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Published: 2009-07-28
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Apaturinae (Lepidoptera: Nymphalidae) from the Korean Peninsula: Synonymic Lists and Keys to Tribes, Genera and Species

Department of Ecology and Evolutionary Biology, University of Connecticut, 75 North Eagleville Road, Storrs, CT 06269, USA
Lepidoptera taxonomy Apaturini Pseudergolini Hestina persimilis species complex biology distribution

Abstract

This paper provides keys to tribes, genera and species and synonymic lists for the 12 butterfly species belonging to eight genera of Apaturinae (Lepidoptera: Nymphalidae) from the Korean Peninsula: Apatura ilia (Denis and Schiffermüller), Apatura metis Freyer, Apatura iris (Linnaeus), Hestina japonica (Felder and Felder), Hestina assimilis (Linnaeus), Sasakia charonda (Hewitson), Mimathyma schrenckii (Ménétriès), Mimathyma nycteis (Ménétriès), Sephisa princeps (Fixsen), Chitoria ulupi (Doherty), Dilipa fenestra (Leech) and Dichorragia nesimachus (Doyère). Species accounts include brief summaries of each taxon’s biology, taxonomy and life history. The distributional ranges of the Hestina persimilis species complex are discussed and illustrated.

References

  1. The publications that could not be checked are marked with an asterisk (*).

    Alphéraky, S. (1892) Lépidoptères rapportés de la Chine et de la Mongolie par G.N. Potanine (Suite). Mémoires sur les Lépidoptères rédigés par N.M. Romanoff, 6, 1–81, pls. 1–3.

    Bang-Haas, O. (1936) Neubeschreibungen und Berichtigungen der Palaearktischen Macrolepidopterenfauna XXV. Entomologische Zeitschrift, 50, 345–348.

    *Boisduval, J.B. ([1846]) explic, pl. 139 bis In: Cuvier, C. (Ed.), Le règne animal distribué d'après son organisation pour servir de base à l’histoire naturelle des animaux, et d’introduction a l’anatomie comparée. Édition par un réunion de disciples de Cuvier, M.M. Audouin, Blanchard, Deshayes, D’Orbigny, Doyère, Dugès Duvernoy, Laurillard, Milne-Edwards, Roulin et Valenciennes. 6 (Vol. 4), Roy. 8vo. Paris.

    Bollow, C. (1930) Apatura. In: Seitz, A. (Ed.), Die Gross-Schmetterlinge der Erde, eine systematische Bearbeitung der bis jetzt bekannten Gross-Schmetterlinge, I. Abteilung: Die Gross-Schmetterlinge des palaearktischen Faunengebietes, Band I: Die palaearktischen Tagfalter, supplementband. Alfred Kernen, Stuttgart. 399 pp., 16 pls.

    Bryk, F. (1938) In: Stichel, H. (Ed.), Nymphalidae: I. Dioninae, Anetiinae, Apaturinae. Lepidopterorum catalogus, Vol. 86. pp. 1–374.

    Butler, A.G. ([1869]) A monographic revision of the Lepidoptera hitherto included in the genus Adolias, with descriptions of new genera and species. Proceedings of the Scientific Meetings of the Zoological Society of London, 1868, 599–615, pl. 45.

    Butler, A.G. (1873) Descriptions of new species of Lepidoptera. Cistula entomologica, 1, 151–177.

    Butler, A.G. (1874) Descriptions of some new species and a new genus of diurnal Lepidoptera, in the collection of Herbert Druce, Esq. The Transactions of the Entomological Society of London, 1874, 423–436.

    Butler, A.G. (1883) On Lepidoptera from Manchuria and the Corea. The Annals and Magazine of Natural History, (5), 11, 109–117.

    Chou, I. [ed.] (1994) Monographia Rhopalocerorum Sinensium, 1–2. Henan Scientific and Technological Publishing House, Zhengzhou. 854 pp. (In Chinese)

    Chou, I. (1998) Classification and identification of Chinese butterflies. Henan Scientific and Technological Publishing House, Zhengzhou. 349+90 pp. (In Chinese)

    Dantchenko, A., Sourakov, A., & Emmel, T.C. (1996) Notes on the life history of Sephisa princeps in eastern Russia (Lepidoptera: Nymphalidae). Holarctic Lepidoptera, 3, 47–57.

    *Denis, J.N.C.M. & Schiffermüller, I. (1775) Ankündung eines systematischen Werkes von den Schmetterlingen der Wienergegend. Augustin Bernardi, Wien. 323 pp., 2 pls.

    Doherty, W. (1889) Notes on Assam butterflies. Journal of the Asiatic Society of Bengal, 58, 118–134, pl. 10.

    Doi, H. (1919) [A list of butterflies from Korea.] Chosen Iho, 58, 115–128. (In Japanese)

    Doi, H. (1928) [The species of the genus Apatura from Korea.] Journal of Chosen Natural History Society, (7), 49–50. (In Japanese)

    Doi, H. (1931) A list of Rhopalocera from Mount Shouyou, Keiki-Do, Korea. Journal of Chosen Natural History Society, (12), 42–47. (In Japanese)

    Doi, H. (1933) [Miscellaneous notes on insects (3).] Journal of Chosen Natural History Society, (15), 85–96. (In Japanese)

    Doubleday, E. (1845) Descriptions of new or imperfectly described diurnal Lepidoptera. The Annals and Magazine of Natural History, 16, 232–236.

    *Doyère, L. ([1840]) pl. 139 bis In: Cuvier, C. (Ed.), Le règne animal distribué d'après son organisation pour servir de base à l’histoire naturelle des animaux, et d’introduction a l’anatomie comparée. Edition accompagnée de planches gravées, representant les types de tous les genres, les caractères distinctifs des divers groupes, et les modifications de structure sur lesquelles repose cette classification; par une réunion de disciples de Cuvier. (Edn 3), Atlas, Les insectes, avec un atlas, par Audouin, Blanchard, Doyère, et Milne-Edwards, 2.

    Druce, H.H. (1911) Description of a new nymphaline butterfly from British India. The Transactions of the Entomological Society of London, 1911, 187–188, pl. 29.

    Dubatolov, V.V. & Korshunov, J.P. (1984) New data on taxonomy of the butterflies of the USSR (Lepidoptera, Rhopalocera). In: [Arthropods and helmints.] (Nauka Press. Siberian Dept., Novosibirsk) New and little known species of Siberian fauna, 17, 51–57. (In Russian with English summary)

    Fabricius, J.C. (1807) Nach Fabricii systema glossatorum, tom. I. In: Illiger, J.C. (Ed.), Die neueste Gattungs-Eintheilung der Schmetterlinge aus den Linnéischen Gattungen Papilio und Sphinx. Magazin für Insektenkunde, Braunschweig, 6, 279–289.

    Felder, C. & Felder, R. (1859) Lepidopterologische Fragmente. Wiener entomologische Monatschrift, 3, 178–186, pls. 3–4.

    Felder, C. & Felder, R. (1862) Observationes de Lepidopteris nonnullis Chinae centralis et Japoniae. Wiener entomologische Monatschrift, 6, 22–32.

    Fixsen, C. (1887) Lepidoptera aus Korea. Mémoires sur les Lépidoptères rédigés par N.M. Romanoff, 3, 233–319, pls. 13–14.

    Freitas, A.V.L. & Brown, K.S. Jr. (2004) Phylogeny of the Nymphalidae (Lepidoptera). Systematic Biology, 53, 363–383.

    Freyer, C.F. (1829) Beiträge zur Geschichte europäischer Schmetterlinge mit Abbildungen nach der Natur, 2. Augsburg. 166 pp., 48 pls.

    Fruhstorfer, H. (1903) Neue Nymphaliden aus Ostasien. Deutsche entomologische Zeitschrift, Iris, 16, 22–42.

    Fruhstorfer, H. (1908) Lepidopterologisches Pêle-Mêle, V. Neue und seltene Rhopaloceren der Insel Formosa. Entomologische Zeitschrift, 22, 102–103.

    Gorbunov, P.Y. (2001) The butterflies of Russia: classification, genitalia, keys for identification (Lepidoptera: Hesperoidea and Papilionoidea). Russian Academy of Sciences, Institute of Plant and Animal Ecology, Ekaterinburg.

    Hall, A. (1935) New forms of Nymphalinae and Ithomiinae. The Entomologist, 68, 221–227, pl. 6.

    Harada, M. & Igarashi, S. (1993) On hibernating aspects of nymphalid larvae in Asia. Butterflies, 6, 11–18. (In Japanese with English abstract)

    Hemming, F. (1939) On five genera in the Lepidoptera Rhopalocera at present without valid names. The Proceedings of the Royal Entomological Society of London, (B), 8, 39.

    Herrich-Schäffer, G.A.W. ([1845]) In: ([1843–1856]). Systematische Bearbeitung der Schmetterlinge von Europa, zugleich als Text, Revision und Supplement zu Jakob Hübner's Sammlung europäischer Schmetterlinge, 1. Die Tagfalter. G.J. Manz, Regenburg. 164 pp., 134 pls.

    Hewitson, W.C. (1861) Descriptions of some butterflies from the collection of Mr. Wallace. Proceedings of the Scientific Meetings of the Zoological Society of London, 1861, 50–52, pls. 8–9.

    Hewitson, W.C. ([1863]) Illustrations of new species of exotic butterflies, selected chiefly from the collections of W. Wilson Saunders and William C. Hewitson, 3. Van Voorst, London. iv+124 pp., 60 pls.

    Higgins, L.G. & Riley, N.D. (1983) A field guide to the butterflies of Britain and Europe, fourth edition. Collins, London. 384 pp.

    *Hübner, J. ([1806]) Tentamen determinationis digestionis atque denominationis singularum stirpium Lepidopterorum, peritis ad inspiciendum et dijudicandum communicatum, a Jacobo Hübner. Privately published, Augsburg. 2 pp. [rejected for nomenclatural purposes by the International Commission on Zoological Nomenclature. 1926. Smithsonian miscellaneous Collections, 73(4) (Opinion 97), 19.]

    Inomata, T. (1982) Appendix. pp. i–xxiv In: Mori, T., Doi, H., &Cho, F., Coloured butterflies from Korea (rep. edn.). Scientist Inc., Tokyo. 30 pls., 86+23+xxiv pp. (In Japanese)

    Joo, H.-Z., Kim, S.-S., & Sohn, J.-D. (1997) Butterflies of Korea. Kyo-Hak Publishing Co., Ltd., Seoul. 437 pp. (In Korean)

    Kim, H.K. (1965) An ecological studies of the butterflies of Korea with emphasis on the hibernation forms and food plants. Journal of Korean Cultural Research Institute, 5, 241–258. (In Korean with English abstract)

    Kim, H.K. & Mi, S.W. (1956) [Correction of the scientific names of Korean butterfly list.] Commemorative Theses for the 70th Anniversary of Ewha Womans University, Seoul. pp. 377–405. (In Korean)

    Kim, S.-S. & Sohn, J.-D. (1992) Life history of Athymodes nycteis Ménètriès in Korea. Journal of the Amateur Lepidopterists' Society of Korea, 5, 24–28. (In Korean with English abstract)

    Kishida, K. & Nakamura, Y. (1936) A catalogue of Japanese insects, fasc. 9. Lepidoptera. The Entomological World, Tokyo, 4, 433–673. (In Japanese)

    Kollar, V. ([1844]) In: Kollar V. & Redtenbacher, L., Aufzählung und Beschreibung der von Freiherrn Carl v. Hügel auf seiner Reise durch Kaschmir und das Himaleyagebirge gesammelten Insecten. In: von Hügel, F.C. (Ed.), Kaschmir und das Reich der Siek, Vol. 4(2). pp. 393–564, 582–585, pls. 1–28.

    Korshunov, Y.P. & Gorbunov, P. (1995) [Butterflies of the Asian part of Russia. A handbook] (Dnevnye babochki aziatskoi chasti Rossii. Spravochnik). Ural University Press, Ekaterinburg. 202 pp. (In Russian)

    Kristensen, N.P., Scoble, M.J., & Karsholt, O. (2007) Lepidoptera phylogeny and systematic: the state of inventorying moth and butterfly diversity. Zootaxa, 1668, 699–747.

    Kurentzov, A.I. (1937) Some new and interesting scale-winged species (Lepidoptera) from Sikhote-Alin. Bulletin of the Far Eastern Branch of the Academy of Sciences of the USSR, 26, 115–132. (In Russian with English summary)

    Kurentzov, A.I. (1970) The butterflies of the Far East USSR. Nauka, Leningrad. 164 pp., 14 pls. (In Russian)

    de Lamarck, M.C. (1819) Histoire naturelle des animaux sans vertèbres, présentant les caractères généraux et particuliers de ces animaux, leur distribution, leurs classes, leurs familles, leurs genres, et la citation des principales espèces qui s’y rapportent; précédée d’une introduction offrant la détermination des caractères essentiels de l’animal, sa distinction du végétal et des autres corps naturels, enfin, l’exposition des principes fondamentaux de la zoologie, Vol. 6, Part 1. Paris. 232 pp.

    Lee, S.-M. (1971) The butterflies of Mt. Seol-Ak. Cheong-Ho-Rim Entomological Laboratory Note, 1, 1–16. (In Korean)

    Lee, S.-M. (1973) A list of butterflies from Mt. Seol-Ak, Korea. Cheong-Ho-Rim Entomological Laboratory Note, 4, 1–10. (In Korean)

    Lee, S.-M. (1978) Notes on the Purple Butterfly (Apatura ilia Schiffermüller) in Korea (Lep.: Nymphalidae). Korean Journal of Entomology, 8, 39–40. (In Korean with English abstract)

    Lee, S.-M. (1982) Butterflies of Korea. Editorial Committee of Insecta Koreana, Seoul. 125 pp., 63 pls.

    Lee, S.-M. (1992) Explanation of genus Apatura (Nymphalidae) in Korean peninsula. Journal of the Amateur Lepidopterists' Society of Korea, 5, 1–5. (In Korean with English abstract)

    Lee, S.-M. & Takakura, T. (1981) On a new subspecies of the Purple Emperor, Apatura iris (Lepidoptera: Nymphalidae), from the Republic of Korea. Tyô to Ga, 31, 133–141.

    Lee, Y.J. (2005) A list of butterflies from Korea with notes on some changes in scientific names. Lucanus, Seoul, 5, 18–28. (In Korean with English abstract)

    Leech, J.H. (1887) On the Lepidoptera of Japan and Corea.―part I. Rhopalocera. Proceedings of the Scientific Meetings of the Zoological Society of London, 1887, 398–431, pls. 35–36.

    Leech, J.H. (1890a) New species of Lepidoptera from China. The Entomologist, 23, 26–50.

    Leech, J.H. (1890b) New species of Rhopalocera from China. The Entomologist, 23, 187–192.

    Leech, J.H. (1891) New species of Rhopalocera from North-West China. The Entomologist, 24(Supplement), 23–31.

    Leech, J.H. (1893) Butterflies from China, Japan, and Corea, Part II. R.H. Porter, London. pp. 115–285, pls. 14–26.

    Linnaeus, C. (1758) Systema naturae, per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Tomus I. Editio decima, reformata. Holmiae. pp. 1–824.

    Maruda, S. (1929) [A list of Lepidoptera in Kyojyo and its vicinity.] Bulletin of the Agricultural Experiment Station, Government-General of Chosen, 4(2), 125–135. (In Japanese)

    Masui, A. & Inomata, T. (1990) Apaturinae of the world (Lepidoptera, Nymphalidae) -1-. Yadoriga, 143, 2–10. (In Japanese)

    Masui, A. & Inomata, T. (1991) Apaturinae of the world (Lepidoptera, Nymphalidae) -2-. Yadoriga, 146, 2–14. (In Japanese)

    Masui, A. & Inomata, T. (1993) Apaturinae of the world (Lepidoptera, Nymphalidae) -5-. Yadoriga, 155, 2–11.

    Masui, A. & Inomata, T. (1994) Apaturinae of the world (Lepidoptera, Nymphalidae) -6-. Yadoriga, 157, 2–12. (In Japanese with English abstract)

    Matsumura, S. (1907) Thousand insects of Japan, 4. Tokyo. 147 pp.

    Matsumura, S. (1910) Neue Rhopaloceren aus Formosa. Entomologische Zeitschrift, 23, 209.

    Matsumura, S. (1928) New butterflies especially from the Kuriles. Insecta matsumurana, 2, 191–201.

    Matsumura, S. (1931) New species and new forms of butterflies from Japan. Insecta matsumurana, 6, 43–45.

    Ménétriès, M. (1859) Lepidoptères de la Sibérie orientale et en particulier des rives de l'Amour. Bulletin de la Classe Physico-Mathématique de l'Académie Impériale des Sciences de St.-Pétersbourg, 17, 212–221.

    Moore, F. (1858) Descriptions of three new species of diurnal Lepidoptera. The Annals and Magazine of Natural History, (3), 1, 48–49.

    Moore, F. ([1858]) In: Horsfield T. & Moore, F., A catalogue of the lepidopterous insects in the Museum of the Hon. East-India Company, (1). Wm. H. Allen and Co., London. v, [ii]+1–278+IV+11 pp., 18 pls, expl.

    Moore, F. ([1866]) On the lepidopterous insects of Bengal. Proceedings of the Scientific Meetings of the Zoological Society of London, 1865, 755–823, pls. 41–43.

    Moore, F. (1882) List of the Lepidoptera collected by the Rev. J.H. Hocking, chiefly in the Kangra District, N.W. Himalaya; with descriptions of new genera and species.―part I. Proceedings of the Scientific Meetings of the Zoological Society of London, 1882, 234–262, pls. 11–12.

    Moore, F. ([1896]) Lepidoptera Indica, 3. L. Reeve & Co., London. pp. 1–48, pls. 191–205.

    Mori, T., Doi, H., & Cho, F. (1934) Coloured butterflies from Korea. 109 pp., 30 pls. (In Japanese)

    Murayama, S.-I. (1963) Remarks on some butterflies from Japan and Korea, with descriptions of 2 races, 1 form, 4 aberrant forms. Tyô to Ga, 14, 43–49. (In Japanese with English resume)

    Nakayama, S. (1932) A guide to general information concerning Corean butterflies. Bulletin of the Suigen Imperial College of Agriculture and Forestry, 25th Anniversary, Suwon. pp. 366–386, pls. 1–3.

    de Nicéville, L. (1900) On new and little-known Lepidoptera from the Oriental Region. The Journal of the Bombay Natural History Society, 13, 157–176, pls. CC–EE.

    Nire, K. (1918) Gen. Apatura Fabricius, 1807, Gen. Sephisa Moore, 1882 and Gen. Sasakia Moore, 1895. In: A catalogue of the butterflies of Japan. The Zoological Magazine, 30, 93–98.

    Oberthür, C. (1885) M.Ch. Oberthür adresse la note suivante. Bulletin des Séances et Bulletin bibliographique de la Société entomologique de France, 1885, 136.

    Oberthür, C. (1890) Lépidoptères de Chine. Études d'Entomologie: Faunes entomologiques; descriptions d'insectes nouveaux ou peu connus, 13, 35–40, pls. 9–10.

    Okamoto, H. (1924) The insect fauna of Quelpart Island (Saishiu-to). Bulletin of the Agricultural Experiment Station, Government-General of Chosen, 1(2), 47–233.

    Okano, M. (1998) The butterflies of Chejudo (Quelpart Island). Fuji Daigaku Kiyo, 31(2), 1–10, pl. 1.

    Schultz, O. (1904) Über einige in Schlesien gefangene interessante Lepidopteren―Aberrationen aus den Gattungen Apatura F. und Limenitis F. Abhandlungen der Naturforschenden Gesellschaft zu Görlitz, 24, 129–136.

    Seok, D.M. (1934) Papilioj en Koreujo (Unua Raporto). Bulletin of the Kagoshima Imperial College of Agriculture and Forestry, 25th Anniversary, 1, 631–784.

    Seok, D.M. (1936) Pri la Du Novaj Specoj de Papilioj, Neptis okazimai kaj Zephyrus ginzii. The Zoological Magazine, 48, 60–65. (In Japanese with Esperanto résumé)

    Seok, D.M. (1937) On two new subspecies of butterflies. Zephyrus, 7, 29–32. (In Japanese)

    Seok, D.M. (1938) On two new forms of Korean butterflies. Zephyrus, 7, 241–243. (In Japanese)

    Seok, D.M. (1939a) Travels after butterflies in the plateau Kaima Korea. The Entomological World, Tokyo, 7, 168–186. (In Japanese)

    Seok, D.M. (1939b) A synonymic list of butterflies of Korea (Tyōsen). Korea Branch of the Royal Asiatic Society, Seoul. xxxl+391 pp., 2 pls.

    Seok, D.M. (1942) Papilioj en Songdo, Koreujo (revizita eldono). Journal of Chosen Natural History Society, 9(35), 86–94. (In Japanese)

    Seok, D.M. (1973) The distribution maps of butterflies in Korea. Po Chin Chai, Ltd., Seoul. 517 pp. (In Korean)

    Seok, D.M. & Umitatsu, T.J. (1942) Butterflies of S. Heiandō. Journal of Chosen Natural History Society, 9(37), 185–192. (In Japanese)

    Shimagami, K. (2000) Geographical variations of Dichorragia nesimachus (Doyére) in the regions around the Korean Peninsula. Gekkan-Mushi, 352, 12–27.

    Shin, Y.-H. (1975) Notes on the butterflies of Kwangnung, Korea (supplement). Journal of Research Institute of Science and Technology, Kyung Hee University, Seoul, 3, 41–47. (In Korean with English abstract)

    Shirôzu, T. (1938) [Explanation of a rare butterfly Apatura ilia mikuni Wileman.] Bulletin of Miyazaki College of Agriculture and Forestry, 25, 1–15. (In Japanese)

    Shirôzu, T. (1955) New or little known butterflies from the North-Eastern Asia, with some synonymic notes, III. Sieboldia, 1, 229–236, pls. 30–34.

    Snellen, P.C.T. (1894) Lepidopterologische Aanteekeningen. Tijdschrift voor Entomologie, 37, 67–72.

    Sohn, J.-D. (1995) Life history of Sephisa princeps Fixsen (Lepidoptera: Nymphalidae) in Korea. Journal of the Lepidopterists' Society of Korea, 8, 1–6. (In Korean with English abstract)

    Sohn, J.-D. & Kim, S.-S. (1990) Life history of Apatura iris Linnaeus in Korea. Journal of the Amateur Lepidopterists' Society of Korea, 3, 40–44. (In Korean with English summary)

    Sohn, J.-D. & Kim, S.-S. (1993) Life histroy of Mimathyma schrenckii (Ménétriès) in Korea. Journal of the Amateur Lepidopterists' Society of Korea, 6, 4–8. (In Korean with English abstract)

    Staudinger, O. & Rebel, H. (1901) Catalog der Lepidopteren des palaearctischen Faunengebietes, 1. Theil: Famil. Papilionidae–Hepialidae. R. Friedländer & Sohn, Berlin. xxxii+368 pp.

    Stichel, H. (1908) ApaturaHestina. pp. 160–193 In: Seitz, A. (Ed.), Die Gross-Schmetterlinge der Erde, eine systematische Bearbeitung der bis jetzt bekannten Gross-Schmetterlinge, I. Abteilung: Die Gross-Schmetterlinge des palaearktischen Faunengebietes, Band I: Die palaearktischen Tagfalter: Textband / Tafelband. Stuttgart. 379 pp., 89 pls.

    Sugitani, I. (1932a) Some butterflies from N.E. Corea, new to the fauna of the Japanese Empire. Zephyrus, 4, 15–30. (In Japanese)

    Sugitani, I. (1932b) On two Corean nymphalid butterflies. Zephyrus, 4, 100–102. (In Japanese)

    Takakura, T. (1981) The Lesser Purple Emperors of Japan. Nature and Insects, Tokyo, 16(2), 4–9. (In Japanese)

    Tuzov, V.K. [ed.], Bogdanov, P.V., Churkin, S.V., Devyatkin, A.L., Dantchenko, A.V., Murzin, V.S., Samodurov, G.D., & Zhdanko, A.B. (2000) Guide to the butterflies of Russia and adjacent territories, 2: Libytheidae, Danaidae, Nymphalidae, Riodinidae, Lycaenidae. Pensoft, Sofia-Moscow. 580 pp.

    Tytler, H.C. (1926) Notes on some new and interesting butterflies from India and Burma. The Journal of the Bombay Natural History Society, 31, 579–590, pls. 1–3, 5.

    Wahlberg, N., Weingartner, E., & Nylin, S. (2003) Towards a better understanding of the higher systematics of Nymphalidae (Lepidoptera: Papilionoidea). Molecular Phylogenetics and Evolution, 28, 473–484.

    Wahlberg, N., Brower, A.V.Z., & Nylin, S. (2005) Phylogenetic relationships and historical biogeography of tribes and genera in the subfamily Nymphalinae (Lepidoptera: Nymphalidae). Biological Journal of the Linnean Society, 86, 227–251.

    Watkins, H.T.G. (1928) New species of Parhestina (Lep., Nymphalidae). The Annals and Magazine of Natural History, (10), 2, 91–93.

    Westwood, J.O. ([1850]) In: Doubleday, E. & Westwood, J.O. [1848–1852], The genera of diurnal Lepidoptera: comprising their generic characters, a notice of their habits and transformations, and a catalogue of the species of each genus. Illustrated by W.C. Hewitson, Vol. 2. Longman, Brown, Green, and Longmans, London. pp. 251–534, pls. 31–79, supplementary plate.

    Wileman, A.E. (1910) Some new butterflies from Formosa and Japan. The Entomologist, 43, 92–93.

    Yoon, I.-H. & Kim, S.-S. (1989) Some notes on the life history of Dilipa fenestra Leech (Lepidoptera, Nymphalidae). Journal of the Amateur Lepidopterists' Society of Korea, 2, 60–63. (In Korean with English summary)

    Zhang, M., Cao, T., Zhang, R., Guo, Y., Duan, Y., & Ma, E. (2007) Phylogeny of Apaturinae butterflies (Lepidoptera: Nymphalidae) based on mitochondrial cytochrome oxidase I gene. Journal of Genetics and Genomics, 34, 812–823.