographic checklist of oribatids from Western Mediterranean (Acari, Oribatida). Annali del Museo Civico di Storia Naturale “G. Doria”, 91, 459–498.

    Subías, L.S. & Minguez, M.E. (1986) Lauroppia similifallax n.gen., n.sp. (Acari, Oribatida, Oppiidae) y descripcion de la España central. Boletín de la Asociación Española de Entomología, 10, 51–58.

    Subías, L.S. & Rodriguez, P. (1988) Oppiidae (Acari, Oribatida) de los sabinares (Juniperus thurifera) de España, VIII. Medioppiinae Subías y Minguez. Boletín de la Asociación Española de Entomología, 12, 27–43.

    Subías, L.S., Arillo, A. & Subías, J. (1997) The genus Arthrodamaeus Grandjean (Acariformes, Oribatida, Gymnodamaeidae). Acarologia, 38, 297–307.

    Trägårdh, I. (1902) Beiträge zur kenntnis der Schwdischen Acaridefauna I. Lappländische Trombiiden und Oribatiden. Bihang till Kongliga Svenska Vetenskaps-Akademiens Handlingar, 28, 1–26.

    Trägårdh, I. (1910) Acariden aus dem Sarekgebirge. Naturwissenschaftliche Untersuchungen des Sarekgebirges in Schwedisch-Lappland, 4, 375–580.

    Usher, M.B., Booth, R.G. & Sparkes, K.E. (1982) A review of progress in understanding the organization of communities of soil arthropods. Pedobiologia, 23, 126–144.

    Valle, A. (1955) Su due nuove specie di Acari raccolti all’sola di Zannone. Atti della Società Italiana di Scienze Naturali, 94, 3–4.

    Vigna Taglianti, A., Audisio, P.A., Belfiore, C., Biondi, M., Bologna, M.A., Carpaneto, G.M., De Biase, A., De Felici, S., Piattella, E., Racheli, T., Zapparoli, M. & Zoia, S. (1993) Riflessioni di gruppo sui corotipi fondamentali della fauna W-paleartica ed in particolare italiana. Biogeographia, Lavori della Società Italiana di Biogeografia (n.s.), 16 [1992], 159–179.

    Vigna Taglianti, A., Audisio, P.A., Biondi, M., Bologna, M.A., Carpaneto, G.M., De Biase, A., Fattorini, S., Piattella, E., Sindaco, R., Venchi, A. & Zapparoli, M. (1999) A proposal for a chorotype classification of the Near East fauna, in the framework of the Western Palearctic region. Biogeographia, Lavori della Società Italiana di Biogeografia (n.s.), 20, 31–59.

    Vreeken-Buijs, M.J., Hassink, J. & Brussaard, L. (1998) Relationships of soil microarthropod biomass with organic matter and pore size distribution in soils under different land use. Soil Biology and Biochemistry, 30, 97–106.

    Wardle, D.A. (2002) Community and ecosystems. Linking the aboveground and belowground components. Monographs in Population Biology. Princeton University Press, Princeton, 392 pp.

    Webb, N.R. (1994) Post-fire succession of cyptostigmatic mites (Acari, Cryptostigmata) in a Calluna-heathland soil. Pedobiologia, 38, 138–145.

    Weigmann, G. (2006) Acari, Actinochaetida. Hornmilben (Oribatida). Goecke & Evers, Keltern, 520 pp.