Abstract
The endemic South African spider genus Austrachelas Lawrence, 1938 is revised. The type species, A. incertus Lawrence, 1938, and A. natalensis Lawrence, 1942, are redescribed, and their males described for the first time. Seven new species are described: A. bergi n. sp. (female only), A. kalaharinus n. sp. (male only), A. merwei n. sp. (both sexes), A. pondoensis n. sp. (both sexes), A. reavelli n. sp. (both sexes), A. sexoculata n. sp. (male only), and A. wassenaari n. sp. (both sexes). A cladistic analysis performed including Austrachelas and various species of the corinnid subfamilies Trachelinae, Castianeirinae, Corinninae and Phrurolithinae, and species from the Liocranidae, Gallieniellidae, Gnaphosidae and Lamponidae, suggests that Austrachelas is currently misplaced. Its transfer to the Gallieniellidae is proposed.References
Bosselaers, J. (2002) A cladistic analysis of Zoropsidae (Araneae), with the description of a new genus. Belgian Journal of Zoology, 132, 141–154.
Bosselaers, J. & Jocqué, R. (2000) Studies in Corinnidae: transfer of four genera and description of the female of Lessertina mutica Lawrence 1942. Tropical Zoology, 13, 305–325.
Bosselaers, J. & Jocqué, R. (2002) Studies in Corinnidae: cladistic analysis of 38 corinnid and liocranid genera, and transfer of Phrurolithinae. Zoologica Scripta, 31, 241–270.
Bosselaers, J., Urones, C., Barrientos, J.A. & Alberdi, J.M. (2009) On the Mediterranean species of Trachelinae (Araneae, Corinnidae), with a revision of Trachelas L. Koch, 1872 on the Iberian Peninsula. Journal of Arachnology, 37, 15–38.
Bremer, K. (1988) The limits of amino acid sequence data in angiosperm phylogenetic reconstruction. Evolution, 42, 795–803.
Bremer, K. (1994) Branch support and tree stability. Cladistics, 10, 295–304.
Coddington, J.A. & Scharff, N. (1994) Problems with zero-length branches. Cladistics, 10, 415–423.
Dippenaar-Schoeman, A.S. & Jocqué, R. (1997) African spiders: an identification manual. Plant Protection Research Institute Handbook 9. Agricultural Research Council, Pretoria, 392 pp.
Goloboff, P.A. (1993) Estimating character weights during tree search. Cladistics, 9, 83–91.
Goloboff, P.A. (1999) Analyzing large data sets in reasonable times: solutions for composite optima. Cladistics, 15, 415–428.
Goloboff, P.A. (2000) The family Gallieniellidae (Araneae, Gnaphosoidea) in the Americas. Journal of Arachnology, 28, 1–6.
Goloboff, P.A., Carpenter, J.M., Arias, J.S. & Miranda Esquivel, D.R. (2008b) Weighting against homoplasy improves phylogenetic analysis of morphological data sets. Cladistics, 24, 758–773.
Goloboff, P.A., Farris, J.S. & Nixon, K.C. (2003) TNT, Tree Analysis using New Technology. Version 1.1, sponsored by the Willy Hennig Society. Available from: http://www.cladistics.com (2008-12-30).
Goloboff, P.A., Farris, J.S. & Nixon, K.C. (2008a) TNT, a free program for phylogenetic analysis. Cladistics, 24, 774–786.
Grismado, C.J. (2004) Una nueva especie del género Meriola Banks de Argentina (Araneae, Corinnidae, Trachelinae). Revista Ibérica de Aracnología, 10, 233–235.
Haddad, C.R. (2006) Spinotrachelas, a new genus of tracheline sac spiders from South Africa (Araneae: Corinnidae). African Invertebrates, 47, 85–93.
Haddad, C.R. & Lyle, R. (2008) Three new genera of tracheline sac spiders from southern Africa (Araneae: Corinnidae). African Invertebrates, 49, 37–76.
Harbach, R.E. & Kitching, I.J. (1998) Phylogeny and classification of the Culicidae (Diptera). Systematic Entomology, 23, 327–370.
Hawkins, J.A., Hughes, C.E. & Scotland, R.W. (1997) Primary homology assessment, characters and character states. Cladistics, 13, 275–283.
Jocqué, R. & Dippenaar-Schoeman, A.S. (2006) Spider Families of the World. Musée Royal de l'Afrique Centrale, Tervuren, 336 pp.
Kaila, L. (1999) Phylogeny and classification of the Elachistidae s.s. (Lepidoptera: Gelechioidea). Systematic Entomology, 24, 139–169.
Lawrence, R.F. (1938) A collection of spiders from Natal and Zululand. Annals of the Natal Museum, 8, 455–524.
Lawrence, R.F. (1942) A contribution to the araneid fauna of Natal and Zululand. Annals of the Natal Museum, 10, 141–190.
Lehtinen, P. (1967) Classification of the cribellate spiders and some allied families, with notes on the evolution of the suborder Araneomorpha. Annales Zoologici Fennici, 4, 199–468.
Lyle, R. (2008) A review of the Afrotropical tracheline sac spiders (Araneae: Corinnidae), with revisions of three genera. Unpublished MSc dissertation, University of the Free State, Bloemfontein, 364 pp.
Lyle, R. & Haddad, C.R. (2006a) A revision of the Afrotropical tracheline sac spider genus Thysanina Simon, 1910 (Araneae: Corinnidae). African Invertebrates, 47, 95–116.
Lyle, R. & Haddad, C.R. (2006b) Somatic morphology supports the transfer of Austrachelas Lawrence to Gallieniellidae (Arachnida: Araneae). Proceedings of the Microscopy Society of Southern Africa, 36, 57.
Maddison, W.P. (1993) Missing data versus missing characters in phylogenetic analysis. Systematic Biology, 42, 576–581.
Maddison, W.P., Donoghue, M.J. & Maddison, D.R. (1984) Outgroup analysis and parsimony. Systematic Zoology, 33, 83–103.
Nixon, K.C. (2002) Winclada version 1.00.08. [Computer programme and documentation]. Available from: http://www.cladistics.com (2008-12-30).
Nixon, K.C. & Carpenter, J.M. (1996) On consensus, collapsibility, and clade concordance. Cladistics, 12, 305–321.
Platnick, N.I. (1984) Studies on Malagasy spiders, 1. The family Gallieniellidae. American Museum Novitates, 2801, 1–17.
Platnick, N.I. (1990) A new species of Legendrena (Araneae: Gallieniellidae) from Madagascar. Journal of the New York entomological Society, 98, 499–501.
Platnick, N.I. (1995) New species and records of the ground spider family Gallieniellidae (Araneae, Gnaphosoidea) from Madagascar. Journal of Arachnology, 23, 9–12.
Platnick, N.I. (2002) A revision of the Australiasian ground spiders of the families Ammoxenidae, Cithaeronidae, Gallieniellidae, and Trochanteriidae (Araneae: Gnaphosoidea). Bulletin of the American Museum of Natural History, 271, 1–243.
Platnick, N.I. (2009) The World Spider Catalog, version 9.5. The American Museum of Natural History. Available from: http://research.amnh.org/entomology/spiders/catalog (2009-05-11).
Platnick, N.I. & Ewing, C. (1995) A revision of the tracheline spiders (Araneae, Corinnidae) of Southern South America. American Museum Novitates, 3128, 1–41.
Platnick, N.I. & Shadab, M.U. (1974) A revision of the tranquillus and speciosus groups of the spider genus Trachelas (Araneae, Clubionidae) in North and Central America. American Museum Novitates, 2553, 1–34.
Sanderson, M.J. & Donoghue, M.J. (1989) Patterns of variation in levels of homoplasy. Evolution, 42, 1781–1795.
Simon, E. (1897) Histoire Naturelle des Araignées. Tome 2. Roret, Paris, 1080 pp.
Swofford, D.L. (2002) PAUP* Phylogenetic Analysis under Parsimony (*and other methods) version 4.0 beta 10. Sinauer Associates, Sunderland, MA.
Tucker, R.W.E. (1920) Contributions to the South African Arachnid Fauna. II. On some new South African spiders of the families Barychelidae, Dipluridae, Eresidae, Zodariidae, Heracliidae, Urocteidae, Clubionidae. Annals of the South African Museum, 17, 439–488.
Tucker, R.W.E. (1923) The Drassidae of South Africa. Annals of the South African Museum, 19, 251–437.
Ubick, D., Paquin, P., Cushing, P. & Roth, V. (Eds.) (2005) Spiders of North America: an identification manual. American Arachnological Society, 377 pp.
Warui, C. & Jocqué, R. (2002) The first Gallieniellidae (Araneae) from eastern Africa. Journal of Arachnology, 30, 307–315.
Watrous, L.E. & Wheeler, Q.D. (1981) The out-group comparison method of character analysis. Systematic Zoology, 30, 1–11.
Wilkinson, M. (1995) Arbitrary resolutions, missing entries, and the problem of zero-length branches in parsimony analysis. Systematic Biology, 44, 108–111.
Wunderlich, J. (2008) Fossil and extant spiders. Beitr Araneology 5, Hirschberg, 850 pp.